Cretaceous Record Research Articles

New materials of lithostrotian titanosaurs (Dinosauria: Sauropoda) from the Upper Cretaceous of central Patagonia

Continental Cretaceous deposits exposed in central Patagonia, Argentina, preserve a rich and important record of titanosaurian evolution that spans the entire Upper Cretaceous. Recently, a new lithostragraphic unit, the Lago Colhué Huapi Formation (Coniacian – Maastrichtian), was located in the Golfo San Jorge Basin. Here, we describe new titanosaurian sauropod material from that formation (UNPSJB-Pv 1051). The material consists of a partially articulated left hind limb and disarticulated but associated skeletal elements. They are confidently referred to Titanosauria, and within that clade, to Lithostrotia, probably occupying a derived position. We interpret the bone concentration as an assemblage of hydraulic origin deposited in a fluvial channel. This new material begins the incipient fossil record of the Lago Colhué Huapi Formation, thereby also increasing the titanosaurian diversity of the latest Cretaceous. Additionally, the new titanosaurian enhances evidence in support of strong faunal similarities among the Allen and Marilia formations, chronological equivalents in the Cretaceous of South America. Furthermore, the Lago Colhué Huapi specimen adds to the Late Cretaceous record of Titanosauria and augments our knowledge of central Patagonian terrestrial vertebrate assemblages during this interval.

Facies characterization and sequential evolution of an ancient offshore dunefield in a semi-enclosed sea: Neuquén Basin, Argentina

This study analyses a 30-m-thick, sand-dominated succession intercalated between offshore mudstones in the Lower Cretaceous record of the Neuquen Basin, Argentina, defining facies associated with unidirectional currents as sand dunes (simple and compound), rippled sand sheets and heterolithic sheets. These facies associations are related to the development of an offshore, forward-accreting dunefield developed as a response to the onset of a tidal-transport system. The reported stratigraphic record results from the combination of the gradual downcurrent decrease of the current speed together with the long-term climbing of the entire system. Maximum amplification of the tidal effect associated with incoming oceanic tides to this epicontinental sea would develop at the time of more efficient connection between the basin and the open ocean. Thus, the onset of the offshore tidal system approximately corresponds to the time of maximum flooding conditions (or immediately after). The short-term evolution of the tidal-transport system is more complex and characterized by the vertical stacking of small-scale cycles defined by the alternation of episodes of construction and destruction of the dunefield. The development of these cycles could be the response to changes in tidal current speed and transport capacity.

A new turtle confirms the presence of Bothremydidae (Pleurodira) in the Cenozoic of Europe and expands the biostratigraphic range of Foxemydina.

Pleurodira is a clade of Gondwanan turtles that reached Europe at the beginning of the Late Cretaceous. It is recognized as the most abundant and diverse group of freshwater turtles in the uppermost Cretaceous record of this continent, being represented by several members of Bothremydidae. Two well-preserved and relatively complete skulls are studied in this paper. They come from lower Eocene levels of the French locality of Saint-Papoul (Aude). These specimens are recognized as attributable to a new taxon, Tartaruscola teodorii gen. et sp. nov., identified as a bothremydid. This new form constitutes the only known unambiguous and valid representative of Bothremydidae in the Cenozoic of Europe. The new taxon is diagnosed by several autapomorphies and also by an exclusive combination of characters. It is one of the few members of Bothremydini identified in the Cenozoic record and the only one described for the Eocene outside of the Ouled Abdoun Basin of Morocco. Tartaruscola teodorii is identified as a member of Foxemydina, a clade recognized here as exclusive of the European record, and whose presence in post-Mesozoic levels is demonstrated in this paper. The European Foxemydina T. teodorii and the African Bothremydina Bothremys kellyi, both from the Ypresian, are the youngest members of Bothremydini hitherto known.

Upper-crustal cooling of the Wrangellia composite terrane in the northern St. Elias Mountains, western Canada

This study presents the long-term exhumation history of the Wrangellia composite terrane of the remote and ice-covered northern St. Elias Mountains in southwest Yukon, northwest British Columbia, and adjacent Alaska. Detrital zircon and apatite fission-track age distributions are presented from 21 glacial catchments. The detrital sampling approach allows for a large spatial coverage (∼30,000 km 2 ) and access to material eroded beneath the ice. An additional five bedrock samples were dated by zircon fission-track analysis for a comparison with detrital results. Our new thermochronology data record the Late Jurassic–mid-Cretaceous accretion of the Wrangellia composite terrane to the former North American margin and magmatism, which reset the older thermal record. The good preservation of the Jurassic–Cretaceous record suggests that Cenozoic erosion must have been limited overall. Nonetheless, Eocene spreading-ridge subduction and Oligocene–Neogene cooling in response to the ongoing Yakutat flat-slab subduction are evident in the study area despite its inboard position from the active plate boundary. The results further indicate an area of rapid exhumation at the northern end of the Fairweather fault ca. 10–5 Ma; this area is bounded by discrete, unmapped structures. The area of rapid exhumation shifted southwest toward the plate boundary and the center of the St. Elias syntaxis after 5 Ma. Integrating the new data with published detrital thermochronology from the southern St. Elias Mountains reveals an evolving concentration of deformation and exhumation, possibly within a large-scale, transpressional structure providing important constraints for geodynamic models of syntaxes.

New record of ornithopod dinosaur from the Plottier Formation (Upper Cretaceous), Patagonia, Argentina

Ornithopods are the least known dinosaurs within the Upper Cretaceous record of Argentina. For this reason every new record is very important to know their evolution in South America. Here, we describe a new remain of an indeterminate ornithopod recovered in the Petrobrasaurus quarry of the Puesto Hernández area, northeastern Neuquén province (Argentina), late Coniacian–early Santonian in age. MAU-Pv-PH-458 is the northernmost bone record of an ornithopod in Argentina. This is a fragmentary neural arch from the middle section of the dorsal series of similar size to Macrogryphosaurus gondwanicus. MAU-Pv-PH-458 has typical ornithopod characters such as a lateromedial narrow neural spine and transverse processes dorsoposteriorly to posteriorly oriented. It shares with Macrogryphosaurus the presence of a deep channel between the bases of the postzygapophysis, which is a continuation of the channel that separates the postzygapophyses in posterior view. MAU-Pv-PH-458 increases the ornithopod record from the Plottier Formation.

Magnetostratigraphy of Upper Cretaceous strata in Grand Staircase-Escalante National Monument, southern Utah: The Santonian–Campanian Stage boundary, reassessment of the C33N/C33R magnetochron boundary, and implications for regional sedimentation patterns within the Sevier Foreland Basin

Samples were collected for magnetostratigraphic analysis from the Upper Cretaceous Straight Cliffs and Wahweap formations within Grand Staircase-Escalante National Monument, southern Utah, in an attempt to locate the C34n–C33r and C33r–C33n boundaries; the former approximates the Santonian–Campanian Stage boundary. Locating both of these horizons within the Monument provides for refined temporal resolution of the many new fossil localities discovered in these units, while also adding to our understanding of the overall Straight Cliffs–Wahweap sequence stratigraphic architecture. Results indicate that the John Henry and Drip Tank members of the Straight Cliffs Formation are of normal polarity, and represent the later part of C34n, the Cretaceous Normal Polarity Super-Chron. The C34n–C33r boundary apparently resides in a significant hiatus determined to occur between the Drip Tank Member of the Straight Cliffs Formation and the lower part of the overlying Wahweap Formation. In stratigraphically ascending order, the informally named lower, middle, and most of the upper members of the Wahweap Formation are of reversed polarity. The C33r–C33n polarity change occurs in the upper part of the upper member, and C33n continues stratigraphically upward through the capping sand member of the Wahweap Formation and into the overlying Kaiparowits Formation. The presence of over 200 m of reversely magnetized strata above a level in the lower part of the middle member dated to 79.9 ± 0.3 Ma, which coincides with the GTS2012 date for the C33r–C33n boundary of 79.9 Ma, suggests that the published age of that boundary may be in error. A new date of approximately 78.91 Ma is proposed, in turn allowing for the thick sequence of reversely magnetized strata above the 79.9 Ma level. Age revision of the Kaiparowits Basin's Late Cretaceous record demonstrates that during late Santonian through early middle Campanian time, except in central Utah, a close correlation in time and facies successions with other successions throughout the Western Interior Basin suggests a eustatic influence on the depositional patterns. Starting in the late middle Campanian, the Kaiparowits record, along with those both to the north and south, appear to diverge in style and facies successions. We consider this to have resulted from a fundamental change in depositional controls triggered by the Laramide Orogeny. The anomalous central Utah record appears to have been profoundly influenced by a massive salient (Nebo-Charleston thrust system) in the Sevier Fold and Thrust belt that was active for much of the Late Cretaceous.

Why so many dipnoans? A multidisciplinary approach on the Lower Cretaceous lungfish record from Tunisia

The Lower Cretaceous record of vertebrates from Africa is problematic as the majority of fossil localities lack adequate stratigraphic and paleoecological data when compared with coeval Laurasian deposits. Thereby, our comprehension of paleocommunities and paleobiogeographic patterns may be affected by the lack of multidisciplinary approach. Among taxonomically and paleoecological significant clades, lungfishes (Sarcopterygii, Dipnoi) are commonly found in the Cretaceous fresh water, brackish and marginal-marine deposits of Gondwana, although identifiable elements are limited to isolated tooth plates. We provide the first taxonomic identification of dipnoans from the Ain el Guettar Formation of southern Tunisia (Oum ed Diab member, Albian). Identification of tooth plates based on morphological parameters and phylogenetic analyses indicates the co-occurrence in a discrete stratigraphic unit of at least five lineages referable to Equinoxiodus, Neoceratodus, Asiatoceratodus and/or Ferganoceratodus, Ceratodus, and Lavocatodus. This unusually high diversity is unparalleled in the fossil record and is also challenged by an actualistic comparison with extant taxa. We suggest that a series of taphonomic factors significantly inflated observed lungfish diversity in the estuarine and marginal-marine deposits of the Oum ed Diab member. Therefore, we recognize the fossil fauna as representative of a larger, inland paleo-hydrographic system. This study confirms the paleoecological scenario resulted from the analyses on terrestrial reptiles from the Oum ed Diab member.

Burmese amber fossils bridge the gap in the Cretaceous record of polypod ferns

Cretaceous fossils of derived leptosporangiate ferns (Polypodiales) are exceedingly rare yet they are needed to confirm the hypothesis of a Cretaceous diversification of polypod ferns as predicted by DNA-based divergence time estimates. Here we show that diverse polypod ferns existed in the mid-Cretaceous woodlands of Myanmar. We describe isolated sporangia with a vertical, broken annulus ring containing a differentiated stomium as well as distinguished pinnae fragments. Krameropteris resinatus gen. et sp. nov. is described based on a pinnae-fragment with free branched veins, exindusiate sori and polypod sporangia with trilete spores, and assigned to the early diverging polypod fern lineage Dennstaedtiaceae. Integration of the new fossil evidence in a phylogenetic framework provides support to the hypothesis of a Cretaceous diversification of polypod ferns.

Late Cretaceous (Campanian) actinopterygian fishes from the Kristianstad Basin of southern Sweden

Abstract Although a diverse range of aquatic vertebrates are documented from the Upper Cretaceous (mid-Campanian) marine strata of the Kristianstad Basin in southern Sweden, only chondrichthyans and marine amniotes have been described in detail to date. In contrast, coeval actinopterygians are virtually unreported, yet their remains are extremely abundant at most sampled localities. A comprehensive assessment of these fossils has identified the first Late Cretaceous actinopterygian fauna from the Fennoscandian Shield, incorporating indeterminate lepisosteids, the durophagous pycnodontid Anomoeodus subclavatus , the predatory pachycormid Protosphyraena sp., a large ichthyodectid, pachyrhizodontids resembling Pachyrhizodus , the enchodontid Enchodus cf. gladiolus and indeterminate small teleosts. These taxa are diagnosed mainly from isolated teeth and scales, implying substantial taphonomic loss prior to burial. Moreover, the prolific recovery of actinopterygian skeletal remnants in recent excavations suggests that historical collecting biases, rather than ecological paucity, have contributed to their under-representation in the Swedish Cretaceous record. Palaeobiogeographically, the Kristianstad Basin actinopterygians show compositional resemblance to assemblages from the Northern European Platform and the Western Interior Seaway of North America, advocating distributional communication across the Boreal proto-Atlantic Ocean.

Inflorescences of Mauldinia sp. (Lauraceae) and associated fruits from the Cenomanian of Languedoc Roussillon, France

Inflorescences of Mauldinia (Lauraceae) are described from the middle Cenomanian Pauletian facies of Gard, Languedoc-Roussillon, south-eastern France. This is the first record of the genus from this part of Europe. Mauldinia sp. is described based on peduncles and lateral units. Isolated, slender peduncles show spirally arranged scars or rarely attached lateral units. Isolated lateral units consist of cladode-like structures showing up to seven flower/fruit scars. Some isolated fruits were found associated with peduncles and lateral units. Fruits are ovoid fruits contain a single, elongated seed. The pericarp consists of epicarp, mesocarp and endocarp. The seed consists of two cotyledons containing well-developed reserve cells. Mauldinia sp. from Gard differs from known Mauldinia species by having longer internodes along the peduncle and by the shape of the lateral units. Sedimentological and palaeontological evidence suggests that Mauldinia sp. from Gard grew in a protected, littoral environment with limited marine influence. This report provides new information to the relatively poorly known Cretaceous record of angiosperms in France. The occurrence of Mauldinia in the lower Upper Cretaceous of Gard corroborates the widespread occurrence of the genus in marginal littoral environments during the mid-Cretaceous.

A new teiid lizard from the Late Cretaceous of the Haţeg Basin, Romania and its phylogenetic and palaeobiogeographical relationships

A new lizard genus and species is described based on a three-dimensionally preserved partial skull and associated lower jaws from the Pui Islaz locality (Late Cretaceous, early Maastrichtian) in the Haţeg Basin, western Romania. Barbatteius vremiri gen. et sp. nov. is diagnosed by a unique combination of symplesiomorphies and synapomorphies. A nested set of synapomorphies support assigning Barbatteius to Teiidae as the first unambiguous Late Cretaceous record of this family from Laurasia. Barbatteius differs from other teiids by having more extensive osteodermal sculpture on the skull roof and suspensorium, and by a pentagonal occipital osteoscute exhibiting more or less parallel lateral margins. Barbatteius is a large-bodied lizard, estimated to be up to 800 mm in total length. It has weakly heterodont dentition, but without enlarged posterior crushing teeth, suggesting that it fed on arthropods, small vertebrates and plants. The mix of taxa with affinities to Euramerica (paramacellodid and borioteiioid lizards) and Gondwana (madtsoiid snakes and the teiid Barbatteius) currently known for the Maastrichtian squamate assemblage from Haţeg Basin supports the growing realization that ‘Haţeg Island’ has a complex palaeobiogeographical history.http://zoobank.org/urn:lsid:zoobank.org:pub:75C2D80F-8DDB-4FB2-9844-1552D626F63D

Island life in the Cretaceous - faunal composition, biogeography, evolution, and extinction of land-living vertebrates on the Late Cretaceous European archipelago.

The Late Cretaceous was a time of tremendous global change, as the final stages of the Age of Dinosaurs were shaped by climate and sea level fluctuations and witness to marked paleogeographic and faunal changes, before the end-Cretaceous bolide impact. The terrestrial fossil record of Late Cretaceous Europe is becoming increasingly better understood, based largely on intensive fieldwork over the past two decades, promising new insights into latest Cretaceous faunal evolution. We review the terrestrial Late Cretaceous record from Europe and discuss its importance for understanding the paleogeography, ecology, evolution, and extinction of land-dwelling vertebrates. We review the major Late Cretaceous faunas from Austria, Hungary, France, Spain, Portugal, and Romania, as well as more fragmentary records from elsewhere in Europe. We discuss the paleogeographic background and history of assembly of these faunas, and argue that they are comprised of an endemic ‘core’ supplemented with various immigration waves. These faunas lived on an island archipelago, and we describe how this insular setting led to ecological peculiarities such as low diversity, a preponderance of primitive taxa, and marked changes in morphology (particularly body size dwarfing). We conclude by discussing the importance of the European record in understanding the end-Cretaceous extinction and show that there is no clear evidence that dinosaurs or other groups were undergoing long-term declines in Europe prior to the bolide impact.

The origin and early evolution of metatherian mammals: the Cretaceous record.

Metatherians, which comprise marsupials and their closest fossil relatives, were one of the most dominant clades of mammals during the Cretaceous and are the most diverse clade of living mammals after Placentalia. Our understanding of this group has increased greatly over the past 20 years, with the discovery of new specimens and the application of new analytical tools. Here we provide a review of the phylogenetic relationships of metatherians with respect to other mammals, discuss the taxonomic definition and diagnosis of Metatheria, outline the Cretaceous history of major metatherian clades, describe the paleobiology, biogeography, and macroevolution of Cretaceous metatherians, and provide a physical and climatic background of Cretaceous metatherian faunas. Metatherians are a clade of boreosphendian mammals that must have originated by the Late Jurassic, but the first unequivocal metatherian fossil is from the Early Cretaceous of Asia. Metatherians have the distinctive tightly interlocking occlusal molar pattern of tribosphenic mammals, but differ from Eutheria in their dental formula and tooth replacement pattern, which may be related to the metatherian reproductive process which includes an extended period of lactation followed by birth of extremely altricial young. Metatherians were widespread over Laurasia during the Cretaceous, with members present in Asia, Europe, and North America by the early Late Cretaceous. In particular, they were taxonomically and morphologically diverse and relatively abundant in the Late Cretaceous of western North America, where they have been used to examine patterns of biogeography, macroevolution, diversification, and extinction through the Late Cretaceous and across the Cretaceous-Paleogene (K-Pg) boundary. Metatherian diversification patterns suggest that they were not strongly affected by a Cretaceous Terrestrial Revolution, but they clearly underwent a severe extinction across the K-Pg boundary.

Sauropod and Small Theropod Tracks from the Lower Jurassic Ziliujing Formation of Zigong City, Sichuan, China, with an Overview of Triassic–Jurassic Dinosaur Fossils and Footprints of the Sichuan Basin

A dinosaur footprint assemblage from the Lower Jurassic Ziliujing Formation of Zigong City, Sichuan, China, comprises about 300 tracks of small tridactyl theropods and large sauropods preserved as concave epireliefs (natural molds). The theropod footprints show similarities with both the ichnogenera Grallator and Jialingpus. Three different morphotypes are present, probably related to different substrate conditions and extramorphological variation. A peculiar preservational feature in a morphotype that reflects a gracile trackmaker with extremely slender digits, is the presence of a convex epirelief that occurs at the bottom of the concave digit impressions. It is possibly the result of sediment compaction underweight load when the pes penetrated the substrate, being a resistant residue during exhumation and weathering. The sauropod tracks belong to a trackway with eight imprints consisting of poorly preserved pes and manus tracks and a better preserved set, probably all undertracks. The narrow-gauge trackway pattern resembles the ichnogenus Parabrontopodus well known from the Jurassic but other features such as the minor heteropody are different. The assemblage enriches the dinosaur record from the famous Zigong locality and the evidence from the Lower Jurassic in this area that was restricted to a few skeletal remains and footprints. Furthermore it proves the presence of small theropods, whereas skeletons of the group, well- known from the Middle-Upper Jurassic of Zigong, are of medium to large size only. Remarkable is the dominance of saurischians in these assemblages, which is characteristic of Jurassic dinosaur communities whereas the Cretaceous record shows an increase of ornithopod groups. An overview of the dinosaur trace and body fossil record of the Sichuan Basin supports this view. The paleoenvironment can be designated as a low-latitude tropical freshwater lake as it is indicated by bivalve shells.

A million-year-scale astronomical control on Late Cretaceous sea-level

An evaluation of the global synchronicity and duration of rd sea-level fluctuations during the Cretaceous greenhouse has been hampered by poor constraints on potential climatic and tectonic drivers, and limitations of geochronology and chronostratigraphic correlation. To provide insight into the na- ture of such sea-level fluctuations, here we present a new Late Cretaceous record from the Jordanian Levant Platform, comprising a detailed physical-, bio-, chemo- and sequence stratigraphy. Carbonate content of these strata reflects overall sequence stratigraphic development, and demonstrates a dramatic 3 rd -order-scale cycle that is also apparent in the δ 13 C record. Updated radioisotopic constraints and astrochronologic testing pro- vide support for the inference of an ~ 1 million year long sea-level oscillation associated with this 3 rd -order cycle, which likely reflects a long-period obliquity (1.2 Myr) control on eustasy and stratigraphic sequence development, linked to the global carbon cycle. The observation of cyclic sea-level fluctuations on this time scale suggests sustained global modulation of continental fresh-water-storage. The hypothesized link between astronomical forcing and sea-level forms a baseline approach in the global correlation of sequence bound- aries.

Eodortoka morellana gen. et sp. nov., the first pan-pleurodiran turtle (Dortokidae) defined in the Lower Cretaceous of Europe

Dortokidae is a relatively poorly-known clade of Pan-Pleurodira, exclusively recognized in Europe. It is the only clade of Laurasiatic pan-pleurodirans known in the post-Jurassic record. Its presence has been recognized in several locations and ages. However, only two nominated valid taxa have been described so far: the western European Upper Cretaceous Dortoka vasconica, and the Romanian late Paleocene Ronella botanica. A new dortokid is studied here, Eodortoka morellana gen. et sp. nov. This new taxon is the oldest nominated dortokid, coming from the lower Aptian (Lower Cretaceous) of the Spanish town of Morella. It shows several primitive characters not shared with the so far described dortokids, such as the presence of mesoplastra and the absence of overlap of the second pair of pleural scutes onto the first pair of costal plates. The comparison of Eodortoka morellana gen. et sp. nov. with other specimens allows us to recognize the presence of two members of Dortokidae in the Iberian Lower Cretaceous record. This study shed light on the early evolution of Dortokidae, as well as on the systematic and paleobiogeography of this clade.

Late Cretaceous record of large soft-bodied coleoids based on lower jaw remains from Hokkaido, Japan

Late Cretaceous record of large soft-bodied coleoids based on lower jaw remains from Hokkaido, Japan

Late Cretaceous Record of Large Soft-Bodied Coleoids Based on Lower Jaw Remains from Hokkaido, Japan

The origin and phylogenetic relationships of most modern coleoid groups have not yet been explained by reliable fossil evidence, in large part because of the reduction or disappearance of a calcified chambered shell during their evolutionary history. Herein we describe two exceptionally large coleoid lower jaws from the Upper Cretaceous strata in Hokkaido, Japan. On the basis of the comparison of gross morphology and morphometric data of the lower jaws of modern and fossil coleoids, we assigned the two lower jaws to the following new taxa: Nanaimoteuthis hikidai sp. nov. of the order Vampyromorpha (superorder Octobrachia) and Haboroteuthis poseidon gen. et sp. nov. of the order Teuthida (superorder Decabrachia). The lower jaw of N. hikidai is distinguished from other species of the same genus from the Upper Cretaceous of Vancouver Island (Canada) and Hokkaido by having a broader, more anteriorly curved hood of the outer lamella. The lower jaw of H. poseidon seemingly exhibits mosaic features like those of...

A new sphenodontian (Lepidosauria: Rhynchocephalia) from the Late Triassic of Argentina and the early origin of the herbivore opisthodontians.

Sphenodontians were a successful group of rhynchocephalian reptiles that dominated the fossil record of Lepidosauria during the Triassic and Jurassic. Although evidence of extinction is seen at the end of the Laurasian Early Cretaceous, they appeared to remain numerically abundant in South America until the end of the period. Most of the known Late Cretaceous record in South America is composed of opisthodontians, the herbivorous branch of Sphenodontia, whose oldest members were until recently reported to be from the Kimmeridgian-Tithonian (Late Jurassic). Here, we report a new sphenodontian, Sphenotitan leyesi gen. et sp. nov., collected from the Upper Triassic Quebrada del Barro Formation of northwestern Argentina. Phylogenetic analysis identifies Sphenotitan as a basal member of Opisthodontia, extending the known record of opisthodontians and the origin of herbivory in this group by 50 Myr.

Additions to the sauropod dinosaur fauna of the Cenomanian (early Late Cretaceous) Kem Kem beds of Morocco: Palaeobiogeographical implications of the mid-Cretaceous African sauropod fossil record

Determining the relationships of mid-Late Cretaceous African taxa is central to understanding the timing and resultant palaeobiogeographical patterns of Gondwanan fragmentation. The early Late Cretaceous (Cenomanian) Kem Kem beds of southeastern Morocco preserve a diverse vertebrate fauna, including sauropod dinosaurs. Sauropod material includes the holotype of the rebbachisaurid diplodocoid Rebbachisaurus garasbae and fragmentary remains representing indeterminate titanosauriforms and rebbachisaurids. Here, we describe two new specimens from the Kem Kem beds. A dorsal neural arch with complex internal pneumaticity is tentatively attributed to a somphospondylan titanosauriform. A caudal vertebra possessing several rebbachisaurid synapomorphies is excavated by a large and pervasive lateral pneumatic foramen, a feature undocumented in other rebbachisaurids. However, caudal vertebrae are currently unknown for the sympatric R. garasbae, so this element could be referable to that taxon or a second, previously unknown, rebbachisaurid species. Interestingly, this new caudal vertebra displays a mosaic of features otherwise restricted to limaysaurine or nigersaurine rebbachisaurids, suggesting a placement basal to these clades, which is the position usually recovered for R. garasbae in phylogenetic analyses. A review of the mid-Cretaceous African sauropod fossil record removes the Cretaceous record of dicraeosaurids from Africa, restricting this clade to a single post-Jurassic occurrence in Argentina. All diagnostic sauropod remains can be attributed to titanosauriforms or rebbachisaurids. Whereas rebbachisaurids were seemingly restricted to northwestern Africa and disappeared post-Cenomanian, titanosauriforms were widespread across the African continent and survived until the latest Cretaceous. The development of the mid-Cretaceous Trans-Saharan Seaway might have acted as a dispersal barrier for rebbachisaurids and other vertebrate groups. In contrast, titanosauriforms might have been able to cross this barrier, but it is possible that they were also unable to disperse, and that northwestern African titanosauriforms were not closely related to taxa from the rest of the African continent. New materials and a better understanding of titanosaur interrelationships will be crucial in teasing these scenarios apart.