Deciduous Stands Research Articles

Effects of Disturbance and Climate Change on Ecosystem Performance in the Yukon River Basin Boreal Forest

A warming climate influences boreal forest productivity, dynamics, and disturbance regimes. We used ecosystem models and 250 m satellite Normalized Difference Vegetation Index (NDVI) data averaged over the growing season (GSN) to model current, and estimate future, ecosystem performance. We modeled Expected Ecosystem Performance (EEP), or anticipated productivity, in undisturbed stands over the 2000–2008 period from a variety of abiotic data sources, using a rule-based piecewise regression tree. The EEP model was applied to a future climate ensemble A1B projection to quantify expected changes to mature boreal forest performance. Ecosystem Performance Anomalies (EPA), were identified as the residuals of the EEP and GSN relationship and represent performance departures from expected performance conditions. These performance data were used to monitor successional events following fire. Results suggested that maximum EPA occurs 30–40 years following fire, and deciduous stands generally have higher EPA than coniferous stands. Mean undisturbed EEP is projected to increase 5.6% by 2040 and 8.7% by 2070, suggesting an increased deciduous component in boreal forests. Our results contribute to the understanding of boreal forest successional dynamics and its response to climate change. This information enables informed decisions to prepare for, and adapt to, climate change in the Yukon River Basin forest.

Effects of forest management on ground-dwelling beetles (Coleoptera; Carabidae, Staphylinidae) in Central Europe are mainly mediated by changes in forest structure

Forest management is known to influence species diversity of various taxa but inconsistent or even contrasting effects are reported for arthropods. Regional differences in management as well as differences in regional species pools might be responsible for these inconsistencies, but, inter-regional replicated studies that account for regional variability are rare. We investigated the effect of forest type on the abundance, diversity, community structure and composition of two important ground-dwelling beetle families, Carabidae and Staphylinidae, in 149 forest stands distributed over three regions in Germany. In particular we focused on recent forestry history, stand age and dominant tree species, in addition to a number of environmental descriptors. Overall management effects on beetle communities were small and mainly mediated by structural habitat parameters such as the cover of forest canopy or the plant diversity on forest stands. The general response of both beetle taxa to forest management was similar in all regions: abundance and species richness of beetles was higher in older than in younger stands and species richness was lower in unmanaged than in managed stands. The abundance ratio of forest species-to-open habitat species differed between regions, but generally increased from young to old stands, from coniferous to deciduous stands and from managed to unmanaged stands. The response of both beetle families to dominant tree species was variable among regions and staphylinid richness varied in the response to recent forestry history. Our results suggest that current forest management practices change the composition of ground-dwelling beetle communities mainly by favoring generalists and open habitat species. To protect important forest beetle communities and thus the ecosystem functions and services provided by them, we suggest to shelter remaining ancient forests and to develop near-to-nature management strategies by prolonging rotation periods and increasing structural diversity of managed forests. Possible geographic variations in the response of beetle communities need to be considered in conservation-orientated forest management strategies.

Forest herbs show species-specific responses to variation in light regime on sites with contrasting soil acidity: An experiment mimicking forest conversion scenarios

Forest conversion from native deciduous forests to coniferous stands has been performed in many European regions and resulted in dramatic shifts in understorey plant community composition. However, the drivers for changes in specific understorey plant species remained unclear.Here, we experimentally determine the species-specific effects of light availability and chemical soil characteristics, on the vegetative and regenerative performance of five herbaceous forest understorey plants. Topsoil samples from both spruce and deciduous stands at four locations, with two levels of soil acidity, were collected and used in a common garden experiment. Additionally, three different light levels were applied, i.e., ‘light deciduous’, ‘dark deciduous’ (extra light reduction during summer) and ‘evergreen’ (light reduction during winter). In a second experiment we evaluated the germination of two of these species against the acidity and tree species at the site of origin of the soil samples.The light regime affected both the vegetative and regenerative performance of the understorey species: compared to light deciduous, Anemone nemorosa had a significantly lower performance under the evergreen light regime, Convallaria majalis under dark deciduous and Luzula luzuloides and Galium odoratum under both light regimes. The vegetative performance was lower in soil from acid sites for the acid-sensitive species G. odoratum and Primula elatior. Differences between the soils sampled under deciduous or spruce stands had no effect on the vegetative, or the regenerative performance of these species. By contrast, the germination of L. luzuloides and P. elatior was higher in soils sampled in deciduous stands and in neutral sites.Species-specific responses in vegetative and regenerative performance of adult plants to a changed light regime and soil acidification could be a reason for the changed vegetation composition in converted stands. Also lower germination and establishment of forest understorey species in spruce stands could influence the species distribution after conversion.

Using dynamic N-mixture models to test cavity limitation on northern flying squirrel demographic parameters using experimental nest box supplementation.

Dynamic N-mixture models have been recently developed to estimate demographic parameters of unmarked individuals while accounting for imperfect detection. We propose an application of the Dail and Madsen (2011: Biometrics, 67, 577–587) dynamic N-mixture model in a manipulative experiment using a before-after control-impact design (BACI). Specifically, we tested the hypothesis of cavity limitation of a cavity specialist species, the northern flying squirrel, using nest box supplementation on half of 56 trapping sites. Our main purpose was to evaluate the impact of an increase in cavity availability on flying squirrel population dynamics in deciduous stands in northwestern Québec with the dynamic N-mixture model. We compared abundance estimates from this recent approach with those from classic capture–mark–recapture models and generalized linear models. We compared apparent survival estimates with those from Cormack–Jolly–Seber (CJS) models. Average recruitment rate was 6 individuals per site after 4 years. Nevertheless, we found no effect of cavity supplementation on apparent survival and recruitment rates of flying squirrels. Contrary to our expectations, initial abundance was not affected by conifer basal area (food availability) and was negatively affected by snag basal area (cavity availability). Northern flying squirrel population dynamics are not influenced by cavity availability at our deciduous sites. Consequently, we suggest that this species should not be considered an indicator of old forest attributes in our study area, especially in view of apparent wide population fluctuations across years. Abundance estimates from N-mixture models were similar to those from capture–mark–recapture models, although the latter had greater precision. Generalized linear mixed models produced lower abundance estimates, but revealed the same relationship between abundance and snag basal area. Apparent survival estimates from N-mixture models were higher and less precise than those from CJS models. However, N-mixture models can be particularly useful to evaluate management effects on animal populations, especially for species that are difficult to detect in situations where individuals cannot be uniquely identified. They also allow investigating the effects of covariates at the site level, when low recapture rates would require restricting classic CMR analyses to a subset of sites with the most captures.

Effects of Inundated Vegetation on X-Band HH–VV Backscatter and Phase Difference

TerraSAR-X HH and HH-VV polarization data of a wetland in Estonia are used to study the effects of HH-VV phase difference on detecting inundated vegetation. Patches with different tree species composition are selected from two images representing flooded and unflooded conditions. Contribution of the double bounce mechanism is studied via phase difference and backscatter analysis. An increase in both HH and HH-VV backscatter was observed for all study cases in flooded conditions, as compared to unflooded periods. Over the different patches, HH backscatter increased from 2.3 dB to 8.2 dB, while for HH-VV - from 3 dB to 9.8 dB. The HH-VV channel allowed for a better separation between flooded and un flooded forest than HH, with the difference between the HH-VV and HH backscatter increase in inundated conditions ranging from 0.2 dB for coniferous stands with a tree height of less than 10 m to 1.6 dB for deciduous stands with a tree height of less than 10 m. A larger separation on both channels was in general observed in deciduous stands as the leaf-off season allowed for a better penetration depth. A considerable phase shift between HH and VV channels ranging from 10° to 28° was observed due to inundation, suggesting a strong contribution from the double bounce mechanism. For unflooded patches, the difference in backscatter between the two acquisition dates ranged from -0.8 dB to 1 dB for HH, and from 0 dB to 2 dB for HH-VV channel. Phase shift for unflooded patches varied from 5° to 8°.

Productivity of a John Deere Harvester Unit in Deciduous Stands

The aim of the presented paper was to determine the productivity of a John Deere 1070D harvester in deciduous coppices of Quercus petraea and Fagus sylvatica, in thinnings over 50 years of age. The research area was located in Tribeč mountain ridge, western Slovakia. The terrain incline did not exceed 20%. The productivity of the harvester in oak stands was 9.97 m3.h−1 without downtimes (breaks, failures, etc.). Performance with breaks was 4.98 m3.PMH−1 (productive machine hour). The productivity of the harvester was influenced by a large portion of passes of the harvester, which constituted 23.47% of the time of the operating cycle. The large number of passes was caused by the fact that the operator did not respect the technologic process and a high portion of tree extraction (8.45%). Furthermore the productivity was decreased by frequent failures of the machine and downtimes, which could be prevented by proper and regular maintenance. The productivity in beech stands was 6.36 m3.h−1 without downtimes and 5.35 m3.PMH−1 with downtimes included. Lower productivity was caused by the numerous and thick branches and frequent presence of trees with multiple tree tops, ergo unsuitable structure of the stand. Absence of a worker with a chainsaw was considered to be a disadvantage, because this worker is often a necessity when harvesters operate in deciduous stands.

Variation in post-wildfire regeneration of boreal mixedwood forests: underlying factors and implications for natural disturbance-based management

To test the direct regeneration hypothesis and support natural disturbance-based forest management we characterized the structure and composition of boreal mixedwood forests regenerating after large wildfires and examined the influence of pre-fire stand composition and post-fire competing vegetation. In stands which had been deciduous (Populus sp.)-dominated, conifer (white spruce)-dominated, or mixed pre-fire we measured regeneration stocking (presence in 10 m2 plots), density and height 10–20 years post-burn in five wildfires in Alberta, Canada. Most plots regenerated to the deciduous or mixed stocking types; plots in the older fire and in stands that were pure conifer pre-fire had higher amounts of conifer regeneration. Surprisingly, regeneration in pre-fire ‘pure’ white spruce stands was most often to pine, although these had not been recorded in the pre-fire inventory. Pre-fire deciduous stands were the most resilient in that poplar species dominated their post-fire regeneration in terms of stocking, density and height. These stands also had the highest diversity of regenerating tree species and the most unstocked plots. High grass cover negatively affected regeneration density of both deciduous and conifer trees. Our results demonstrate the natural occurrence of regeneration gaps, pre- to post-fire changes in forest composition, and high variation in post-fire regeneration composition. These should be taken into consideration when developing goals for post-harvest regeneration mimicking natural disturbance.

The Relative Abundance of the Juvenile Phase of the Eastern Red-Spotted Newt at Harvard Forest Prior to the Arrival of the Hemlock Woolly Adelgid

The invasive insect pest Adelges tsugae (Hemlock Woolly Adelgid) threatens the ecologically unique Tsuga canadensis (Eastern Hemlock)-dominated forest type throughout its range. Relatively little is known about how the loss of this forest type will affect the relative abundance of amphibians. This study assessed the relative abundance of the juvenile phase of Notophthalmus viridescens viridescens (Eastern Red-spotted Newt, Red Eft) in Eastern Hemlock-dominated stands (n = 5) and mixed deciduous stands (n = 5) at Harvard Forest in Petersham, MA, using both transect surveys of the forest floor surface (n = 368 Red Eft observations over four seasons), and intensive searches of quadrats (n = 27 Red Eft observations over two seasons). Using data from transect surveys, the average relative abundance of Red Efts was more than two times greater in Eastern Hemlock-dominated stands than in mixed deciduous stands, however the differences were not statistically significant (P = 0.146). Quadrat surveys yielded relative abundance estimates for Red Efts that were more than 5 times greater in Eastern Hemlock-dominated stands than in mixed deciduous stands, but again the differences were not statistically significant (P = 0.213).

A Photogrammetric Workflow for the Creation of a Forest Canopy Height Model from Small Unmanned Aerial System Imagery

The recent development of operational small unmanned aerial systems (UASs) opens the door for their extensive use in forest mapping, as both the spatial and temporal resolution of UAS imagery better suit local-scale investigation than traditional remote sensing tools. This article focuses on the use of combined photogrammetry and “Structure from Motion” approaches in order to model the forest canopy surface from low-altitude aerial images. An original workflow, using the open source and free photogrammetric toolbox, MICMAC (acronym for Multi Image Matches for Auto Correlation Methods), was set up to create a digital canopy surface model of deciduous stands. In combination with a co-registered light detection and ranging (LiDAR) digital terrain model, the elevation of vegetation was determined, and the resulting hybrid photo/LiDAR canopy height model was compared to data from a LiDAR canopy height model and from forest inventory data. Linear regressions predicting dominant height and individual height from plot metrics and crown metrics showed that the photogrammetric canopy height model was of good quality for deciduous stands. Although photogrammetric reconstruction significantly smooths the canopy surface, the use of this workflow has the potential to take full advantage of the flexible revisit period of drones in order to refresh the LiDAR canopy height model and to collect dense multitemporal canopy height series.

Can pinewoods provide habitat for a deciduous forest specialist? A two-scale approach to the habitat selection of Bechstein's bat

Populations of Myotis bechsteinii in Sierra de Cazorla, Segura y Las Villas Natural Park (southwestern Spain) survive in a large, old-growth coniferous woodland, which is in contrast with known ecological preferences of the species. We tracked ten lactating females and studied patterns of habitat selection by Bechstein's bat (Myotis bechsteinii) at two spatial scales: macrohabitat and microhabitat, regarding forest essence (coniferous vs. deciduous). The tracked individuals always foraged within forested areas and did not use areas devoid of trees. At the macrohabitat level, no positive selection of deciduous stands was apparent, suggesting selection studies of coarse resolution may not be able to capture subtle selection patterns. At the microhabitat level Myotis bechsteinii selected deciduous patches within the coniferous matrix, therefore, our results corroborate the perception of this species as dependent of services provided by deciduous woodlands. Larger foraging home ranges and commuting distances as compared with other Mediterranean localities suggest that our studied population inhabits a marginal or suboptimal environment in terms of habitat quality. We argue that this population of Bechstein's bat has survived as a relict one probably as a consequence of fragmentation and transformation of deciduous forest ecosystems in the Mediterranean range.

Trophic ecology of mesostigmatan and oribatid mites in harvested and control coniferous and deciduous stands of the boreal mixedwood forest determined using 15N stable isotopes

In boreal forest ecosystems, thick forest floors are a key habitat for microorganisms, micro- and mesofauna that drive soil biochemical processes. While they are the least abundant of soil mesofauna, mesostigmatan mites play a central role in soils by regulating population dynamics of other meso- and microfauna and by affecting nitrogen mineralization rates. The most abundant and diverse oribatid mites are especially important for decomposition processes in boreal forests. In this study, we evaluated the trophic ecology of mesostigmatan and oribatid mites from Canadian boreal forests by using body δ15N signatures as indicators of their trophic position. We extracted mites from the forest floors of control and clearcut stands from coniferous and deciduous stands dominated by white spruce (Picea glauca (Moench) Voss), and trembling aspen (Populus tremuloides Michx.), respectively. The mite values spanned 10.2 δ units in 15N, and were enriched by at least 1.2‰ compared to forest floor (baseline) values. When compiled to known eating habits from published feeding experiments, isotopic values placed the mites in three main trophic guilds, including detritivorous, omnivorous (overlapping with predators), and predator mites. Isotopic nitrogen fractionation within the mites did not seem to be affected by their habitat (spruce vs. aspen), or clearcutting practices. Instead, in this boreal forest environment, the well-defined degree of isotopic fractionation within the food web may be solely dependent on the mite feeding relationships.

Hygroscopic properties of the litter of coniferous and deciduous stands in central Siberia

The hygroscopic properties of the litter of coniferous (larch and pine) and deciduous (aspen and birch) stands were studied in terms of isotherms of water-vapor sorption and with use of thermal analysis methods (TG, DSC). It was found that most of the monolayer capacity belongs to subhorizons OL and OF of the larch and pine forest litters. The larch plantation litter has the highest hydrophilicity. DSC results show that the water is most strongly associated with the litter substance in subhorizon OL in coniferous plantations and OH in deciduous ones. The moisture supply in the litter of coniferous stands is greater than in deciduous ones.

High plant species richness indicates management-related disturbances rather than the conservation status of forests

There is a wealth of smaller-scale studies on the effects of forest management on plant diversity. However, studies comparing plant species diversity in forests with different management types and intensity, extending over different regions and forest stages, and including detailed information on site conditions are missing. We studied vascular plants on 1500 20m×20m forest plots in three regions of Germany (Schwäbische Alb, Hainich-Dün, Schorfheide-Chorin). In all regions, our study plots comprised different management types (unmanaged, selection cutting, deciduous and coniferous age-class forests, which resulted from clear cutting or shelterwood logging), various stand ages, site conditions, and levels of management-related disturbances. We analyzed how overall richness and richness of different plant functional groups (trees, shrubs, herbs, herbaceous species typically growing in forests and herbaceous light-demanding species) responded to the different management types. On average, plant species richness was 13% higher in age-class than in unmanaged forests, and did not differ between deciduous age-class and selection forests. In age-class forests of the Schwäbische Alb and Hainich-Dün, coniferous stands had higher species richness than deciduous stands. Among age-class forests, older stands with large quantities of standing biomass were slightly poorer in shrub and light-demanding herb species than younger stands. Among deciduous forests, the richness of herbaceous forest species was generally lower in unmanaged than in managed forests, and it was even 20% lower in unmanaged than in selection forests in Hainich-Dün. Overall, these findings show that disturbances by management generally increase plant species richness. This suggests that total plant species richness is not suited as an indicator for the conservation status of forests, but rather indicates disturbances.

Puistu esimese rinde võrastiku alguse kõrguse hindamine lennukilidari mõõdistusandmete järgi / A simple model to estimate forest canopy base height from airborne lidar data

Abstract Airborne laser scanner (ALS) measurements from two test sites in Estonia were used to estimate forest canopy-base height (HVL). The ALS data was collected by Estonian Land Board using Leica ALS50-II scanner. The HVL was estimated by using mode value and standard deviation of the ALS pulse reflection position height distribution. The pulse reflections which had height less than 0.5 m over the estimated digital terrain model were excluded from the analysis. In situ measurements of canopy base height (HVA) were carried out in 20 mesotrophic Norway spruce and silver birch forest stands in Järvselja and in 45, mostly Scots pine dominant, mesotrophic forest stands in Aegviidu. Determination coefficients of linear regression between HVL and HVA for both test sites were over 0.8 and the residual standard errors of the models were less than two meters. The influence of forest understory vegetation to the estimation of HVL was tested by excluding the near-to-ground vegetation reflections which had height less than 1.5 m. The test results revealed no significant impact of forest understory to the HVL models. The cross validation showed that the HVL models were independent of test sites and tree species composition. The Järvselja data based HVL model had 1.3 m negative bias if applied to Aegviidu forests and the Aegviidu data based HVL model had 1.4 m positive bias if applied to Järvselja forests. In the Aegviidu test site, difference of HVL models of coniferous and deciduous stands was tested and the difference was found not to be significant

Rapid assessment of wind storm-caused forest damage using satellite images and stand-wise forest inventory data

Abstract: This paper introduces a method for rapid forest damage assessment using satellite images and stand-wise forest inventory data. Two Landsat 5 Thematic Mapper (TM) images from June and September 2010 and data from a forest stand register developed within the frameworks of conventional stand-wise forest inventories in Lithuania were used to assess the forest damage caused by wind storms that occurred on August 8, 2010. Satellite images were geometrically and radiometrically corrected. The percentage of damage in terms of wind-fallen or broken tree volume was then predicted for each forest compartment within the zone potentially affected by the wind storm, using the non-parametric k-nearest neighbor technique. Satellite imagery-based difference images and general forest stand characteristics from the stand register were used as the auxiliary data sets for prediction. All auxiliary data were available from existing databases, and therefore did not involve any added data acquisition costs. Simultaneously, aerial photography of the area damaged by the wind storm was carried-out and color infrared (CIR) orthophotos with a resolution of 0.5 x 0.5 m were produced. A precise manual interpretation of the effects of the wind storm was used to validate satellite image-based estimates. The total wind damaged volume in pine dominating forest (~1.180.000 m3) was underestimated by 2.2%, in predominantly spruce stands (~233.000 m3) by 2.6% and in predominantly deciduous stands (~195.000 m3) by 4.2%, compared to validation data. The overall accuracy of identification of wind-damaged areas was around 95-98%, based solely on difference data from satellite images gathered on two dates.

Edge Effects on Soil Acidification in Forests on Sandy Soils Under High Deposition Load

This study investigated how forest soil acidification is affected by edge proximity. We measured pH(KCl) and exchangeable K, Ca, Mg and Al concentrations of the mineral topsoil (0–30 cm) from the exposed edge to the interior (128 m from the edge) of three deciduous and four coniferous forest stands. From the front edge to the interior of the deciduous stands, the pH(KCl) values decreased at 0–5 cm soil depth (from 3.07 to 2.98) but increased at 5–10 cm (from 3.26 to 3.32) and 10–30 cm (from 3.48 to 3.75) depth. In the coniferous stands, pH(KCl) values declined from edge to interior at all soil depths, i.e. from 3.10 to 2.89, from 3.26 to 3.06 and from 3.54 to 3.31 at 0–5, 5–10 and 10–30 cm, respectively. The concentrations of exchangeable cations decreased from edge to interior, with larger differences in the coniferous (of up to 265 %) than in the deciduous stands (up to 99 %). At forest edges, enhanced soil acidification due to higher potentially acidifying deposition could be counteracted in the upper mineral soil by higher base cation throughfall and litterfall, faster litter decomposition, higher soil organic matter content, lower nitrate leaching from the soil and/or lime fertiliser drift. Nonetheless, deeper in the soil of the deciduous stands, these buffer processes seem unable to counteract soil acidification due to potentially acidifying deposition at the edges. Edge effects on soil acidity are important since they can translate into effects on plant communities, soil biota, nitrogen cycling and carbon sequestration.

Influence of stand composition on predatory mite (Mesostigmata) assemblages from the forest floor in western Canadian boreal mixedwood forests

Mesostigmatan mites dwelling in forest floors are the least abundant of soil mesofauna, however, they play an important role by regulating population sizes of other microarthropods. In this study, we evaluate the impact of stand type on the structure and community composition of predator mites from the Mesostigmata order in forest floors from undisturbed deciduous stands dominated by Populus tremuloides, coniferous stands dominated by Picea glauca, and mixed stands in the western Canadian boreal forest. We observed differences in species richness, dominance and community composition among the stand types. Coniferous and mixed stands had higher mesostigmatan species richness than deciduous stands. Coniferous stands had a different community composition than deciduous and mixed stands, indicating the importance of coniferous trees in structuring mesostigmatan assemblages. Although forest floor pH varied within a narrow range, it was important in explaining variations in mesostigmatid community composition. Mesostigmatan assemblages were also distinguished according to forest floor thickness, probably reflecting species-specific habitat preferences within accumulated litter. Maintaining coniferous trees will be necessary for conserving the natural patterns of mesostigmatan mites in western Canadian boreal mixedwood forests.

Influence of canopy budget model approaches on atmospheric deposition estimates to forests

Accurate quantification of total nitrogen and acidifying deposition is a major source of uncertainty in determining the exceedance of critical loads in forest ecosystems. Monitoring of atmospheric deposition is frequently based on throughfall measurements in combination with the canopy budget model to calculate ion-exchange fluxes between the forest canopy and incident rainfall water. Various approaches for each step in the canopy budget model have been reported and compared, but combinations of different approaches were not yet assessed. Therefore, the present study quantified the range of estimated dry deposition and total deposition resulting from all possible combinations of canopy budget model approaches for three typical case studies: (i) total nitrogen and potentially acidifying deposition onto a forest canopy, (ii) the ratio of these deposition variables between adjacent coniferous and deciduous stands and (iii) the parameters of a deposition time trend analysis. The time step, type of precipitation data and tracer ion used in the model had a significant effect on the findings in the three case studies. In addition, including or excluding canopy leaching of weak acids and canopy uptake of nitrogen during the leafless season largely affected the results, while including or excluding canopy uptake of nitrate generally showed no effect. In general, the use of wet-only precipitation data can be recommended, along with sodium as a tracer ion and the inclusion of weak acids. We conclude that further research should focus on the assumptions of inertness of the tracer ion and the equal deposition efficiency of base cations and the tracer ion and on the quantification of weak acids in rainfall and throughfall water. Since local or tree-species specific effects might influence the results obtained in this study, a similar analysis is recommended for other tree species and regions when using the canopy budget model.

Functional analysis of vegetation on alpine treeline ecotone in the Julian and Kamnik-Savinja Alps in Slovenia

Our study focused on the functional aspects of plant species and vegetation at the transition from larch (Larix decidua Mill.) forest to mountain pine (Pinus mugo Turra) stands on the alpine treeline ecotone. With increasing elevation, living conditions grow harsher, which is reflected in the plant species and functional trait composition of plant communities. At four different localities in the Slovenian Alps, releves along an altitudinal gradient and according to vegetation type were made (European larch forests, larch trees-mountain pine shrubs, mountain pine shrubs), using standard Central European phytocoenological method. In the upper mountain pine belt, few differential species were found, since most species represented in mountain pine stands also occurred in the lower two vegetation belts, while there were many differential species in the lower forest belt. Species with considerable competitive ability and moderate stress tolerance dominated the upper treeline ecotone, whereas ruderality is poorly expressed. The importance of stress tolerance in plant strategies increased slightly in the mountain pine belt. Changes in the representation of some functional traits attributes were detected by vegetation belts, but complete species turnover did not occur. Changes in dominant life form involved greater cover of chamaephytes and nanophanerophytes in the upper mountain pine belt. Species with evergreen leaves dominated mountain pine stands and deciduous forest stands. The share of species with scleromorphic leaves increased in the direction of the mountain pine belt while the share of species with mesomorphic and hygromorphic leaves declined. Mountain pine stands create good conditions for the regeneration of tree species and colonisation by ecologically more demanding forest species while species of alpine grasslands withdraw on open areas. Since today’s treeline is lowered due to past human activity, an upward shift is expected. Also, considering the great importance of competition strategy, current conditions in mountain pine stands are favourable for future forest succession.

Species’ thermal preferences affect forest bird communities along landscape and local scale habitat gradients

The spatial distributions of species, and the resulting composition of local communities, are shaped by a complex interplay between species’ climatic and habitat preferences. We investigated this interaction by analyzing how the climatic niches of bird species within given communities (measured as a community thermal index, CTI) are related to vegetation structure. Using 3129 bird communities from the French Breeding Bird Survey and an information theoretic multimodel inference framework, we assessed patterns of CTI variation along landscape scale gradients of forest cover and configuration. We then tested whether the CTI varies along local scale gradients of forest structure and composition using a detailed data set of 659 communities from six forests located in northwestern France. At landscape scale, CTI values decreased with increasing forest cover, indicating that bird communities were increasingly dominated by cold‐dwelling species. This tendency was strongest at low latitudes and in landscapes dominated by unfragmented forest. At local scale, CTI values were higher in mature deciduous stands than in conifer or early stage deciduous stands, and they decreased consistently with distance from the edge of forest. These trends underpin the assertion that species’ habitat use along forest gradients is linked with their climatic niche, although it remains unclear to what extent it is a direct consequence of microclimatic variation among habitats, or a reflection of macroscale correlations between species’ thermal preferences and their habitat choice. Moreover, our results highlight the need to address issues of scale in determining how habitat and climate interact to drive the spatial distribution of species. This will be a crucial step towards accurate predictions of changes in the composition and dynamics of bird communities under global warming.