We have used a global time-dependent magnetohydrodynamic (MHD) simulation of the magnetosphere and particle tracing calculations to determine the access of solar wind ions to the magnetosphere and the access of ionospheric O + ions to the storm-time near-Earth plasma sheet and ring current during the September 24–25, 1998 magnetic storm. We found that both sources have access to the plasma sheet and ring current throughout the initial phase of the storm. Notably, the dawnside magnetosphere is magnetically open to the solar wind, allowing solar wind H + ions direct access to the near-Earth plasma sheet and ring current. The supply of O + ions from the dayside cusp to the plasma sheet varies because of changes in the solar wind dynamic pressure and in the interplanetary magnetic field (IMF). Most significantly, ionospheric O + from the dayside cusp loses access to the plasma sheet and ring current soon after the southward turning of the IMF, but recovers after the reconfiguration of the magnetosphere following the passage of the magnetic cloud. On average, during the first 3 h after the sudden storm commencement (SSC), the number density of solar wind H + ions is a factor of 2–5 larger than the number density of ionospheric O + ions in the plasma sheet and ring current. However, by 04:00 UT, ∼4 h after the SSC, O + becomes the dominant species in the ring current and carries more energy density than H + ions in both the plasma sheet and ring current.