This investigation on the carbohydrate values within Vitis vinifera is included in a general study of nutrition. Presumably, a better understanding of the physiology of the grapevine would result in improved viticultura! practices. It would serve as a suitable basis for evaluating the analytical results obtained by studying the carbohydrate nutrition of the vine and also for interpreting data on pathologically affected vines. Certain points were pertinent. Investigators have often suggested that carbohydrates are translocated during the dormant period. If this translocation does take place, its bearing on the time of pruning would be important. Neither Richey and Bowers (19) nor Schrader (22) found such translocation in the Concord grape. It would, indeed, appear unlikely; the callus closure of the sieve plates during the dormant season seems to render the accepted channel of elaborated food transport unavailable. Another important point was the significance of hemicellulose as a reserve foo?l. Data on this have already been published (35). There were no indications that hemicellulose was available as a reserve in the grape even under the severe deficiency conditions resulting from repeated defoliation. A third point was brought forth by findings such as those of Davis (5) with the sugar prune. According to Davis, there is a comparatively large spring increase of starch in the bark and wood of the spurs?an increase not explained by the decrease in sugar content. Richey and Bowers (19) and Schrader (22) also found that the total carbohydrate content of Concord grape stems became markedly greater during the dormant period. Such findings might hypothetically be explained by one or several assumptions: (1) by transformation to or from other materials, such as fat, hemicellulose, or unknown substances; (2) by transport of sugars from another part of the tree; or (3) by sampling and analytical error. Davis in an unpublished analysis of the same material could not account for this increase by the disappearance of hemicellulose. The data of Richey and Bowers (19) and of Schrader (22) presumably include the hemicellulosic fraction in the total carbohydrates. A potential source of such materials might be a fatty reserve; and there remains the possibility of such unevaluated substances as Swarbrick (28) and Sinnott (24) have observed in microchemical studies, or some fraction of the comparatively great amount of nonsugar alcohol-soluble substance observed by Niklewski (16). Schrader (22) has found much of this soluble nonsugar material in Concord grape shoots. According to Smyth (27), $n extremely large quantity of it occurs in young apple wood. The improbability of the translocation hypothesis has already ? This is the second of a series of papers on the carbohydrate metabolism of Vitis vinifera.