Complete Row Research Articles

Redescription of an endemic mahseer, Tor mahanadicus (David, 1953) from Mahanadi River basin, India based on integrated morphological and molecular techniques.

Tor mahanadicus was originally described as Tor mosal mahanadicus with inadequate information on its morphological traits and no designated type specimen. Currently, T. mahanadicus is synonymized with Tor putitora, solely based on partial molecular data despite significant morphological differences. In this study, we performed an integrated morphological and molecular analysis to redescribe T. mahanadicus from the Mahanadi River. Tor mahanadicus is distinguished from all known Indian Tor species by the presence of 2 complete rows of scales between pelvic fin origin and lateral line, small eye (15.3-16.9% in head length), and a wide mouth gap (21.7-23.8% in head length). Moreover, it undoubtedly distinguished from the closely related species Tor putitora by having a wider body depth (26.8-28.2% in standard length vs. 14.5-19.9%), short snout length (23.3-26.4% in head length vs. 28.0-29.3%) and wide inter orbit space (30.5-37.3% in head length vs. 27.6-28.5%). Additionally, the molecular phylogenetic tree generated from a combination of three genes demonstrates a monophyletic clade separate from the Tor putitora clade. Based on the distinct morphological traits and mitochondrial gene sequences, we established Tor mahanadicus as separate species under the genus Tor.

Detection of Cracks in Turbomachinery Blades by Online Monitoring

AbstractThe presence of a crack in a blade can change the natural frequencies of that blade. It has long been a goal to detect blade cracks by assessing the change in a measured vibration frequency of the blade over time. It has been found that prior frequency assessment methods can be less accurate than is desirable to reliably detect the relatively small frequency changes that are typically associated with blade crack sizes of practical interest. This paper describes a method in which potential temporal changes in the frequencies of individual blades are assessed by periodically analyzing complete rows of blades using mistuning analysis techniques that treat the blade rows as coupled systems, in contrast to other techniques that consider each blade individually in turn. This method, while computationally complicated and challenging, has been found to be capable of detecting blade root cracks that are much smaller than those that can be detected using other techniques. Moreover, this method has been demonstrated to detect cracks that are much smaller than the critical size for mechanical separation of the blade from the rotor. This improved frequency assessment technique has been used to identify more than 30 blades with frequency changes that were considered to be potential indicators of blade cracks. Subsequent inspections verified indications in all of those blades. In addition to providing operational guidance, the frequency change data were used to infer the time periods during which crack growth had occurred.

New insight of the genus Cephalomyopsis (Caviomorpha, Cephalomyidae): Systematic revision and paleobiogeographical implications

Cephalomyopsis hypselodontus is a member of the family Cephalomyidae, originally described based on some isolated remains from the Colhuehuapian locality of Bryn Gwyn (Chubut Province, Argentina). Some specimens from the Deseadan localities of Cabeza Blanca (Chubut Province, Argentina) and Lacayani (La Paz Department, Bolivia), first described as Cephalomyidae morphotype 2 and 3, were later included in the genus. Also, a specimen found in Arroyo Avalos (Corrientes Province, Argentina) was included in the genus, but a taxonomic revision at the species level was never conducted. In this contribution, I revise all the specimens included in the genus, as well as new specimens found in Bryn Gwyn. The new specimens include the most juvenile teeth known so far and the first complete upper tooth row known for the genus. As a result, I define a single species for the genus and discuss the implication of this new systematic proposal. The species has a wide geographical distribution and a long biochron, the longest among cephalomyids. Finally, I discuss the dissimilarities between different localities in Argentina and Bolivia and the implications in the evolutionary history of Cephalomyidae and Chinchilloidea.

The modular Weyl–Kac character formula

We classify and explicitly construct the irreducible graded representations of anti-spherical Hecke categories which are concentrated one degree. Each of these homogeneous representations is one-dimensional and can be cohomologically constructed via a BGG resolution involving every (infinite dimensional) standard representation of the category. We hence determine the complete first row of the inverse parabolic p-Kazhdan–Lusztig matrix for an arbitrary Coxeter group and an arbitrary parabolic subgroup. This generalises the Weyl–Kac character formula to all Coxeter systems (and their parabolics) and proves that this generalised formula is rigid with respect to base change to fields of arbitrary characteristic.

Scalable distributed least square algorithms for large-scale linear equations via an optimization approach

This paper proposes scalable distributed least square algorithms for solving large-scale linear algebraic equations via multi-agent network. We consider two kinds of different decomposition structures of Ax=b, in which each agent only knows a sub-block of linear equation instead of the complete row or column information in the existing results. By introducing one extra variable, the least square problem of linear equation can be transformed into a distributed constrained optimization problem. Based on the augmented Lagrange method, two continuous-time scalable distributed primal–dual algorithms are developed, in which all the states of agents have fewer dimension and can be even scalars. Subsequently, the discrete-time distributed algorithms with constant step-sizes are proposed by using the Euler discretization method, and the explicit bounds of constant step-sizes are provided. Based on the KKT condition and Lyapunov stability, we show that the proposed continuous-time and discrete-time distributed algorithms collaboratively obtain a least square solution with a linear convergence speed, and also own an additional property that verifies whether the obtained solution is an exact solution. Finally, some simulation examples are carried out to verify the effectiveness of the proposed algorithms.

Robust Correspondence Analysis

AbstractCorrespondence analysis is a method for the visual display of information from two-way contingency tables. We introduce a robust form of correspondence analysis based on minimum covariance determinant estimation. This leads to the systematic deletion of outlying rows of the table and to plots of greatly increased informativeness. Our examples are trade flows of clothes and consumer evaluations of the perceived properties of cars. The robust method requires that a specified proportion of the data be used in fitting. To accommodate this requirement we provide an algorithm that uses a subset of complete rows and one row partially, both sets of rows being chosen robustly. We prove the convergence of this algorithm.

Protura (Arthropoda, Hexapoda) of the Middle East, with the description of a new species

In 2021 as part of a research on apterygotes in Iran 18 specimens of Protura were collected; 14 of them were identified as Acerella muscorum (Ionesco, 1930), one was an Acerentomon belonging to the affine group, three were female specimens of a new species of Acerentulus described in this paper. Like other species belonging to confinis group, Acerentulus iranicus sp. nov. has a complete row of posterior setae (18) on tergite VII, a long foretarsal sensillum a almost reaching seta γ3, and sensilla b and c almost of the same length, both passing the base of seta γ3. The closest species to A. iranicus sp. nov. is A. setosus Szeptycki, 1993 from Western Caucasus (Russia). This species differs from the new species for its smaller pseudoculi, the ratio between mesothoracic setae P1 and P2, the presence of a doubled pore on tergite XII, and the longer foretarsal sensillum a. The partial description of an undetermined specimen of Acerentomon belonging to the affine group is outlined too. A key to the Protura species known from the Middle East is provided. http://www.zoobank.org/urn:lsid:zoobank.org:pub:A0243956-A081-4D5D-8FEA-361CA7123852

How to Lay Bricks: Local-Global Alignment Predicts Preference for Shifted Tile Patterns.

We examine the aesthetic characteristics of row tile patterns defined by repeating strips of polygons. In experiment 1 participants rated the perceived beauty of equilateral triangle, square and rectangular tilings presented at vertical and horizontal orientations. The tiles were shifted by one-fourth increments of a complete row cycle. Shifts that preserved global symmetry were liked the most. Local symmetry by itself did not predict ratings but tilings with a greater number of emergent features did. In a second experiment we presented row tiles using all types of three- and four-sided geometric figures: acute, obtuse, isosceles and right triangles, kites, parallelograms, a rhombus, trapezoid, and trapezium. Once again, local polygon symmetry did not predict responding but measures of correspondence between local and global levels did. In particular, number of aligned polygon symmetry axes and number of aligned polygon sides were significantly and positively correlated with beauty ratings. Preference was greater for more integrated tilings, possibly because they encourage the formation of gestalts and exploration within and across levels of spatial scale.

Optical RAM Row With 20 Gb/s Optical Word Read/Write

During the past years, optical memories have emerged as the main way to enable fast access times and increased memory bandwidth in synergy with optical interconnections. The migration, however, to fully functional and practical optical RAMs and cache memories will require an additional yet essential step: Optical memories have to get organized into an optical memory subsystem for delivering seamless cooperation between storage and peripheral decoding units, exploiting also the wavelength dimension for efficient RAM architectural layouts. In this paper we demonstrate the first experimentally verified Wavelength Division Multiplexing (WDM) -enabled optical RAM Row subsystem comprising both the peripheral decoding circuitry and a 4-bit optical RAM Row for storing 20Gb/s 4-bit WDM-formatted optical data words. The proposed scheme comprises a single shared multi-λ Semiconductor Optical Amplifier Mach-Zehnder Interferometer (SOA-MZI) Access Gate (AG) for granting access-control to the complete RAM Row, a passive Column Decoder (CD) that directs the incoming WDM-formatted data words to the respective RAM cells that are in turn based on four broadband Indium Phosphide (InP) photonic integrated Flip-Flops (FFs). Our experimental verification firstly investigates the broadband capability of the elementary InP monolithic RAM cell at 5 Gb/s, revealing error-free Write functionality with <4.5 dB power penalty across the whole C-band confirming in this way the potential of efficient scaling to multi-cell WDM-enabled RAMs. Then, experimental validation of the complete 4-bit WDM-enabled optical RAM Row with 20 Gb/s WDM-formatted data words is performed, featuring simultaneous Access gating, decoding and storage operation for all four RAM cells, having a peak power penalty within the range of [4.6-5.6] dB for Write and [2.2-2.9] dB for Read functionalities, respectively, at a Bit Error Rate of 10 <sup xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink">−9</sup> .

Two additional new species of Clypeocaenis Soldán, 1978 (Ephemeroptera: Caenidae) from the Western Ghats of Peninsular India.

Two new mayfly species of Clypeocaenis (Ephemeroptera: Caenidae) are described based on nymphal and adult characters from Kaveri River, Kodagu, Karnataka, Southern India. The main characteristics that distinguish Clypeocaenis kaveri sp. nov. from other Oriental species are: (i) fore femur with 4 long spines at middle of inner margin, a row of bifid spines at outer margin and row of bifid spines at subapical surface; (ii) dorsal surface of femora with complete transverse row of bifid setae located 3/4 distance from the base; (iii) forceps smaller than penis lobe. The main characteristics that distinguish Clypeocaenis napoklu sp. nov. from other Oriental species are: (i) fore femur with 2 long spines at middle of inner margin and row of thin setae on outer margin; segment 2 with bifid spine at apex.(ii) dorsal surface of femora with incomplete transverse row of bifid setae located 2/3 distance from the base; (iii) forceps little longer than penis lobe. A key for the nymphs of described species has been provided.

New remains of Glyptodontidae (Mammalia, Xenarthra) from the Salicas Formation (late Miocene, Messinian), La Rioja Province, Argentina

In South America, glyptodonts are recorded since the late Eocene to the early Holocene. Their fossil record is represented mainly by disarticulated osteoderms during the Eocene and Oligocene, whereas more complete skeletons appeared since the early Miocene. The characters from the dorsal surface of the osteoderms (e.g., figures, foramina, sulci) are used for systematic recognition and phylogenetical relationships of Glyptodontidae. The aim of this contribution is to analyze a new collection of glyptodontid osteoderms from the Salicas Formation (late Miocene) at La Rioja Province, Argentina. The glyptodontids osteoderms herein presented are assigned to 1) Stromaphorus compressidens (CRILAR-Pv 113 and CRILAR-Pv 114) given the osteoderms have a central figure surrounded by a first row of 12–14 peripheral figures and, by a not always a complete second row of peripheral figures; and 2) Eosclerocalyptus cf. proximus (CRILAR-Pv 115) given the osteoderms have a circular to subelliptical central figure surrounded by a first row of 10 peripheral figures and by some peripheral figures located on a second row. The record of S. compressidens from Salicas Formation herein presented constitutes the first record of the species for the late Neogene of La Rioja Province; whereas the record of E. cf. proximus corroborates the presence of the genus in the Salicas Formation. The mammal assemblage recorded at the El Degolladito area suggests a latest Miocene age (Messinian) for the Salicas Formation at this locality.

Мaize hybrids for grain and silage in agro-ecological conditions of the Republic of Srpska

Maize is used in human and animal nutrition, in industrial processing (nutritional products, pharmaceutical industry, alcohol industry, chemical industry, textile industry). Economical significance of maize is derived from its yield, namely its total production of grain maize, as well as silage. Today, together with potatoes and sugar cane, maize is used for production of bioethanol, a very important source of renewable energy. Maize is grown worldwide on surfaces of 192,05 million ha, on which around 1,108 billion tonnes of grain is produced. The largest producers of genetically modified corn in the world are USA, Brasil, Argentina, while in the EU GM corn is grown in Spain and Portugal. Maize is the most important field crop in the Republic of Srpska and is grown on 46.7 % of sown areas. In a ten-year period (2008-2017) the least growing area on which the maize was grown were in 2012 (120,901 ha), and the largest in 2008 (148,539 ha). The limiting factor in achieving high yield are meteoroidal conditions (high temperatures, draught or excessive rainfall) because of which average yield variation is from 2.7 to 5.3 t ha-1 . The biggest cereals and maize production in Republic of Srpska, the one of 1,657,556 tonnes, was achieved in 2016 which, compared to 2015 was an increase of 46%, or 520,000 tonnes, and was affected by a record breaking maize production of 1.178.423 tonnes. In organic corn production the sweet and popcorn are the most bred. The improvement in agricultural production mostly depends on perfecting the existing and developing the new and appropriate technical solutions. The accent is increased on protection of the environment, production of healthy and quality food and decrease in production expenses, and all of this can be achieved by applying the conservation treatment of soil. Growing corn in complete and regular crop rows, especially in which there are legume plants, can significantly affect the decrease in weediness and increase in grain yield. The corn is often grown in combined sowing with other plants, most often with soy, beans and fodder pumpkin. Maize breeding in past period created many hybrids, which enabled a permanent yield increase. The methods of conventional breeding are efficient but slow in creating a germplasm tolerant on existing range of climate conditions and are not suitable for rapid improvement of multiple stress tolerance. Molecular breeding enables increase in speed and efficiency of breeding. The only institution in Bosnia and Herzegovina which deals with breeding and creating new species and hybrids of grown plants, adapted to local agroecological conditions is the PI Agricultural Institute, Banja Luka. So far, the Institute created nine maize hybrids, the most significant being BL 43.

A new species of Eremias (Squamata: Lacertidae) from the arid mountains of Pakistan.

A new, morphologically distinctive lacertid lizard of the genus Eremias (Rhabderemias) is described from the arid mountains of northwestern Balochistan Province in Pakistan. Eremias kakari sp. nov. has an isolated distribution and can be easily distinguished from all other species of mainly desert subgenus Rhabderemias (E. andersoni, E. cholistanica, E. fasciata, E. lineolata, E. pleskei, E. scripta, E. vermiculata). Apart from other differences, E. kakari sp. nov. can be distinguished from geographically close members of the subgenus Rhabderemias (E. cholistanica, E. fasciata, and E. scripta) by having a single row of subdigital lamellae and a complete row of lateral scales and hence three scales around the penultimate phalanx of 4th toe. The new species is morphologically (dorsal pattern) very similar to E. fasciata but can be distinguished from this species for having 22-26 subdigital lamellae under 4th toe, 48-55 dorsal scales across midbody, ventrals in 11-14 oblique longitudinal series across the belly, 17-21 femoral pores and 17-21 scales in the 9th-10th annulus posterior to the postcloacal granules. The new species is currently known only from the type locality situated in the Toba Kakar Range, near to Tanishpa village. However, we expect that Eremias kakari sp. nov. would have a broader range in northwestern Pakistan and southeastern Afghanistan. An identification key for the Pakistani Eremias, together with other remarks to the new species, is presented.

Thecodont tooth attachment and replacement in bolosaurid parareptiles.

Permian bolosaurid parareptiles are well-known for having complex tooth crowns and complete tooth rows in the jaws, in contrast to the comparatively simple teeth and frequent replacement gaps in all other Paleozoic amniotes. Analysis of the specialized dentition of the bolosaurid parareptiles Bolosaurus from North America and Belebey from Russia, utilizing a combination of histological and tomographic data, reveals unusual patterns of tooth development and replacement. The data confirm that bolosaurid teeth have thecodont implantation with deep roots, the oldest known such example among amniotes, and independently evolved among much younger archosauromorphs (including dinosaurs and crocodilians) and among synapsids (including mammals). High-resolution CT scans were able to detect the density boundary between the alveolar bone and the jawbone, as confirmed by histology, and revealed the location and size of developing replacement teeth in the pulp cavity of functional teeth. Evidence provided by the paratype dentary of Belebey chengi indicates that replacement teeth are present along the whole tooth row at slightly different stages of development, with the ontogenetically more developed teeth anteriorly, suggesting that tooth replacement was highly synchronized. CT data also show tooth replacement is directly related to the presence of lingual pits in the jaw, and that migration of tooth buds occurs initially close to these resorption pits to a position immediately below the functional tooth within its pulp cavity. The size and complex shape of the replacement teeth in the holotype of Bolosaurus grandis indicate that the replacement teeth can develop within the pulp cavity to an advanced stage while the previous generation remains functional for an extended time, reminiscent of the condition seen in other amniotes with occluding dentitions, including mammals.

First Snout with Complete Teeth Row of Small Titanosaur in Indo-Pakistan Found from the Latest Maastrichtian Vitakri Formation of Pakistan; Associated Cranial and Postcranial Skeletons of &amp;lt;i&amp;gt;Saraikimasoom vitakri&amp;lt;/i&amp;gt; (Poripuchia, Stocky Titanosauria, Sauropoda) from Pakistan and Referred Fossils from India

Titanosaurs articulated cranial elements especially jaw bones with articulated complete teeth row are extraordinarily rare. Here the holotypic snout with articulated jaw bones and complete teeth row of Saraikimasoom vitakri are being presented which has international significance and contributes to understanding the evolutionary relationships and paleobiogeographic history of the vertebrates of Indo-Pakistan subcontinent. The completeness of the skeleton, in particular the presence of a well-preserved skull has a significant influence on resolution of titanosaur phylogeny. Here Saraikimasoom vitakri holotypic snout from South Kinwa, and referred jaw with articulated pointed and rounded teeth along with associated postcranial fossils from north Kinwa, and other associated materials from Mari Bohri, mid-Sangiali, Zubra Basti Nala and Grut localities of Pakistan are being presented. All these assemblages show consistent results.

Mammal Molar Size Ratios and the Inhibitory Cascade at the Intraspecific Scale.

SynopsisMammalian molar crowns form a module in which measurements of size for individual teeth within a tooth row covary with one another. Molar crown size covariation is proposed to fit the inhibitory cascade model (ICM) or its variant the molar module component (MMC) model, but the inability of the former model to fit across biological scales is a concern in the few cases where it has been tested in Primates. The ICM has thus far failed to explain patterns of intraspecific variation, an intermediate biological scale, even though it explains patterns at both smaller organ-level and larger between-species biological scales. Studies of this topic in a much broader range of taxa are needed, but the properties of a sample appropriate for testing the ICM at the intraspecific level are unclear. Here, we assess intraspecific variation in relative molar sizes of the cotton mouse, Peromyscus gossypinus, to further test the ICM and to develop recommendations for appropriate sampling protocols in future intraspecific studies of molar size variation across Mammalia. To develop these recommendations, we model the sensitivity of estimates of molar ratios to sample size and simulate the use of composite molar rows when complete ones are unavailable. Similar to past studies on primates, our results show that intraspecific variance structure of molar ratios within the rodent P. gossypinus does not meet predictions of the ICM or MMC. When we extend these analyses to include the MMC, one model does not fit observed patterns of variation better than the other. Standing variation in molar size ratios is relatively constant across mammalian samples containing all three molars. In future studies, analyzing average ratio values will require relatively small minimum sample sizes of two or more complete molar rows. Even composite-based estimates from four or more specimens per tooth position can accurately estimate mean molar ratios. Analyzing variance structure will require relatively large sample sizes of at least 40–50 complete specimens, and composite molar rows cannot accurately reconstruct variance structure of ratios in a sample. Based on these results, we propose guidelines for intraspecific studies of molar size covariation. In particular, we note that the suitability of composite specimens for averaging mean molar ratios is promising for the inclusion of isolated molars and incomplete molar rows from the fossil record in future studies of the evolution of molar modules, as long as variance structure is not a key component of such studies.

Latent Complete Row Space Recovery for Multi-View Subspace Clustering

Multi-view subspace clustering has been applied to applications such as image processing and video surveillance, and has attracted increasing attention. Most existing methods learn view-specific self-representation matrices, and construct a combined affinity matrix from multiple views. The affinity construction process is time-consuming, and the combined affinity matrix is not guaranteed to reflect the whole true subspace structure. To overcome these issues, the Latent Complete Row Space Recovery (LCRSR) method is proposed. Concretely, LCRSR is based on the assumption that the multi-view observations are generated from an underlying latent representation, which is further assumed to collect the authentic samples drawn exactly from multiple subspaces. LCRSR is able to recover the row space of the latent representation, which not only carries complete information from multiple views but also determines the subspace membership under certain conditions. LCRSR does not involve the graph construction procedure and is solved with an efficient and convergent algorithm, thereby being more scalable to large-scale datasets. The effectiveness and efficiency of LCRSR are validated by clustering various kinds of multi-view data and illustrated in the background subtraction task.

A new species of Xenylla Tullberg, 1869 (Collembola: Hypogastruridae)from Peru, with a key to Neotropical species of the genus.

A new species belonging to genus Xenylla Tullberg, 1869 from Peru is described and illustrated. The new species, Xenylla gwiazdowiczi sp. nov., can be distinguished from previously described Xenylla species by combination of the following characters: mucro separated from dens, presence of complete row of setae c on dorsal side of the head, four thick sensilla on antennal segment IV, 19,19,18 setae on tibiotarsi of legs I-III, slightly serrated setae on abdominal segments V-VI, presence of two weakly clavate tenent hairs on each leg. The new species resembles X. spinosissima Najt Rubio, 1978 and X. humicola (Fabricius, 1780). Similarities between them have been discussed. An identification key to Neotropical species of the genus is provided.

Characterization of age-dependent variability in the flank scales of two scorpaeniformes fishes by applying light and scanning electron microscopy imaging

The morphology of scales from four flank regions in two species of Scorpaeniforme fishes of the family Platycephalidae was studied by applying light and scanning electron microscopy, and the age-dependent variability of scale structures was discussed. The scales of four flank regions from three size classes in both species were ctenoid. The posterior margin of all the ctenoid scales was formed by two rows of cteni, while the three complete rows of cteni were not observed in the examined scales. The cteni of both rows were similar in morphology. Some scale characteristics showed alteration during the fish growth, including the shape of the anterior region of scale in Platycephalus indicus, the presence of lepidonts in Grammoplites suppositus, and the general shape of scale as well as the indices SCL.SCW and JSW.SL in both species. Also, the number of cteni in posterior margin of scales was increased during fish development, while their general morphology did not change significantly. The number of cteni in the scales of small and large fishes in G. suppositus was found to be obviously lower than P. indicus. By considering the function of lepidonts in improving the hydrodynamic efficiency of swimming, it can be assumed that G. suppositus is not probably very active in swimming as P. indicus. As a conclusion, modification in the ornamentations of the posterior region are subject to alteration during the growth of the fish. Besides its developmental changes, some scale characters in the adult forms have taxonomic significance, including the square-shaped scales in P. indicus vs. the oval-shaped scale in G. suppositus; a total of 26–31 radii in the anterior field in P. indicus vs. 11–12 radii in G. suppositus; the presence of many (54–58) cteni in the posterior part in P. indicus vs. a moderate (19–23) cteni in G. suppositus. These characters could be used as additional morphological data to be used for studying systematics and even phylogeny of the flathead fishes.

A new genus of Pycnodontidae (Actinopterygii) from the Upper Jurassic of France and Germany, included in a phylogeny of Pycnodontiformes

AbstractThe new genus Thiollierepycnodus is erected here to accommodate ‘Gyrodus’ wagneri or ‘Proscinetes’ wagneri, a taxon known since the 19th century from the marine Kimmeridgian Plattenkalk of Cerin (France) and since 2014 at the excavation site of the Naturkunde-Museum Bamberg in Wattendorf (Germany). Thiollierepycnodus is placed in Pycnodontidae (Upper Jurassic to Eocene), which is the most advanced family of Pycnodontiformes. With 154–152 Myr (Upper Jurassic), Thiollierepycnodus belongs to the stratigraphically oldest Pycnodontidae, still possessing some primitive features such as four incisiform teeth in the dentary, a crown of bifurcated anterior dentary teeth, reduced scale bars posterior to the dorsal fin origin, but the area of pterygiophores without scales and nine to 10 postanal ventral keel scales and approximately 10 complete scale rows posterior to the cloaca. A phylogenetic analysis leads to the placement of this taxon in Proscinetinae. This and the introduction of two additional taxa from the Cretaceous of Lebanon in the phylogeny are further steps to enlighten the broader evolutionary patterns of Pycnodontidae.