The 145th symposium of the Wenner-Gren Foundation took place June 1–8, 2012, in Haringe Slott near Stockholm, Sweden. The primary goal of the symposium was to reframe discussions of behavioral evolution among Neanderthals and early modern humans. We hoped to replace conventions of a single scale of evolutionary progress (in which the primary benchmark is “modern human behavior”) with a more Darwinian framework that could allow for independent evolutionary trajectories in different areas. The 15 participants included archaeologists researching material culture and subsistence in Eurasia, Africa, and China; physical anthropologists; a demographer; a geneticist; modelers of cultural evolution; and a climatologist. Participants were asked to draw on evidence in their areas of expertise, focusing on evolutionary trends in both modal tendencies and levels of variation/diversity within various regions during the interval in which the Middle Stone Age and Middle Paleolithic developed, spread, and eventually disap...