Deadwood Volume Research Articles

A Permanent Research Platform for Ecological Studies in Intact Temperate Mountainous Forests from Slătioara UNESCO Site and Its Surroundings, Romania

This paper describes a permanent research platform (PRP) designed and implemented in “Codrul secular Slătioara” and its surroundings (2205.85 ha), having also the role of introductory paper for future research articles based on data collected from this platform. “Codrul secular Slătioara” is known as one of the largest temperate mountainous intact forests of Europe and, in 2017, it was included in UNESCO World Heritage List, as part of the “Ancient and Primeval Beech Forests of the Carpathians and Other Regions of Europe”. Moreover, the PRP overlaps other three scientific reserves, the share of strictly protected forests exceeding 70%. This platform has a multiple role, being developed for research, conservation and educational activities. The PRP was designed for an ecological analysis of the intact forest ecosystems. It contains 193 circular sample plots, each of them of 500 m2, and it is structured on two levels. The first level contains 58 sample plots corresponding to a square grid of 500 × 500 m, stretching over the entire forested area, and the second level contains 135 plots, placed according to a square grid of 100 × 100 m, covering 136 ha within the core area of the UNESCO site. We measured the characteristics of 8296 living trees, 1743 standing dead trees, 1900 dead wood trunks, 3214 saplings, and the abundance–dominance indices of flora species. Thus, we identified 14 tree species, 17 shrub species, and 248 other cormophyte species forming the herbaceous layer. In terms of volume, the main tree species are Norway spruce, silver fir and European beech. The tallest species are Norway spruce (56 m) and silver fir (51 m). The average volume of living trees is 659 m3·ha−1, with a maximum of 1441 m3·ha−1. The mean total dead wood volume is about 158 m3·ha−1, with sample plots where the total dead wood volume exceeds 600 m3·ha−1. After presenting the results of preliminary data processing, the paper describes the main research topics to be further considered, based on the PRP, and the foresights related to the PRP’s monitoring and development.

Quality of substrate and forest structure determine macrofungal richness along a gradient of management intensity in beech forests

The richness of saproxylic species is commonly found to be compromised in managed forests. However, it is often less clear if this reflects local scarcity of high quality deadwood objects or changes to the overall forest structure. Further, anthropogenic and natural drivers working at the landscape scale may have a strong impact. Hence, it is difficult to give detailed advice on how to conserve biodiversity in a cost-effective manner. Here we explored macrofungal richness on deadwood across a gradient of forest management intensity in Danish Fagus sylvatica forests. We aimed to disentangle the importance of habitat quality per se from the effects of variables recorded at the forest stand and landscape scales. Data was collected from 40 beech stands, each representing one of four broad management types: conventionally managed, near-to-nature managed, recently unmanaged and long unmanaged. Stands were aggregated within six larger forest clusters, to account for variation at the landscape scale. Fungal species were recorded as fruitbodies on deadwood at five decay stages. We found that the four management types showed strong differences in fungal richness, deadwood volume and forest structure, with long unmanaged stands having notably higher structural complexity and species richness. Fungal species richness and presence of red-listed fungi appeared to be mainly driven by deadwood volume, tree species and decay stage, but some stand-level variables such as canopy gaps, basal area and stand age were also of importance. Lying coarse deadwood had the highest species richness while standing coarse woody debris and fine deadwood had lower species density. Saproxylic fungal richness on individual deadwood objects was largely affected by resource quality and size, which is often limited in managed forests. Our results indicate that the best measure to increase fungal species richness, especially in managed stands, is to increase the amount of large diameter deadwood.

Modeling of Dead Wood Potential Based on Tree Stand Data

Here we present a framework for identifying areas with high dead wood potential (DWP) for conservation planning needs. The amount and quality of dead wood and dying trees are some of the most important factors for biodiversity in forests. As they are easy to recognize on site, it is widely used as a surrogate marker for ecological quality of forests. However, wall-to-wall information on dead wood is rarely available on a large scale as field data collection is expensive and local dead wood conditions change rapidly. Our method is based on the forest growth models in the Motti forest simulator, taking into account 168 combinations of tree species, site types, and vegetation zones as well as recommendations on forest management. Simulated estimates of stand-level dead wood volume and mean diameter at breast height were converted into DWP functions. The accuracy of the method was validated on two sites in southern and northeastern Finland, both consisting of managed and conserved boreal forests. Altogether, 203 field plots were measured for living and dead trees. Data on living trees were inserted into corresponding DWP functions and the resulting DWPs were compared to the measured dead wood volumes. Our results show that DWP modeling is an operable tool, yet the accuracy differs between areas. The DWP performs best in near-pristine southern forests known for their exceptionally good quality areas. In northeastern areas with a history of softer management, the differences between near-pristine and managed forests is not as clear. While accurate wall-to-wall dead wood inventory is not available, we recommend using DWP method together with other spatial datasets when assessing biodiversity values of forests.

Driving factors of conifer regeneration dynamics in eastern Canadian boreal old-growth forests.

Old-growth forests play a major role in conserving biodiversity, protecting water resources, and sequestrating carbon, as well as serving as indispensable resources for indigenous societies. Novel silvicultural practices must be developed to emulate the natural dynamics and structural attributes of old-growth forests and preserve the ecosystem services provided by these boreal ecosystems. The success of these forest management strategies depends on developing an accurate understanding of natural regeneration dynamics. Our goal was therefore to identify the main patterns and drivers involved in the regeneration dynamics of old-growth forests with a focus on boreal stands dominated by black spruce (Picea mariana (L.) Mill.) and balsam fir (Abies balsamea (L.) Mill.) in eastern Canada. We sampled 71 stands in a 2 200 km2 study area located within Quebec's boreal region. For each stand, we noted tree regeneration (seedlings and saplings), structural attributes (diameter distribution, deadwood volume, etc.), and abiotic (slope and soil) factors. The presence of seed-trees located nearby and slopes having moderate to high angles most influenced balsam fir regeneration. In contrast, the indirect indices of recent secondary disturbances (e.g., insect outbreaks or windthrows) and topographic constraints (slope and drainage) most influenced black spruce regeneration. We propose that black spruce regeneration dynamics can be separated into distinct phases: (i) layering within the understory, (ii) seedling growth when gaps open in the canopy, (iii) gradual canopy closure, and (iv) production of new layers once the canopy is closed. These dynamics are not observed in paludified stands or stands where balsam fir is more competitive than black spruce. Overall, this research helps explain the complexity of old-growth forest dynamics, where many ecological factors interact at multiple temporal and spatial scales. This study also improves our understanding of ecological processes within primary old-growth forests and identifies the key factors to consider when ensuring the sustainable management of old-growth boreal stands.

Ecological assessment of dead wood volume in natural deciduous forests in the Vita river valley in Holosiivskyi National Nature Park

The volume of dead wood by fractions, components, species composition and decay stages in natural lime-ash-oak forest in Vita river valley within the protected zone of the Holosiivskyi National Nature Park in northern part of forest-steppe of Ukraine is studied and presented. The study was conducted on permanent sample plot by identifying and measuring components of standing dead wood (standing dead trees, snags) and lying (fallen dead trees, logs and tree branches) dead wood. Total volume of deadwood is 94.2 m3 ·ha–1, and consists of standing dead wood (23.9 m3 ·ha–1) and lying deadwood (70.3 m3 ·ha–1). Common ash (Fraxinus excelsior L.) has the highest proportion of total dead wood volume – 43.2% (40.7 m3 ·ha–1) followed by common oak (Quercus robur L.) with 32.0% (30.1 m3 ·ha–1). Hornbeam (Carpinus betulus L.) and linden (Tilia cordata Mill.) have 10.1 and 8.8 % respectively of total dead wood volume. The relative sharing of wych elm (Ulmus glabra Huds.) and black alder (Alnus glutinosa (L.) Gaerth.) is low. Common ash has the highest proportion of standing dead wood volume – 59.4%, the proportion of hornbeam is much lower – 23.8%. Standing dead wood mainly belongs to decay stage I (81.6%). The volume of lying dead wood consists mainly of common oak (40.8%) and common ash (37.7%). In general lying dead wood belongs to decay stages I–V, although decay stages III and IV dominate. Among tree species only common ash has lying dead wood of all five decay stages, although dead wood of III and IV decay stages dominates with proportions 35.2% and 29.3% respectively. Lying dead wood of other tree species belongs to one-three decay stages with domination one decay stage. Dead wood displayed a wide range of variation in fractions and components, tree species and decay stages may have an important role in creating a high diversity of habitats and substrates for wildlife organisms in studied forest ecosystems

Volume techniques for estimating standing and lying dead wood in Okomu national park, Edo state, Nigeria

Dimensions are essential characteristics to estimate standing and lying dead wood volumes. Therefore, the study is aimed at estimating standing and lying dead wood volumes with a view to identify the variation that exist amongst volume equations in Okomu National Park. Hossfeld’s equation and other volume equations should be considered suitable for estimating volume of standing and lying deadwood, depending on the available variable.

Stand characteristics and dead wood in urban forests: Potential biodiversity hotspots in managed boreal landscapes

Urban forests are usually not intensively managed and may provide suitable environments for species threatened by production forestry. Thus, urban forests could have the potential of enhancing biodiversity both within cities and at a larger landscape scale. In this study, we investigated stand structures of boreal urban forests to assess them in terms of naturalness and biodiversity conservation potential. We sampled two types of urban spruce-dominated stands: random urban stands as representatives of average urban forests, and valuable urban stands known to host high polypore richness and assumed to represent urban biodiversity hotspots. Urban forests were compared to rural forests with different levels of naturalness. Living and dead trees and cut stumps were measured from all studied stands. Urban forests had generally diverse living tree structures with abundant large-diameter trees. Random urban forests had more dead wood (median 10.1 m3 ha−1) than production forests (2.7 m3 ha−1) but still considerably less than protected, former production forests (53.9 m3 ha−1) or semi-natural forests (115.6 m3 ha−1). On the other hand, valuable urban forests had relatively high median volume of dead wood (88.2 m3 ha−1). We conclude that the combination of diverse stand composition and the presence of old-growth characteristics in boreal urban forests form a strong baseline from which their biodiversity value can be further developed, e.g. by leaving more fallen or cut trees to form dead wood. We propose that urban forests could become significant habitats for biodiversity conservation in the future.

Carbon Stock in Deadwood: The Mountain Birch (Betula pubescens subsp. czerepanovii) Forests in the Khibiny Mountains (Russia)

ABSTRACT Deadwood is an important component for conserving carbon stock (C-stock) and maintaining species diversity in forests. In the last years, some studies emphasized that the estimation of deadwood C-stock is a crucial aspect to understand the role of boreal forests as potential carbon sink. The aim of the present study is to provide new data about deadwood volume and C-stock in the mountain birch forests – pure mountain birch stands and mixed stands – of the Kola Peninsula (Russia). In 28 sampling plots randomly located in the Khibiny Mountains, quantitative and qualitative data on deadwood (logs, snags, stumps) were collected and processed. The results show on average 126 logs, 127 snags, and 225 stumps per hectare of mountain birch forest. The average deadwood volume is 23.9 m3 ha−1, while the deadwood C-stock in the mountain birch forests of the Khibiny Mountains is estimated at 3.10 Mg ha−1. The pure mountain birch stands are characterized by an average deadwood volume of 18.4 m3 ha−1, while the mixed stands have an average deadwood volume of 28.6 m3 ha−1. These results highlight an important contribution of deadwood of the boreal forests as carbon storage to mitigate climate change.

Deadwood Characteristics in Mature and Old-Growth Birch Stands and Their Implications for Carbon Storage

As one of the most abundant tree species in the hemiboreal zone, birch is important from both commercial and biodiversity perspectives. While old-growth deciduous stands are important for biodiversity conservation with an emphasis on deadwood availability, the role that deadwood in these stands plays in carbon sequestration remains unclear. We studied mature (71–110 years old) and old-growth (121–150 years old) birch stands on fertile mineral soils. The marginal mean deadwood volume was 43.5 ± 6.4 m3 ha−1 in all mature stands, 51.3 ± 7.1 m3 ha−1 in recently unmanaged mature stands, and 54.4 ± 4.4 m3 ha−1 in old-growth stands; the marginal mean deadwood carbon pool for each stand type was 5.4 ± 0.8 t·ha−1, 6.3 ± 0.9 t·ha−1, and 7.9 ± 0.6 t·ha−1, respectively. Deadwood volume was not related to stand productivity in terms of stand basal area, stand height, or stand age. The difference between mature and old-growth stands remained non-significant (p < 0.05). A high volume of deadwood was almost continuously present throughout the landscape in assessed unmanaged sites; moreover, 88% of sample plots in old-growth stands and 63% of sample plots in mature stands had a deadwood volume higher than 20 m3·ha−1. Old-growth stands had a slightly greater volume of large deadwood than unmanaged mature stands; in both, almost half of the deadwood was more than 30 cm in diameter and approximately one-fifth had a diameter greater than 40 cm. Both groups of stands had similar proportions of coniferous and deciduous deadwood and lying and standing deadwood. Old-growth stands had a higher volume of recently and weakly decayed wood, indicating increased dieback during recent years.

Variation in Deadwood Microsites in Areas Designated under the Habitats Directive (Natura 2000)

The continuing decline in biodiversity presents a major environmental protection challenge. The conservation of sufficiently extensive and diverse habitats requires an array of coordinated actions, often involving large areas. While a set of conservation objectives have been defined for the Natura 2000 network, no universal methods of accomplishing them have been specified, and so they must be designed by individual Member States. Deadwood volume and the density of large deadwood pieces are widely used for evaluating the quality of forest habitat types designated under the Habitats Directive. In the present study, data from 5557 sample plots were used to evaluate the mean values of the two deadwood indicators as well as the ratio of deadwood volume to living tree volume for each of the 13 habitat types in Poland. In addition, a logistic regression model was constructed to evaluate the effects of terrain, site, and tree stand characteristics as well as protection type on deadwood volume in Natura 2000 areas. Mean deadwood volume varied greatly between habitat types, with the lowest values found for Central European lichen Scots pine forests (91T0–2.5 m3 ha−1) and Old acidophilous oak woods (9190–4.4 m3 ha−1), and the highest for Riparian mixed forests (91F0–43.1 m3 ha−1) and Acidophilous Picea forests of the montane to alpine levels (9410–55.4 m3 ha−1). The ratio of deadwood volume to living tree volume ranged from approx. 1%–17%. Additionally, the presence of large deadwood differed among habitat types: in some, there were no deadwood pieces with a diameter of ≥50 cm, while their maximum density was 6.1 pieces ha−1. The logistic regression model showed that the likelihood of a habitat type to have a ‘favorable conservation status’ as defined by deadwood abundance (a threshold of at least 20 m3 ha−1 according to Polish manuals on habitat type evaluation) increased with sample plot elevation, site fertility, and moisture, as well as stand age and volume. Positive effects were also observed for forests under strict and active protection versus managed forests. Planned efforts are necessary to enhance the quality of habitats with insufficient deadwood, especially in managed forests. Special attention should be given to areas that are readily accessible due to gentle terrain and low site moisture. Furthermore, younger stands on less fertile sites may require intervention to promote deadwood accumulation. We recommend retaining a certain proportion of mature stands until natural death and decomposition. Increasing the density of large deadwood is currently one of the most pressing conservation needs in most habitat types.

Near-natural forests harbor richer saproxylic beetle communities than those in intensively managed forests

Globally, boreal forests cover 33 percent of Earth's forested area, constitute the largest terrestrial carbon source, and house an important biodiversity of forest-dependent species, many of which are saproxylic taxa reliant on a wide variety of dead wood. This dead wood is most prevalent in old-growth forest, yet intensive management means that little old growth is left, and the remaining forests rarely attain the necessary age and structure to resemble these original forests. There is increasing recognition, however, that forests that have not been subjected to modern, stand-replacing harvest methods (like clear-cutting), and in which management activities are no longer occurring, may be able to provide habitat similar to old-growth forest for many species (older mean age, higher deadwood volumes, and more age-class heterogeneity). These are called ‘near-natural’ forests.We use linear mixed models and redundancy analysis (RDA) to compare beetle species richness and community composition using a large, mixed data set, containing more than one third of all beetle species known from Norway, from window trap records in three types of forest: intensively managed mature stands, recent clear-cuts, and ‘near-natural forests’.Even though location, study substrate and other sampling details vary substantially between sites, the signal from management still comes out as a significant factor explaining the patterns on beetle diversity. We find that species richness of saproxylic, natural-forest indicator, and red-listed beetle species is higher in both near-natural forests and recent clear-cuts than in managed mature forests. Community composition also differs significantly between the three forest types, and these differences are correlated with forest age and volume at the landscape (1 km radius) level.These findings provide support for the idea that near-natural forests provide habitats that are distinct from those in managed forests and which are important for species of conservation concern. Clear-cuts briefly harbor some of these same species, probably due to the sun-exposed coarse woody debris (CWD) that is available immediately after felling, but this effect is transient. Conservation of existing near-natural forests, many of which are currently without legal protection, should therefore be prioritized.

Rocky pine forests in the High Coast Region in Sweden: structure, dynamics and history

Almost all forests in Sweden are managed and only a small fraction are considered natural. One exception is low productive forests where, due to their limited economical value, natural dynamics still dominate. One example is the Scots pine (Pinus sylvestris L.) forests occurring on rocky and nutrient-poor hilltops. Although these forests represent a regionally common forest type with a high degree of naturalness, their dynamics, structure and history are poorly known. We investigated the structure, human impact and fire history in eight rocky pine forests in the High Coast Area in eastern Sweden, initially identified as good representatives of this forest type. This was done by sampling and measuring tree sizes, -ages, fire-scarred trees, as well as dead wood volumes and quality along three transects at each site. The structure was diverse with a sparse layer of trees (basal area 9 m2 and 640 trees larger than 10 cm ha-1) in various sizes and ages; 13 trees ha-1 were more than 300 years old. Dead wood (DW), snags and logs in all stages of decay, was present and although the actual DW (pine) volume (4.4 m3 ha-1) and number of units (53 ha-1) was low, the DW share of total wood volume was 18% on average. Dead wood can be present for several centuries after death; we found examples of both snags and logs that had been dead more than 300 years. Frequent fires have occurred, with an average cycle of 40 years between fires. Most fires occurred between 1500-1900 and many of them (13) during the 1600s. However, fires were probably small since most fire years were only represented at one site and often only in one or a few samples. The rocky pine forests in the High Coast Area are representative of undisturbed forests with low human impact, exhibiting old-growth characteristics and are valuable habitats for organisms connected to sun-exposed DW. Management of protected rocky pine forests may well include small-scale restoration fires and the limited DW volumes should be protected.

The importance of being random! Taking full account of random effects in nonlinear sigmoid hierarchical Bayesian models reveals the relationship between deadwood and the species richness of saproxylic beetles

Hierarchical models are used to study the relationship between a response variable and a predictor in structured data. Random effects are meant to capture the structured part of variability among groups of observations. In ecology, random effects are usually incorporated into the intercept. Their application to the other parameters of the curve, especially in nonlinear curves, has been understudied. However, applying random effects to different parameters of the function is of interest, as it allows us to account for variations in the shape of the relationship over groups of observations.Our study was based on Bayesian models linking the local quantity of deadwood to the local species richness of saproxylic beetles in French forests. Our hypothesis was that it was important to account for inter-forest variations of the relationship to better fit the data. Since a sigmoidal curve seemed adapted to studying this relationship from an ecological point of view, we paid special attention to commonly used sigmoidal functions, but also included two new ones for biogeography originating from ecophysiology (one sigmoid with estimated asymptotes and one with estimated asymptotes allowing asymmetry). We applied various settings of random effects to these different mean functions. We compared, evaluated and interpreted the models and results according to several criteria (WAIC, comparison of significance of the difference in terms of LOOic, goodness-of-fit p-values and magnitude of the effect).We first found that models without random effects were systematically the worst and that the best model was not necessarily the one with random effect incorporated into the intercept, as is usually done in ecology. Secondly, we found that, in most cases, for a given mean function, the best model had several random effects, and the model with the most random effects performed nearly as well as the best models. Furthermore, the inclusion of random effects revealed statistically significant relationships between deadwood volume and species richness. Thirdly, we revealed a complementarity between the different assessment criteria, each one giving important information for the selection and interpretation of the models. In conclusion, future forest biodiversity management studies should incorporate random effects into the modeling framework so that more robust conclusions can be made about the relationships, based on complementary post-fitting analysis criteria.

Patterns and drivers of deadwood volume and composition in different forest types of the Austrian natural forest reserves

Deadwood is an important structural feature in forests providing essential resources for various organisms. Both forest management and conservation are increasingly focusing on the integration of deadwood into forest management. Deadwood volume and composition are driven by forest type, stand age, natural tree mortality, tree species composition and harvesting intensity. Here, we used inventory data from 28 unmanaged natural forest reserves (NFR) in Austria to analyse the patterns and drivers of: (i) the volume of standing and lying deadwood, (ii) the diversity of deadwood in different forest types. Eight forest types are located in the investigated NFRs covering a wide range of vegetation types with altitudes of 140–1825 m asl.The volumes of living wood and deadwood differed markedly between forest types. The average deadwood volume per forest type ranged from 23 m3 ha−1 in larch forests to 109 m3 ha−1 in spruce-fir-beech forests. Likewise, deadwood diversity (species diversity as well as diversity in diameter classes and degree of decomposition) differed significantly among forest types, with the highest deadwood diversity found in beech and spruce forests and the lowest in hardwood floodplain forests, carbonate pine forests and larch forests.Our results show that volume and composition of deadwood vary greatly among different forest types. Regression models revealed that the availability of deadwood was mainly driven by tree- and stand related factors (e.g. forest type, diameter at breast height and volume of living stand), whereas site-related (e.g. altitude) and climatic factors (e.g. mean annual precipitation) had a minor influence. The variables tree species diversity, aspect and slope showed no significance and were therefore not integrated into the final model.This study provides insights into deadwood availability and diversity in NFRs on a national scale, providing reference data for unmanaged temperate forests and aiding decision-making in nature conservation and forest management, since NFRs are reference areas for close-to-nature forestry.

Increasing numbers of wintering forest birds in Swedish Lapland 1986–2017 show stronger correlations with forest development than with local weather

After a long period of decline, the number of forest birds has increased in Sweden in recent decades. Whether this trend is due to an increase in forested area, forest quality, climate change, or a combination of these factors, remains unclear. Here, we compared forest bird data from a local winter point count route around Storuman in Swedish Lapland between 1986 and 2017, with the development of regional forest composition and local weather conditions. We suggest that rather than changes in average annual, winter, or summer local temperatures or precipitation, the main drivers behind increasing numbers of wintering forest birds in this part of Sweden are an increase in the area of denser forest and dead wood volume, and a decrease in open ground area without forest vegetation. While there may be supplementary explanations behind the increasing numbers of forest birds, such as reduced agriculture, decreasing local human population, or stronger photosynthesis, our results indicate that local land use has been favourable for forest birds in recent decades in this area.

Structure and Dynamics of Deadwood in Pine and Oak Stands and their Role in CO2 Sequestration in Lowland Forests of Central Italy

There are many data values describing deadwood in primary forests, however, there is much less information concerning managed forests, particularly in the Mediterranean area. Whole non-living woody biomass (deadwood) is the main component of forest types that plays a vital role in improving and maintaining biodiversity. The dynamics of deadwood and CO2 sequestration by deadwood were examined in three Mediterranean lowland forests: pine forest Pinus pinea L.), evergreen oak forest (Quercus ilex and Quercus suber L.), and deciduous oak forest (Quercus cerris L.). The aim of this study was to carry out a quantitative and qualitative evaluation of the deadwood to gather useful information for the Mediterranean forest management, and to provide some useful data that can be integrated into the methods of estimation for carbon stored in dead components of Mediterranean forest types. The investigations focused on the characterization of the deadwood, to determine which traits are dominant and their potentially functionality within the forest type. Results indicated the deciduous oak type had the highest amount of stand volume (379 m3/ha), deadwood volume (161.8 m3/ha), and C storage (31.43 t/ha). The major component of dead wood was the standing dead trees or snags. There was a higher volume of deadwood in the deciduous oak forest than in the pine and evergreen oak forests. In addition, the deciduous oak forest had a higher snag creation index, a higher fallen log creation index, and a higher past management index compared to pine and evergreen oak forest types. Deadwood volume increased as the decay class in the deciduous oak forest increased, while this trend decreased in the evergreen oak and pine forests. The amount of deadwood was affected by the forest type and forest management regime. Dynamic and past management of deadwood indices indicated that their structure was still in the initial phase of creation and decay in the pine and evergreen oak forests. A comparison with other studies on similar forest types brought out that the variation range of the main parameters for the management of deadwood fell within the variation of the parameters studied. However, the values of these parameters cover a broad range. The population of each forest type is extremely sensitive to the different evolutionary periods of the forest dynamics.

Formerly managed forest reserves complement integrative management for biodiversity conservation in temperate European forests

In Central Europe, the predominant conservation strategy in forests is integrative management, seeking to balance economic interests with conservation goals. This is complemented by unmanaged strict forest reserves, most often small and with a history of forest management. Whether and how such reserves contribute to conservation when the surrounding forest is under progressive integrative management remains unclear.We compared forest structure and biodiversity of several taxa between formerly managed forest reserves and stands under progressive integrative management in beech and beech-oak forests. Alpha diversity was higher in reserves for birds and bats and higher in managed stands for plants and beetles, with no significant differences for fungi. Community composition differed between reserves and managed stands for plants, wood-decomposing fungi, beetles and birds. Reserves had 17 indicator species, including three red-listed species, and managed stands had 34, including one red-listed species. Diversity metrics differed between reserves and managed stands for both beech and beech-oak forests.Our results indicate that progressive integrative management and reserves, even when located in formerly managed stands, are complementary approaches benefitting different taxa and hosting partly different communities. Higher numbers of plants and beetles in managed stands were associated with higher light availability, as reserves in our study were undisturbed mature stands characterized by low light availability and low deadwood volumes. To benefit light and deadwood demanding species, new reserves should include early or late successional stands. If this is not feasible, restoration measures prior to designation or where possible under current protection status should be discussed.

How Well Do Stakeholder-Defined Forest Management Scenarios Balance Economic and Ecological Forest Values?

Research Highlights: We show the difference in the long-term effects on economic and ecological forest values between four forest management scenarios of a large representative forest landscape. The scenarios were largely formulated by stakeholders representing the main views on how to manage north-European forests. Background and Objectives: Views on how to balance forest management between wood production and biodiversity differ widely between different stakeholder groups. We aim to show the long-term consequences of stakeholder-defined management scenarios, in terms of ecological and economic forest values. Materials and Methods: We simulated management scenarios for a forest landscape in Sweden, based on the management objectives and strategies of key stakeholders. We specifically investigated the difference in economic forest values coupled to wood supply and ecological indicators coupled to structural biodiversity between the scenarios over a 100-year period. The indicators were net present value, harvest, growing stock and increment, along with deadwood volume, the density of large trees, area of old forests and mature broadleaf-rich forests. Results: We show that the scenarios have widely different outcomes in terms of the studied indicators, and that differences in indicator outcome were largely due to different distributions in management regimes, i.e., the proportion of forest left unmanaged or under even-aged management or continuous cover forest, as well as specific retention practices. Retention and continuous cover forestry mitigate the negative effects that clear-cut forestry has upon biodiversity. Conclusions: We found that an increase in the forest area under the continuous cover forestry regime could be a cost-efficient way to increase structural diversity in managed boreal forests. On the other hand, no single management regime performed best with respect to all indicators, which means that a mixture of several management regimes is needed to balance conflicting objectives. We also show that the trade-off between economic and ecological indicators was not directly proportional, meaning that an increase in structural biodiversity may be obtained at a proportionally low cost with appropriate management planning.

Dead wood diversity in a Norway spruce forest from the Calimani National Park (Eastern Carpathians)

The role of dead wood in forests encompasses the preservation of biodiversity, the productivity, and the storage of carbon. In this study, we aimed to investigate the variation of the dead wood volume in a subalpine Norway spruce forests. Using a regular grid, the size of the wood, ordered in a five-class scale for snags and logs and a six-class scale for stumps, was recorded. The relationship between the dead wood volume and site respectively stand characteristics by some predictive factors (stand age, altitude, human influence and site productivity) was modelled using mixed linear/nonlinear models. Dead wood volume (74.3 m3·ha-1) accounted for 18.3% of the volume (live trees and dead wood). Regarding the dead wood only, snags amounted to 44.8% of the volume and logs and stumps amounted to 46.5, and 8.7%, respectively. Snags (57.5% of the total) and logs (42.8%), were most common in the 2nd decay class, while stumps were most common in the 6th (51.2%). The distribution of the number of dead wood pieces, in terms of diameter, was obtained according to the Meyer theoretical distribution for snags and to the Gamma theoretical distribution for logs and stumps. This study completes the dead wood database from Norway spruce forests in the subalpine area to find the common features across the Carpathians Mountains. It contributes to the understanding of the dead wood dynamics in a young National Park, considering the year of establishment, consists of forests where forestry work was carried out and stands that were exempt from cutting.

Optimal Continuous Cover Forest Management with a Lower Bound Constraint on Dead Wood

AbstractWe analyze economically optimal continuous cover forestry with dead wood as a biodiversity indicator. We study mixed-species stands consisting of Norway spruce (Picea abies [L.] Karst.), birch (Betula pendula Roth.), and other broadleaves (e.g., oak [Quercus sp.], maple [Acer sp.]). The analysis is based on an economic description of continuous cover forest management using an empirically estimated size-structured transition matrix model. We use size-specific decomposition rates for dead wood, with the lower limit on total dead wood volume varying between 0 and 40 m3 ha–1. The optimization problem is solved in its general dynamic form using gradient-based interior point methods. Increasing the dead wood volume requirement affects total stand density only slightly, but increases stand heterogeneity as other broadleaves are grown in higher numbers. In addition, increasing the dead wood requirement has only a minor effect on the total felled volume, but harvests shift from timber harvests to biodiversity fellings to maintain the required dead wood volume. In the optimal steady state with a high dead wood requirement, two harvesting cohorts emerge: one for timber harvests and the other for biodiversity fellings. Increasing the dead wood requirement decreases steady-state net timber income by up to 30 percent compared to the unconstrained solution.