Abstract. The Tibetan Plateau (TP) contains the headwaters of major Asian rivers that sustain billions of people and plays an important role in both regional and global climate through thermal and mechanical forcings. Understanding the characteristics and changes to the hydrological regimes on the TP during the mid-Holocene (MH) will help in understanding the expected future changes. Here, an analysis of the hydroclimates over the headwater regions of three major rivers originating in the TP, namely the Yellow, Yangtze, and Brahmaputra rivers, is presented, using dynamically downscaled climate simulations constructed using the Weather Research and Forecasting Model (WRF) coupled to the hydrological model WRF-Hydro. Green Sahara (GS) boundary conditions have also been incorporated into the global model so as to capture the remote feedbacks between the Saharan vegetation and the river hydrographs over the TP. Model–data comparisons show that the dynamically downscaled simulations significantly improve the regional climate simulations over the TP in both the modern day and the MH, highlighting the crucial role of downscaling in both present-day and past climates. TP precipitation is also strongly affected by the greening of the Sahara, with a particularly large increase over the southern TP, as well as a delay in the monsoon withdrawal. The simulation results were first validated over the upper basins of the three rivers before the hydrological responses to the MH forcing for the three basins were quantified. Both the upper Yellow and Yangtze rivers exhibit a decline in streamflow during the MH, especially in summer, which is a combined effect of less snowmelt and stronger evapotranspiration. The GS forcing caused a rise in temperature during the MH, as well as larger rainfall but less snowfall and greater evaporative water losses. The Brahmaputra River runoff is simulated to increase in the MH due to greater net precipitation.