Continental Landmass Research Articles

Three new species of Amphipoda (Crustacea: Peracarida) from the remote oceanic island of Trindade, off the coast of south-eastern Brazil

ABSTRACT Trindade is a volcanic remote island isolated some 1200 km from the nearest continental landmass. While knowledge of the island’s diversity of crustacean decapod fauna has been notably broadened by extensive investigations, knowledge about its amphipod fauna is still poor. During a 12-year survey (2012–2024) a wealth of amphipod material was obtained in Trindade from various sublittoral habitats at water depths of less than 30 m. Here, we report on three new species of the genera Colomastix Grube, 1861, Pariphinotus Kunkel, 1910 and Eusiroides Stebbing, 1888. Additionally, identification keys to the world species of Eusiroides and Pariphinotus, and to the Brazilian species of Colomastix are provided. http://www.zoobank.org/urn:lsid:zoobank.org:pub:B17266F0-B78E-4768-8FB2-D91BA5F748CA

A distinctive rare earth element signature for pyrite oxidation and glacial weathering

The application of rare earth elements (REE) and neodymium (Nd) isotopes to authigenic Fe-(oxyhydr)oxide phases leached from marine sediments can be used for reconstructing past ocean circulation and chemical weathering patterns on nearby continental landmasses. To explore the behaviour of REE during chemical weathering in glacial environments, we analyzed the authigenic fraction of glacimarine sediments from the continental shelf off northern Svalbard (Arctic Ocean), where the evolution of ice sheet dynamics since the last deglaciation is well constrained. Our results show that leached authigenic fractions in Svalbard sediments exhibit shale-normalized REE patterns characterized by anomalously high mid-REE enrichment relative to both light- and heavy-REE, as expressed by concavity index values (CI > 2.5) that significantly depart from typical REE signatures for leached fractions in marine and river sediments worldwide. Using a compilation of literature data, we provide compelling evidence that the occurrence of pronounced mid-REE enrichment in authigenic fractions of Svalbard marine sediments links to terrestrial oxidation of pyrite following glacial weathering. While future investigation will be required to further understand the detailed mechanism accounting for the observed REE decoupling and its link to pyrite oxidation, we propose that preferential dissolution of mid-REE-enriched rock-bearing minerals such as apatite following glacial erosion, sulfide weathering and subsequent release of sulfuric acid could release a distinctive REE signature in glacial surface environments. Our findings suggest that the use of authigenic REE in the sedimentary record could provide a means for tracing glacial weathering and associated biogeochemical sulfur and iron cycling across geological times.

Subaerial weathering drove stabilization of continents.

Earth's silica-rich continental crust is unique among the terrestrial planets and is critical for planetary habitability. Cratons represent the most imperishable continental fragments and form about 50% of the continental crust of the Earth, yet the mechanisms responsible for craton stabilization remain enigmatic1. Large tracts of strongly differentiated crust formed between 3 and 2.5 billion years ago, during the late Mesoarchaean and Neoarchaean time periods2. This crust contains abundant granitoid rocks with elevated concentrations of U, Th and K; the formation of these igneous rocks represents the final stage of stabilization of the continental crust2,3. Here, we show that subaerial weathering, triggered by the emergence of continental landmasses above sealevel, facilitated intracrustal melting and the generation of peraluminous granitoid magmas. This resulted in reorganization of the compositional architecture of continental crust in the Neoarchaean period. Subaerial weathering concentrated heat-producing elements into terrigenous sediments that were incorporated into the deep crust, where they drove crustal melting and the chemical stratification required to stabilize thecratonic lithosphere. The chain of causality between subaerial weathering and the final differentiationof Earth's crust implies that craton stabilization was an inevitable consequence of continental emergence. Generation of sedimentary rocks enriched in heat-producing elements, at a time in the history of the Earth when the rate of radiogenic heat production was on average twice the present-day rate, resolves a long-standing question of why many cratons were stabilized in the Neoarchaean period.

Osteology, phylogenetic affinities, and palaeobiogeographic significance of the bizarre ornithischian dinosaur Ajkaceratops kozmai from the Late Cretaceous European archipelago

Abstract At the climax of their evolutionary history in the latest Cretaceous, ceratopsian dinosaurs were among the most dominant components of North American and Asian land ecosystems. In other continental landmasses, however, ceratopsians were extraordinarily rare and the affinities of their proposed representatives often turned out to be inconclusive. Arguably the most significant evidence of Ceratopsia from outside North America and Asia is represented by Ajkaceratops kozmai from the Santonian (Upper Cretaceous) of Hungary. We provide a detailed osteological description of Ajkaceratops and highlight its bizarre anatomy. Ajkaceratops has been ‘traditionally’ interpreted to represent a Bagaceratops-like coronosaur, and its occurrence on the European islands was hypothesized to probably result from an early Late Cretaceous dispersal event from Asia. However, while the snout of Ajkaceratops may resemble that of some ceratopsians, closer inspection of the preserved elements indicates that these similarities are largely superficial. While it cannot be ruled out that Ajkaceratops represents a highly peculiar member of the clade, its placement is far from certain. Still, the discovery of Ajkaceratops exemplifies the importance and uniqueness of European dinosaur faunas.

Characteristics, distribution patterns and sources of atmospheric microplastics in the Bohai and Yellow Seas, China

Although the prevalence of microplastics in the atmosphere has recently received considerable attention, there is little information available regarding the distribution of atmospheric microplastics over oceanic regions. In this study, during the summer and autumn months of 2022, we investigated atmospheric microplastics in four marine regions off the eastern coast of mainland China, namely, the southern, middle, and northern regions of the Yellow Sea, and the Bohai Sea. The abundance of atmospheric microplastics in these regions ranged from 1.65 to 16.80 items/100 m3 during summer and from 0.38 to 14.58 items/100 m3 during autumn, although we detected no significant differences in abundance among these regions. Polyamide, chlorinated polyethylene, and polyethylene terephthalate were identified as the main types of plastic polymer. On the basis of meteorological data and backward trajectory model analyses, we established that the atmospheric microplastics detected during summer were mainly derived from the adjacent marine atmosphere and that over the continental landmass in the vicinity of the sampling area, whereas microplastics detected during autumn appear to have originated mainly from the northeast of China. By influencing the settlement and migration of microplastics, meteorological factors, such as relative humidity and wind speed, were identified as potential factors determining the distribution and characteristics of the detected microplastics. Our findings in this study, revealing the origin and fate of marine atmospheric microplastics, make an important contribution to our current understanding of the distribution and transmission of microplastics within the surveyed region and potentially worldwide.

Neoarchean magmatism and Rhyacian reworking in the northeast of the Amapá Block, southeastern Guiana Shield, Brazil: Geochemistry, U–Pb geochronology, and Hf–Nd isotopes

This study investigates the age, geochemical nature, sources, and Rhyacian reworking of the Neoarchean magmatism at the northeastern portion of the Amapá Block, southeastern Guiana Shield (northeast Amazonian Craton). This block consists of a large Archean continental landmass strongly reworked during the Transamazonian orogeny (2.26–1.95 Ga). We provide the very first geochemical results for Archean basement units of the Amapá Block, which are the Mesoarchean Tumucumaque and Neoarchean Guianense complexes, and Neoarchean Mungubas Granite. The first two exhibited geochemical affinity with enriched TTG-type and hybrid granitoids, respectively, while the Mungubas Granite showed affinity with biotite granites. Such affinity with C-type granitoids indicates that the genesis of these units involves partial melting of crustal sources. LA-MC-ICP-MS zircon U–Pb and Lu–Hf, and whole-rock Sm–Nd data were obtained for the Neoarchean Mungubas Granite and Guianense Complex. In both units, zircon U–Pb dating provided a distribution of dates that span ∼50 Ma forming two groups with distinct mean 207Pb/206Pb ages of ∼2.70 Ga and ∼2.65 Ga, respectively. The older age is considered as the crystallization/emplacement age of the Mungubas Granite and the Guianense orthogneiss protolith, while the younger one is ascribed to concealed Pb loss. The recognition of Mesoarchean inherited zircon core and xenocryst with ages of 2.84 Ga and 2.99 Ga suggest that these units reworked older basement rocks from the Amapá Block. Subchondritic ԐHf(2.70 Ga) of zircon and ԐNd(2.70 Ga) of whole-rock, along with their respective Hf-TDMC (3.6–3.2 Ga) and Nd–TDM (3.6 Ga) model ages, for both units, reflect the substantial crustal reworking of predominant Paleoarchean sources during the Neoarchean in the northeastern Amapá Block. In addition, U–Pb dating on titanite crystals from the Guianense orthogneiss yielded an unprecedented ∼2.05 Ga age, arguing that a Late Rhyacian metamorphic event affected the Archean orthogneisses. This 2.05 Ga event coincides with the emplacement of Rhyacian granitoids in the area, as well as with the 2.06–2.04 Ga late orogenic HT granulite facies and amphibolite facies metamorphic events identified, respectively, at the northern border and in the central-southern portion of the Amapá Block.

Persistent warm and cold spells in the Northern Hemisphere extratropics: regionalisation, synoptic-scale dynamics and temperature budget

Abstract. Persistent warm and cold spells are often high-impact events that may lead to significant increases in mortality and crop damage and can put substantial pressure on the power grid. Taking their spatial dependence into account is critical to understand the associated risks, whether in present-day or future climates. Here, we present a novel regionalisation approach of 3-week warm and cold spells in winter and summer across the Northern Hemisphere extratropics based on the association of the warm and cold spells with large-scale circulation. We identify spatially coherent but not necessarily connected regions where spells tend to co-occur over 3-week timescales and are associated with similar large-scale circulation patterns. We discuss the physical drivers responsible for persistent extreme temperature anomalies. Cold spells systematically result from northerly cold advection, whereas warm spells are caused by either adiabatic warming (in summer) or warm advection (in winter). We also discuss some key mechanisms contributing to the persistence of temperature extremes. Blocks are important upper-level features associated with such events – co-localised blocks for persistent summer warm spells in the northern latitudes; downstream blocks for winter cold spells in the eastern edges of continental landmasses; and upstream blocks for winter cold spells in Europe, northwestern North America and east Asia. Recurrent Rossby wave patterns are also relevant for cold and warm spell persistence in many mid-latitude regions, in particular in central and southern Europe. Additionally, summer warm spells are often accompanied by negative precipitation anomalies that likely play an important role through land–atmosphere feedbacks.

Cenozoic stable isotope constraints on the Eurasian continental interior hydroclimate response to high CO2

The response of the terrestrial hydrologic cycle to higher atmospheric CO2 remains poorly constrained, largely due to difficulty in predicting how land surface processes may modify individual hydroclimate parameters such as precipitation (P), evapotranspiration (ET), and runoff (q). To interrogate how the terrestrial hydrologic cycle may change with warming and higher CO2, we utilize the Cenozoic geologic record of terrestrial stable oxygen isotopes (δ18O) as recorded in authigenic minerals, which reflect precipitation δ18O (δ18Op). Values of δ18Op are sensitive to changes in terrestrial hydroclimate, including to the ratio of P/ET and to precipitable water. In short, decreasing P/ET or higher precipitable water produces shallower δ18Op gradients inland, whereas increasing P/ET or lower precipitable water results in steeper δ18Op continental gradients. We compile nearly 15,000 samples of authigenic carbonate and tooth enamel across Eurasia—the largest continental landmass—that span the Cenozoic Era to reconstruct the past spatial distribution and zonal (i.e. east-to-west) gradients of δ18Op. We find that, in epochs with higher CO2, zonal δ18Op gradients increase, suggesting an increase in P/ET across Eurasia despite the expected increase in precipitable water in greenhouse climates. We compare these results to δ18Op gradients simulated by an isotope-enabled climate model (iCESM) forced with both pre-industrial (PI) and 4xPI CO2 mixing ratios. Simulated δ18Op gradients shallow as CO2 rises, in contrast to the reconstructed δ18Op gradients. This data-model mismatch suggests either that Earth system feedbacks not represented in our iCESM simulations, such as changes in Northern Hemisphere ice extent and vegetation, may have fundamentally changed Northern Hemisphere hydroclimate or that land-surface processes under high CO2 climates may be mis-represented in iCESM.

Environmental evolution of western Tierra del Fuego (∼54°S) since ice-free conditions and its zonal/hemispheric implications

By virtue of its position adjacent to the Drake Passage, Tierra del Fuego in South America allows examining the vegetation and environmental history of the southernmost continental landmass outside Antarctica, and the evolution of the Southern Westerly Winds-Southern Ocean (SWW–SO) coupled system since the Last Glacial Maximum (LGM). For that purpose, we studied sediment cores from Lago Charquito, a small closed-basin lake in central-west Tierra del Fuego with a continuous lacustrine record since ∼17.3 ka. Ice-free conditions at the site imply a ∼70 km retreat of the Bahía Inútil glacier lobe from its LGM position during a ∼800-year interval, a trend that continued until its disappearance ∼100 km upstream from L. Charquito, ∼800 years later. Our palynological data show an open landscape dominated by cold-tolerant shrubs and herbs between ∼17.3–12.9 ka, with increases in precipitation of SWW origin at ∼16.3 ka, ∼14.7 ka, between ∼8.7–7.6 ka, and after ∼6.8 ka. Warming at ∼12.9 ka initiated an abrupt afforestation trend that stalled during the early Holocene (∼12–8.7 ka) owing to a precipitation decline and wildfires, and later resumed in response to invigorated SWW. We hypothesize that sparse Nothofagus tree populations inhabited the periphery of the Patagonian Ice Sheet (PIS) during the LGM and migrated toward the Andes contemporaneous with glacier recession as temperature rose during the Last Glacial Termination (T1). We posit that besides establishing topographic and climatic barriers for land biota, the PIS enabled the connectivity of cold-tolerant hygrophilous plant populations along a humid fringe adjacent to its land-based perimeter, despite the presumably dry conditions downwind from the eastern PIS margin. Our results suggest that southward shifts or expansion of the SWW toward or beyond Tierra del Fuego enhanced upwelling and ventilation of deep waters in the SO, northward shifts or weakening had the opposite effect. We observe that the time evolution of atmospheric CO2 concentrations, high-latitude air and sea-surface temperatures, and sea level during T1 fall short in explaining the timing and abruptness of the Bahía Inútil glacier lobe collapse, and quite possibly multiple other glacier lobes from the PIS.

Diversification dynamics of a common deep-sea octocoral family linked to the Paleocene-Eocene thermal maximum

The deep-sea has experienced dramatic changes in physical and chemical variables in the geological past. However, little is known about how deep-sea species richness responded to such changes over time and space. Here, we studied the diversification dynamics of one of the most diverse octocorallian families inhabiting deep sea benthonic environments worldwide and sustaining highly diverse ecosystems, Primnoidae. A newly dated species-level phylogeny was constructed to infer their ancestral geographic locations and dispersal rates initially. Then, we tested whether their global and regional (the Southern Ocean) diversification dynamics were mediated by dispersal rate and abiotic factors as changes in ocean geochemistry. Finally, we tested whether primnoids showed changes in speciation and extinction at discrete time points. Our results suggested primnoids likely originated in the southwestern Pacific Ocean during the Lower Cretaceous ∼112 Ma, with further dispersal after the physical separation of continental landmasses along the late Mesozoic and Cenozoic. Only the speciation rate of the Southern Ocean primnoids showed a significant correlation to ocean chemistry. Moreover, the Paleocene-Eocene thermal maximum marked a significant increase in the diversification of primnoids at global and regional scales. Our results provide new perspectives on the macroevolutionary and biogeographic patterns of an ecologically important benthic organism typically found in deep-sea environments.

Constraining the Nineteenth-Century Temperature Baseline for Global Warming

Abstract Since the Paris Agreement, climate policy has focused on 1.5° and 2°C maximum global warming targets. However, the agreement lacks a formal definition of the nineteenth-century “pre-industrial” temperature baseline for these targets. If global warming is estimated with respect to the 1850–1900 mean, as in the latest IPCC reports, uncertainty in early instrumental temperatures affects the quantification of total warming. Here, we analyze gridded datasets of instrumental observations together with large-scale climate reconstructions from tree rings to evaluate nineteenth-century baseline temperatures. From 1851 to 1900 warm season temperatures of the Northern Hemisphere extratropical landmasses were 0.20°C cooler than the twentieth-century mean, with a range of 0.14°–0.26°C among three instrumental datasets. At the same time, proxy-based temperature reconstructions show on average 0.39°C colder conditions with a range of 0.19°–0.55°C among six records. We show that anomalously low reconstructed temperatures at high latitudes are underrepresented in the instrumental fields, likely due to the lack of station records in these remote regions. The nineteenth-century offset between warmer instrumental and colder reconstructed temperatures is reduced by one-third if spatial coverage is reduced to those grid cells that overlap between the different temperature fields. The instrumental dataset from Berkeley Earth shows the smallest offset to the reconstructions indicating that additional stations included in this product, due to more liberal data selection, lead to cooler baseline temperatures. The limited early instrumental records and comparison with reconstructions suggest an overestimation of nineteenth-century temperatures, which in turn further reduces the probability of achieving the Paris targets. Significance Statement The warming targets formulated in the Paris Agreement use a “pre-industrial” temperature baseline that is affected by significant uncertainty in the instrumental temperature record. During the second half of the nineteenth century, much of the continental landmasses were not yet covered by the observational station network and existing records were often subject to inhomogeneities and biases, thus resulting in uncertainty regarding the large-scale mean temperature estimate. By analyzing summer temperature reconstructions from tree-rings for the Northern Hemisphere extratropical land areas, we examine an independent climate archive with a typically broader and more continuous spatial extent during the “pre-industrial” period. Despite the additional uncertainty when using climate reconstructions instead of direct observations, there is evidence for an overestimation of land temperature during the summer season in early instrumental data. Colder early instrumental temperatures would reduce the probability of reaching the Paris targets.

Climatic data comparison between mid-latitude Azores (North Atlantic) and Tristan da Cunha (South Atlantic) islands

"The Azores islands in the North Atlantic and the Tristan da Cunha islands in the South Atlantic are approximately located at the same latitude, stretching between 36 and 41 degrees North and South respectively. Nevertheless, there are important climatic differences between the two archipelagos. The average annual temperature is higher in the Azores by more than 3 degrees, while the amount of precipitation is higher in Tristan da Cunha by about 600 mm/year on average. The warmer climate of the Azores may be explained by several contributing factors, such as: the smaller distance to the nearest continental landmass – less than 1500 km, compared to more than 2700 km in the case of Tristan da Cunha; the land mass of the islands themselves, as the Azores are much larger; the location of the Azores in the area of the North Atlantic High, with more stable weather and a higher amount of sunshine hours compared to the Tristan da Cunha islands, where westerlies prevail; and, not in the least, the influence of the Gulf Stream in the case of the Azores. The larger amount of precipitation in the case of Tristan da Cunha islands is due to their location on the southern flank of the South Atlantic High, which causes a higher degree of cloudiness and therefore more precipitation. The altitude is also a factor, as both archipelagos feature volcanic mountains rising above 2000 metres, providing a barrier for the air masses and determining orographic precipitation, especially on the windward side of the islands. The climate has had an impact on the settlement and human life on the islands. The Azores were settled immediately after their discovery in the 15th century and there is a rich social and economic life ever since. By contrast, the small community on Tristan da Cunha main island was formed only in the 19th century and has endured many hardships. Keywords: climatic comparison, Azores, Tristan da Cunha, average temperature, precipitation."

Fall migration, oceanic movement, and site residency patterns of eastern red bats (Lasiurus borealis) on the mid-Atlantic Coast

Along the mid-Atlantic coast of the United States, eastern red bats (Lasiurus borealis) are present during fall mating and migration, though little is currently known about most aspects of bat migration. To reveal migration patterns, and understand drivers of over-water flight, we captured and radio-tagged 115 eastern red bats using novel technology, and subsequently tracked and described their movements throughout the region. We compared over-water flight movements to randomly generated patterns using a use-availability framework, and subsequently used a generalized linear mixed effects model to assess the relationship of over-water flight to atmospheric variables. We used hidden Markov models to assess daily activity patterns and site residency. Most bats with long-distance movements traveled in a southwesterly direction, however path vectors were often oriented interior toward the continental landmass rather than along the coastline. We observed that some bats transited wide sections of the Chesapeake and Delaware bays, confirming their ability to travel across large water bodies. This over-water flight typically occurred in the early hours of the night and during favorable flying conditions. If flight over large water bodies is a proxy for over-ocean flight, then collision risk at offshore wind turbines – a major source of migratory bat fatalities – may be linked nightly to warm temperatures that occur early in the fall season. Risk, then, may be somewhat predictable and manageable with mitigation options linking wind-energy operation to weather conditions and seasonality.

Brief history of the island of New Guinea (Papua)

Papua Island is divided into two parts, the eastern part of which is within the continental landmass of Australia, and the western part is located within Indonesia. The two departments are governed in many respects by both. Pacific Islands, and New Guinea.

Geochemistry of granulite facies BIFs from the Luis Alves Craton (Southern Brazil): Implications for Archean-Paleoproterozoic depositional and tectonic settings

Banded Iron Formations (BIF) developed strongly during the Neoarchean – Paleoproterozoic Eras linked with global environmental processes such as the Great Oxidation Event (GOE) and changes in geodynamics, sea level, geobiology, hydrothermal systems, and sediment deposition conditions. BIF development and distribution worldwide are also related to supercontinent cycles, being a powerful tool to understand the evolution of plate tectonics. In this paper, we used field-based geological-structural data, petrography, X-ray diffraction, and lithogeochemical analyses to perform the first characterization of Banded Iron Formations at the Luis Alves Craton (LAC), Southern Brazil. The LAC BIF are classified as oxide and silicate facies, both with granoblastic texture. Oxide facies consists of quartz and magnetite. Besides these minerals, silicate facies contains olivine and ferrosilite, evidencing granulite metamorphism. Based on major elements oxide facies has a hydrothermal signature, with low content of Al2O3 and TiO2, whereas silicate facies register a continental sediment contribution been more enriched in these elements, MgO, MnO, and CaO. Trace-elements also indicate the main source for both silicate and oxide facies, which is seawater with a major influence of low-T hydrothermal sources, besides continental sedimentation sources. The Ce/Ce × anomaly is true negative, representing oxidizing waters, which also may represent a shallow water column depth. The depositional environment is inferred as a shallow, restricted basin. Oxide facies represents sedimentation closer to hydrothermal vents and with weak influence from continental clastic sedimentation, while silicate facies represent sedimentation closer to an adjoining continental landmass. Oxide facies has more Algoma-like characteristics, such as higher Fe and Eu contents, Sm/Yb and Eu/Sm typical of higher hydrothermal fluid inputs, and low siliciclastic contribution. On the other hand, silicate facies resembles Superior-type BIF, with higher siliciclastic contribution, negative Ce/Ce × anomalies, and Eu/Eu* <1.8. In summary, we propose the LAC BIF is representative of a transitional environment from Algoma-to Superior-type. The first depositions of the LAC BIFs may have developed within an Algoma-like environment during the Neoarchean. The waning tectonic and volcanic activity during the Siderian shifted to a platformal environment, with shallower waters that may induce the deposition of carbonates typical of Superior-type deposits. The depositional basin may have been closed by the Rhyacian and those sediments were then pushed down to the lower crust and underwent granulite facies metamorphism.

Continental heat storage: contributions from the ground, inland waters, and permafrost thawing

Abstract. Heat storage within the Earth system is a fundamental metric for understanding climate change. The current energy imbalance at the top of the atmosphere causes changes in energy storage within the ocean, the atmosphere, the cryosphere, and the continental landmasses. After the ocean, heat storage in land is the second largest term of the Earth heat inventory, affecting physical processes relevant to society and ecosystems, such as the stability of the soil carbon pool. Here, we present an update of the continental heat storage, combining for the first time the heat in the land subsurface, inland water bodies, and permafrost thawing. The continental landmasses stored 23.8 ± 2.0 × 1021 J during the period 1960–2020, but the distribution of heat among the three components is not homogeneous. The sensible diffusion of heat through the ground accounts for ∼90 % of the continental heat storage, with inland water bodies and permafrost degradation (i.e. latent heat) accounting for ∼0.7 % and ∼9 % of the continental heat, respectively. Although the inland water bodies and permafrost soils store less heat than the solid ground, we argue that their associated climate phenomena justify their monitoring and inclusion in the Earth heat inventory.

Exploring island syndromes: Variable matrix permeability in Phalaenopsis pulcherrima (Orchidaceae), a specialist lithophyte of tropical Asian inselbergs.

Plants confined to island-like habitats are hypothesised to possess a suite of functional traits that promote on-spot persistence and recruitment, but this may come at the cost of broad-based colonising potential. Ecological functions that define this island syndrome are expected to generate a characteristic genetic signature. Here we examine genetic structuring in the orchid Phalaenopsis pulcherrima, a specialist lithophyte of tropical Asian inselbergs, both at the scale of individual outcrops and across much of its range in Indochina and on Hainan Island, to infer patterns of gene flow in the context of an exploration of island syndrome traits. We sampled 323 individuals occurring in 20 populations on 15 widely scattered inselbergs, and quantified genetic diversity, isolation-by-distance and genetic structuring using 14 microsatellite markers. To incorporate a temporal dimension, we inferred historical demography and estimated direction of gene flow using Bayesian approaches. We uncovered high genotypic diversity, high heterozygosity and low rates of inbreeding, as well as strong evidence for the occurrence of two genetic clusters, one comprising the populations of Hainan Island and the other those of mainland Indochina. Connectivity was greater within, rather than between the two clusters, with the former unequivocally supported as ancestral. Despite a strong capacity for on-spot persistence conferred by clonality, incomplete self-sterility and an ability to utilize multiple magnet species for pollination, our data reveal that P. pulcherrima also possesses traits that promote landscape-scale gene flow, including deceptive pollination and wind-borne seed dispersal, generating an ecological profile that neither fully conforms to, nor fully contradicts, a putative island syndrome. A terrestrial matrix is shown to be significantly more permeable than open water, with the direction of historic gene flow indicating that island populations can serve as refugia for postglacial colonisation of continental landmasses by effective dispersers.

Seismic and ionospheric disturbances caused by the 3 September 2017 underground nuclear test in North Korea

We investigated lithospheric and ionospheric disturbances caused by the 3 September 2017 underground nuclear test (UNT) in North Korea based on data from networks of seismic stations and GPS/GLONASS receivers. We analyzed frequency composition of longitudinal and surface waves detected at more than 40 seismic stations. Low frequencies (∼0.45 ± 0.21 Hz) were found to prevail in the spectrum of longitudinal waves. For both types of waves, frequencies decreased with distance according to the power law. Analysis revealed two trends in the spatial distribution of peak frequencies. First, high and mid-frequencies (0.14–0.50 Hz) are typical for the continental landmass regions, and low frequencies of surface waves (0.13 Hz and below) are more common for the regions adjacent to fringe seas. Second, uneven frequency variations of seismic waves for different azimuths relative to the UNT epicenter (frequencies subside rapidly in the east, southeast, and southwest directions from the epicenter; towards the continental landmass inner parts, frequency variation is much slower). Records of longitudinal waves made at different distances from the epicenter allowed estimating the size of the focal area. Analysis of data from GPS/GLONASS receiving stations near the Korean Peninsula revealed ionospheric disturbances that were most likely caused by UNT. The ionospheric disturbances started to be detected ∼ 8 min after UNT and were observed for about 5 hrs. During the first 1.5–2 hrs after UNT, travelling ionospheric disturbances (TID) were recorded; they propagated from the epicenter at ∼ 600, 250 and 133 m/s. These TIDs had periods of 1–10.5 min and could be associated with acoustic waves induced in the Earth's atmosphere by the underground nuclear test. After TID passage over the UNT site, there was a long-lived (more than 3.5 hrs) region of non-travelling perturbations of ionospheric plasma with the velocity about 7 m/s. This velocity describes not so much the disturbance travel, but the time when the “receiver – satellite” line of sight (LOS) crosses the disturbance. This region can possibly occur due to the formation of standing waves in the atmosphere, development of plasma instabilities or penetration into the ionosphere of an anomalous electric field generated by radioactive substances leaked out onto the surface.

A new subspecies of Papilio saharae Oberthür, 1879 (Lepidoptera: Papilionidae) from Lampedusa, Italy

An investigation into the only representative of the genus Papilio on the island of Lampedusa, the largest island of the Pelagian group, was conducted as part of a broader Mediterranean-wide study on the machaon complex. Over a three-year period, adults, larvae and ova were collected for further research during field visits, while an in-house breeding programme involving wild-collected gravid females was initiated with a view to examine an adequately sized series of specimens. A total of 38 adults, including 23 male specimens, >150 ova, 233 larvae, and 220 pupae were examined. In addition to a thorough morphometric assessment, statistical tests were performed using one-way ANOVA and multivariate analysis. Results demonstrate morphological characters of P. saharae and, to a lesser degree, of P. machaon, suggesting that the taxon is plausibly the outcome of a hybrid swarm whose occurrence on the island was facilitated by Pleistocene lowstands (when the island was physically connected with the north African continental landmass, the taxon's centre of origin). Based on results, involving all four stages of metamorphosis, it is proposed to 'anchor' the taxon to Papilio saharae Oberthür, 1879 and assign it subspecific status, aferpilaggi ssp. nov.

Uplifting mountains and shaking deserts: volcano-tectonic earthquakes revealed by soft-sediment-deformation structures in Upper Cretaceous aeolian deposits

ABSTRACT During the last stages of Gondwana fragmentation, large regions of the newly formed South American continent were covered by extensive deserts. Some parts of this continental landmass were synchronously affected by pronounced tectonism and magmatism, which were responsible for reshaping the regional topography. In this context, the southwestern part of the Sanfranciscana Basin in central Brazil is a key area for understanding this particular period in the geodynamic evolution of the South American continent. Aeolian deposits of the Posse Formation in the basin occur in direct association with volcanic rocks of the Upper Cretaceous Mata da Corda Group. Here, we report evidence of synsedimentary magmatism in direct association with soft-sediment-deformation structures, including flame structures, load casts and pseudonodules, water-escape structures, convolute lamination, faults, breccias, and clastic dikes, developed exclusively in aeolian sandstone and siltstone facies. The deformation features are interpreted as indicative of liquefaction, fluidization, and brittle behavior of the loose to partially lithified, wet sandy–silty sediments. The Late Cretaceous aeolian sedimentation is contemporaneous with the uplift of the Paranaíba High and associated magmatism in the Minas–Goiás Alkaline Province. In this context, these significant volcano-tectonic activities are considered to have triggered ductile to brittle deformation in the reported aeolian deposits.