1 Department of Biology, University of Washington, Seattle, WA, USA 2 Department of Fisheries and Wildlife Sciences, Virginia Polytechnic Institute and State University, Blacksburg, VA, USA 3 Department of Biology, Valdosta State University, Valdosta, GA, USA 4 New Mexico Museum of Natural History, Albuquerque, NM, USA 5 Department of Environmental, Population, and Organismic Biology, University of Colorado, Boulder, CO, USA 6 School of Molecular and Life Sciences, University of Limpopo, Sovenga, South Africa