ABSTRACT Seven portolan charts and atlases (including the anonymous Rex Tholomeus chart) made between 1311 and 1538 were cartometrically analysed. The findings indicate that their geometry was not improved chronologically and that the accuracy of Pietro Vesconte's charts (1311–13) was never surpassed. Portolan charts and atlases appear to be mosaics composed of subsections, each with variable metrics and accuracy levels that are, on average, twice as great in comparison to their Mediterranean and Black Sea areas (as a whole), and it is highly unlikely that they are genuine late medieval products, which is also supported by historical evidence. Their anticlockwise tilts fit well with the tilt of the Gibraltar-Antioch line on modern maps that closely approximate Ptolemy I and Ptolemy II projections centred near Gibraltar and with their meridian tilts for areas around Genoa. If late medieval Italian cartographers had acquired a collection of maps or charts from before the Middle Ages, it is theoretically feasible that they could have achieved the typical geometry of portolan charts by graphically combining their manuscript copies of large-scale regional maps in Mercator or Mercator-like projection, using a small-scale map in conic or pseudoconic projection (on which the convergence of meridians is displayed) as a template.