Central Canadian Arctic Research Articles

Palaeoeskimo Demography on Western Boothia Peninsula, Arctic Canada

Surveys on western Boothia Peninsula in 2004 documented 483 Palaeoeskimo dwellings spanning approximately 3300 years (4500–1200 B.P. in uncalibrated radiocarbon years), about the total time range for Palaeoeskimo groups in the central Canadian Arctic. On the basis of dwelling elevations above sea level and a series of radiocarbon dates, Palaeoeskimo occupation appears to have passed through multiple boom-and-bust cycles. Following the first peopling of the region about 4500 B.P., populations rose to a maximum between about 4200 and 3600 B.P., followed by a crash. A recovery between 3200 and 2500 B.P. led to a second decline, and a final, partial recovery between 1600 and 1200 B.P. was followed by the disappearance of Palaeoeskimo groups. Although climate change cannot be ruled out as a causal factor for these cycles, there is no compelling evidence for such a scenario. Resource over exploitation is equally plausible, although we do not necessarily favor one cause over the other.We interpret the intrasite patterns to indicate that Palaeoeskimo settlements were comprised of dispersed nuclear families or small extended families for most of the year, but annual aggregations involved 100 individuals or more. Minimal social units do not appear to have changed during seasonal aggregations in Pre-Dorset times (4500–2500 B.P.). By Dorset times (2500–700 B.P.), however, minimal social units sometimes melded together to form one or a few larger units living in one or a few large dwellings. The latter may represent the social precursor of later Dorset longhouse aggregations.

A 2000 year midge-based paleotemperature reconstruction from the Canadian Arctic archipelago

A lake sediment core recovered from Lake V57 on Victoria Island, Nunavut, Canada, spanning the last 2000 years, was analyzed for sub-fossil midge remains and organic-matter content (estimated by loss-on-ignition (LOI)). Significant changes in midge community composition occurred during the last 2000 years, with a distinct midge community appearing after 1600 AD. The chironomid community between 0 and 1600 AD was dominated by Heterotrissocladius, Tanytarsus, Abiskomyia, and Paracladius. At approximately 1600 AD, Heterotrissocladius decreased in relative abundance and taxa such as Corynocera ambigua, C. oliveri, Psectrocladius sordidellus type, and Pentanneurini increased in relative abundance. Previously published midge-based inference models for average July air temperature (AJAT) and summer surface–water temperature (SSWT) were applied to the subfossil midge stratigraphy. The AJAT reconstruction indicates relatively cool conditions existed between 1100 and 1600 AD, with exceptional warming occuring after 1600 AD, as lake productivity inferred from organic-matter content increased concomitantly with midge-inferred AJAT and SSWT. The cooler conditions between 1200 and 1600 AD, and the pattern of warming over recent centuries inferred from Lake V57 is broadly consistent with temperature-sensitive biogenic silica records from other sites in the central Canadian Arctic and the treeline zone to the south suggesting a regionally synchronous response to climate forcing.

Pollen-based reconstructions of late Holocene climate from the central and western Canadian Arctic

Two lake-sediment cores from the western and central Canadian Arctic were used to investigate late Holocene climate variability in the region. Both cores were analyzed for pollen, organic matter, biogenic silica, and magnetic susceptibility, and were dated using a combination of 210Pb and 14C techniques. Core MB01, from southwestern Victoria Island, provides a 2600-year-long record. Fossil pollen percentages, along with other parameters, suggest the occurrence of a cold period around 2400 cal year BP (450 BC), followed by slightly warmer conditions by 1800 cal year BP (150 AD), and a return to cooler conditions throughout much of the last millennium. Core SL06, from southern Boothia Peninsula, shows more subtle changes in pollen percentages over its 2500-year duration, but an increase in Cyperaceae and decrease in Oxyria pollen around 1400 cal year BP (550 AD) are indicative of warmer conditions at that time. Quantitative climate reconstructions from these pollen sequences were compared to two other pollen-based climate records from the region and indicate the presence of a widespread wet period ~1500 cal year BP (450 AD), and a cool and dry Little Ice Age. In the reconstructions based on pollen percentage data, the twentieth century summer temperature and annual precipitation in the central and western Canadian Arctic were comparable to that which occurred over the last 2500 years. However, pollen-influx values increase in the most recent sediments, suggesting high plant productivity during the late twentieth century.

Development of a chironomid-based air temperature inference model for the central Canadian Arctic

Subfossil midge remains were identified in surface sediment recovered from 88 lakes in the central Canadian Arctic. These lakes spanned five vegetation zones, with the southern-most lakes located in boreal forest and the northern-most lakes located in mid-Arctic tundra. The lakes in the calibration are characterized by ranges in depth, summer surface-water temperature (SSWT), average July air temperature (AJAT) and pH of 15.5 m, 10.60°C, 8.40°C and 3.69, respectively. Redundancy analysis (RDA) indicated that maximum depth, pH, AJAT, total nitrogen-unfiltered (TN-UF), Cl and Al capture a large and statistically significant fraction of the overall variance in the midge data. Inference models relating midge abundances and AJAT were developed using different approaches including: weighted averaging (WA), weighted averaging-partial least squares (WA-PLS) and partial least squares (PLS). A chironomid-based inference model, based on a two-component WA-PLS approach, provided robust performance statistics with a high coefficient of determination (r2 = 0.77) and low root mean square error of prediction (RMSEP = 1.03°C) and low maximum bias. The use of a high-resolution gridded climate data set facilitated the development of the midge-based inference model for AJAT in a region with a paucity of meteorological stations and where previously only the development of a SSWT inference model was possible.

Evaluating the Response of Cladocera to Recent Environmental Changes in Lakes from the Central Canadian Arctic Treeline Region

ABSTRACTArctic and subarctic ecosystems have undergone considerable environmental changes in recent years as the result of climate warming. Fossil records of freshwater diatoms in lakes throughout the circumpolar Arctic, including the central Canadian Arctic treeline region, have revealed marked directional shifts in diatom assemblages in recent lake sediments. These algal changes have been linked to longer growing seasons, decreased duration of ice cover, and/or increased thermal stability. The effects of these recent environmental changes on higher trophic levels, such as the Cladocera, are still unclear. Using cladoceran remains preserved in the sediments of 50 lakes, which were previously examined for changes in diatoms, we show that significant changes in cladoceran species composition have occurred from pre-industrial times to the present. However, these changes are considerably muted compared to the more substantial changes observed in the diatom record. We found no consistent patterns of change in planktonic cladocerans (i.e. Daphnia, Bosmina) within our study lakes, and the response of the Cladocera to environmental changes does not appear to be strongly coupled to recent changes in diatom communities, thus further confirming that the previously observed diatom changes were related to bottom-up limnological controls. These results highlight the complex response of Arctic freshwater food webs to climate change, and the need for incorporating multi-trophic studies into climate change investigations.

The influence of low‐level thermal inversions on estimated melt‐season characteristics in the central Canadian Arctic

AbstractDaily vertical temperature gradients were examined in order to infer melt‐event characteristics at an elevation relevant to basin‐scale snow and plateau ice‐melt studies in the Canadian Arctic. Surface and upper‐air temperature data from Resolute, Cornwallis Island, was used to estimate vertical lapse rates up to 300 m asl, and to identify the presence of inversions at that altitude. Lapse rates vary throughout the melt season and are substantially less than typically published generalized values. Thermal inversions are more frequent during the melt season than in the periods immediately before and after, suggesting a strong control on intraseasonal temperature patterns. In July, the period of maximum temperature in the Arctic, inversion frequency is highest and closely related to calculated melting degree‐days. Results show that increased summer mean temperature resulted in a substantial lengthening of estimated melt events, as opposed to increased event intensity. Increased inversion frequency leading to shallower vertical lapse rates since the late 1980s is speculated to be the result of synoptic‐scale climate patterns, and is potentially an important meteorological mechanism for enhanced glacial melt since the late 1980s. Copyright © 2008 Royal Meteorological Society

Responses of Fragilarioid-dominated diatom assemblages in a small Arctic lake to Holocene climatic changes, Russell Island, Nunavut, Canada

We present a paleolimnological record spanning the Holocene from a small lake on Russell Island (Lake PW02), in the central Canadian Arctic Archipelago (74.07° N, 97.77° W, 182 m asl). Fragilarioid diatom types in the genera Pseudostaurosira, Staurosira and Staurosirella constitute >90% of valves in fossil samples. Using modern biogeographic data which specify the temperature optima of the Fragilarioid diatom taxa, we present new inferences about the timing of paleoclimatic changes in the central Arctic islands. The early Holocene was characterized by maximum values for sediment organic matter, and lower ratios of Staurosirella pinnata to Staurosira construens v. venter, suggesting warm summer air temperatures between about 9500−6500 cal year BP. Influxes of biogenic silica and diatom valves decreased following 4000 cal year BP, the sediment accumulation rate slowed and diatom taxa of the littoral zone diversified, suggesting cooler summers and more persistent lake ice. Variations in the species composition of the assemblages indicate paleoclimatic changes that are in broad agreement with other paleoenvironmental records from the Arctic including melt records from the Agassiz Ice Cap. Although autecological data remain incomplete for Fragilarioid taxa, our results indicate differences in these taxa in responses to paleoenvironmental change and underline the potential for the increased use of these taxa in paleoenvironmental reconstructions. The record from Lake PW02, as in other records from Arctic lakes with low algal diversity throughout the Holocene, shows a pronounced increase in diatom diversity since the 1920s, and diatom production since the 1970s far exceeds any recorded during the Holocene.

Spatial trends of perfluoroalkyl compounds in ringed seals (Phoca hispida) from the Canadian Arctic

The present study examined spatial trends of perfluoroalkyl compounds (PFCs) in liver samples from 11 populations of ringed seals (Phoca hispida) in the Canadian Arctic from 2002 to 2005. Trophic position and relative carbon sources were compared by analyzing stable nitrogen and carbon isotopes in muscle samples. Geometric mean concentrations of total C9-C15 perfluorinated carboxylates (PFCAs) ranged from 8.8 to 84 ng/g wet weight, and C9-C11 PFCAs predominated. Perfluorooctane sulfonate was the dominant PFC measured, with concentrations ranging from 6.5 to 89 ng/g wet weight, contributing between 29 and 56% of the total PFC concentration. Overall, mean PFC concentrations were similar between populations, and differences were attributed largely to elevated levels in the Gjoa Haven (Rae Strait, central Canadian Arctic archipelago) and Inukjuak populations (eastern Hudson Bay) and to lower concentrations at Pangnirtung (Cumberland Sound, Baffin Island). Mean stable nitrogen isotope ratios (+/-95% confidence intervals) ranged from 14.7 per thousand (+/-0.3 per thousand) at Nain (Labrador) to 17.9 per thousand (+/-0.7 per thousand) at Gjoa Haven, suggesting that all populations were within the same trophic level. Stable carbon isotope ratios varied widely between the seal populations, ranging from -22.9 per thousand (+/-0.2 per thousand) at Gjoa Haven to -17.7 per thousand (+/-0.4 per thousand) at Nain. The delta13C ratios from Gjoa Haven were significantly more depleted than those for other populations and may suggest a terrestrially based carbon source. The depleted stable carbon isotope ratio may explain the elevated PFC concentrations in the Gjoa Haven population. Analysis of covariance indicated that delta13C was a significant covariable for seven of nine seal populations for which delta13C values were available. After adjusting for delta13C values, concentrations of most PFCs generally were statistically greater in the Grise Fiord, Qikiqtarjuaq, Arviat, and Nain populations.

Trace Element Concentrations in Blood of Nesting King Eiders in the Canadian Arctic

The king eider (Somateria spectabilis) is a migratory species of sea duck whose North American population is thought to be declining. We determined levels of cadmium, lead, selenium, and mercury in blood from female king eiders nesting in the central Canadian Arctic from 2001 to 2003. Year-to-year repeatability estimates were calculated from birds sampled in 2 or 3 years. Repeatability coefficients were 0.45, 0.35, 0.58, and 0.25 for cadmium, lead, selenium, and mercury, respectively. The first three were significantly different from zero (p < 0.05), whereas the last approached significance (0.05 < p < 0.1). In 2001 and 2002, we also identified probable wintering locations of a subset of the birds. In both years, cadmium levels were higher and selenium levels were lower in birds inferred to have wintered in the eastern part of their range compared to those that had wintered in the west. There was little evidence that timing of breeding, timing of sampling, or body condition were related to levels of these trace elements, although in 1 of 2 years, lead levels were influenced by body condition and nest initiation date (R(2) = 0.24) and cadmium levels were related to incubation day (partial R(2) = 0.04). Year-to-year repeatability of cadmium and selenium levels among individuals in this population of king eiders was likely influenced by where they wintered.

Stratigraphy and Paleoecology of Quaternary Sediments Along Pasley River, Boothia Peninsula, Central Canadian Arctic

Quaternary sediments exposed along Pasley River consist of a lower marine deltaic sand overlain in succession by complexly interbedded tills and glaciomarine sediments (the lower glacigenic assemblage), by a mid-section fluvial gravel, by an upper marine deltaic sand, and by glaciomarine sediment and till (the upper glacigenic assemblage). The midsection fluvial gravels contain plant and insect fossils indicating a climate as warm as and perhaps warmer than present. The top of the gravel is more than 55 000 years old ; the unit is probably of Sangamonian age (&gt;75 000 ka) and separates Wisconsinan from lllinoian glacial deposits. The deltaic sands that underlie both glacigenic assemblages indicate substantial crustal depression during glacial buildup episodes prior to arrival of ice at the site. This implies that the process of buildup was slow and involved glacier expansion into major marine basins. Glaciomarine beds of the lower glacigenic assemblage locally contain abundant detrital terrestrial organic material as well as marine molluscs. The terrestrial organic detritus, an unusual constituent of glaciomarine sediment, is thought to have been released into the sea from glacier ice. These terrestrial fossil asemblages exhibit compositional differences which vary with the sediment faciès and probably reflect taphonomic factors such as differential buoyancy of the fossils. The upper marine deltaic sands contain some "old " rebedded plant detritus and amber indicating a nearby source of Tertiary sediment, possibly equivalent in age to the Beaufort Formation. Other rebedded fossils from the upper deltaic unit may be the same age as the mid-section fluvial gravels.

Glacial and Sea Level History of Lowther and Griffith Islands, Northwest Territories: A Hint of Tectonics

Lowther and Griffith islands, in the centre of Parry Channel, were overrun by the Laurentide Ice Sheet early in the last glaciation. Northeastward Laurentide ice flow persisted across at least Lowther Island until early Holocene déglaciation. Well constrained postglacial emergence curves for the islands confirm a southward dip of raised shorelines, contrary to the dip expected from the ice load configuration. This and previously reported incongruities may indicate regionally extensive tectonic complications of postglacial rebound aligned with major structural elements in the central Canadian Arctic Islands.

A palaeolimnological record of diatom-community dynamics and late-Holocene climatic changes from Prescott Island, Nunavut, central Canadian Arctic

Palaeolimnological analyses from Lake PW03, a small, dilute lake on crystalline bedrock on Prescott Island, Nunavut, Canada (73°07′ N, 96°41′ W, 243 m a.s.l.), provide a sensitive record of the impacts of Holocene climatic change on diatom communities. A high-resolution sequence from 3500 cal. BP to present indicates that a significant change in diatom assemblages takes place at 2300 cal. BP, when alkaliphilous fragilarioids and planktonic taxa are replaced by benthic, epiphytic and acidophilous diatoms. These changes coincide with the onset of neoglacial cooling and likely reflect more persistent ice-cover and associated acidification of lake water. Maximum diatom species richness occurs during this period of transition. Dense bryophyte communities in the littoral zone provide habitats to sustain diatom production even under the cooler conditions following 2300 cal. BP. The concentration of diatom valves declines at 500 cal. BP, coincident with cooling at the time of the `Little Ice Age', and minor changes in species composition take place at that time. The sediments deposited over the past 150 years show modest increases in fragilarioids and planktonic diatoms, in response to a longer ice-free season under a warmer climate. Diatom valve production increases significantly in these recent sediments but increases in diversity are relatively small. Diatom diversity, production and the species composition of diatom communities do not change simultaneously in response to climatic changes.

Tundra Game Drives: an Arctic-Alpine Comparison

Stone fences and blinds built by prehistoric hunters to gather and ambush elk and bighorn sheep above timberline in the Colorado Front Range are similar in concept and function to structures built by the Copper Inuit and their predecessors for hunting caribou near Bathurst Inlet, in the Central Canadian Arctic. Four principal differences exist: (1) Circular blinds and continuous rock walls are more numerous in the Front Range than in the Arctic, where arcuate breastworks and lines of widely spaced cairns predominate. Differences in prey-species behavior are the most probable explanation. (2) Stone house foundations, meat-drying facilities, meat caches, kayak-storage racks, and fox and wolf traps occur near drive sites along caribou migration routes in the Bathurst Inlet region. The structures imply long-term habitation made possible by a plentiful meat supply. Comparable structures are absent above timberline in the Front Range because people retreated to warmer environments in winter, and because steep terrain and deep snow discouraged return visits to high-altitude caches. (3) The technique was adopted much earlier in Colorado than in the Central Canadian Arctic. The oldest Front Range drive systems were constructed while the Laurentide Ice Sheet still covered the Bathurst Inlet landscape. (4) Pedestrian game-drive hunting was abandoned in the Front Range soon after arrival of the horse (ca. a.d. 1700), but remained an integral part of Copper Inuit subsistence until the mid twentieth century. The rich ethnographic and oral history record of communal hunting in the Arctic is invaluable for interpreting the Colorado structures.

The dynamics of landscape change and snow geese in mid‐continent North America

AbstractThe Mid‐Continent Population of the lesser snow goose, which breeds in the eastern and central Canadian Arctic and sub‐Arctic, and winters in the southern United States and northern Mexico has increased 5–7% annually from the late 1960s to the mid‐1990s, largely because of increased survival in response to an agricultural food subsidy. The rise in numbers complements the increased use of nitrogen fertilizers and a corresponding rise in yields of rice, corn, and wheat along the flyways and on the wintering grounds. In sub‐Arctic migration areas and at Arctic breeding colonies, foraging by high numbers of birds has led to loss of coastal vegetation, adverse changes in soil properties and the establishment of an alternative stable state of exposed sediment, which can be detected with LANDSAT imagery. At a local scale, gosling growth, size and survival decreased in affected areas and other taxa have been adversely affected. The food subsidy on wintering and migration areas appears insufficient to meet reproductive demands as foraging in spring continues to occur on southern Hudson Bay staging and nesting areas. The recent introduction of liberal hunting regulations may reduce population size in the near term, but the revegetation of these coastal ecosystems will take decades to achieve. The present pattern of vegetation loss in these Arctic coastal systems is likely to continue in the forseeable future.

Changes in mercury and cadmium concentrations and the feeding behaviour of beluga ( Delphinapterus leucas) near Somerset Island, Canada, during the 20th century

Beluga ( Delphinapterus leucas) continues to be an important food species for Arctic communities, despite concerns about its high mercury (Hg) content. We investigated whether Hg and cadmium (Cd) concentrations had changed during the 20th century in beluga near Somerset Island in the central Canadian Arctic, using well-preserved teeth collected from historical sites (dating to the late 19th century and 1926–1947) and during subsistence hunts in the late 1990s. Mercury concentrations in both historical and modern teeth were correlated with animal age, but 1990s beluga exhibited a significantly more rapid accumulation with age than late 19th century animals, indicating that Hg concentrations or bioavailability in their food chain had increased during the last century. The geometric mean tooth Hg concentration in modern 30 year old animals was 7.7 times higher than in the late 19th century, which corresponds to threefold higher concentrations in muktuk and muscle. Teeth from 1926 to 1947 were similar in Hg content to the late 19th century, suggesting that the increase had occurred sometime after the 1940s. In contrast, tooth Cd was not correlated with animal age and decreased during the last 100 years, indicating that anthropogenic Cd was negligible in this population. Late 19th century beluga displayed a greater range of prey selection (tooth δ 15N values: 15.6–20.5‰) than modern animals ( δ 15N: 17.2–21.1‰). To prevent this difference from confounding the temporal Hg comparison, the Hg–age relationships discussed above were based on historical animals, which overlapped isotopically with the modern group. Tooth δ 13C also changed to isotopically more depleted values in modern animals, with the most likely explanation being a significant shift to more pelagic-based feeding. Industrial Hg pollution is a plausible explanation for the recent Hg increase. However, without further investigation of the relationship between the range exploitation of modern beluga and their possible exposure to regional marine food chains with (naturally) higher Hg contents than their historical counterparts, we cannot unequivocally conclude that the increase was anthropogenically driven.

HIERARCHICAL HABITAT SELECTION BY TUNDRA WOLVES

Using resource selection functions, we examined habitat selection patterns of tundra wolves (Canis lupus) in the central Canadian Arctic at the level of the home range and within the home range. Esker habitat was relatively preferred by wolves at the home range level of selection, possibly indicating the importance of denning habitat as a limiting factor for tundra wolves. We failed to conclusively tie vegetation communities to movements of wolves within the home range. Eskers make up only about 1-2% of the landscape; hence, a potential conflict between wolf conservation and industrial development in the region may occur as eskers are targeted as a source of granular material for road construction. Results of this study underline the importance of scale dependence in habitat selection. Failure to view habitat selection as a hierarchical process may result in a narrow and possibly misleading notion of the value of habitats to animals.

Limnological characteristics of 56 lakes in the Central Canadian Arctic Treeline Region

Measured environmental variables from 56 lakes across the Central Canadian Treeline Region exhibited clear limnological differences among subpolar ecozones, reflecting strong latitudinal changes in biome characteristics (e.g. vegetation, permafrost, climate). Principal Components Analysis (PCA) clearly separated forested sites from tundra sites based on distinct differences in limnological characteristics. Increases in major ions and related variables (e.g. dissolved inorganic carbon, DIC) were higher in boreal forest sites in comparison to arctic tundra sites. The higher values recorded in the boreal forest lakes may be indirectly related to differences in climatic factors in these zones, such as the degree of permafrost development, higher precipitation and runoff, duration of ice-cover on the lakes, and thicker and better soil development. Similar to trends observed in DIC, substantially higher values for dissolved organic carbon (DOC) were measured in boreal forest lakes than in arctic tundra lakes. This was likely due to higher amounts of catchment-derived DOC entering the lakes from coniferous leaf litter sources. Relative to arctic tundra lakes, boreal forest lakes had higher nutrient concentrations, particularly total nitrogen (TN), likely due to warmer conditions, a longer growing season, and higher precipitation, which would enhance nutrient cycling and primary productivity. Results suggest that modern aquatic environments at opposite sides of the central Canadian arctic treeline (i.e. boreal forest and arctic tundra) exhibit distinct differences in water chemistry and physical conditions. These limnological trends may provide important information on possible future changes with additional warming.

Ecology and spatial distributions of surface-sediment diatoms from 77 lakes in the subarctic Canadian treeline region

Diatom ecology and species compositional patterns across current arctic treeline can provide important paleoecological information associated with climatic and environmental change. In this paper we examine the relationships between measured environmental variables and modern diatom assemblage composition from 77 lakes across the treeline ecozones of the Central Canadian Arctic. The weighted-average optima for selected environmental variables were calculated for 74 of the most common diatoms, and photographic plates of these taxa are included. Our results indicated that both forest-tundra and arctic tundra lakes differed significantly in diatom assemblage composition from boreal forest lakes. In general, planktonic diatom taxa (e.g., Cyclotella species) were more common in forested lakes, which may be due to ecological conditions related to climate. Small, benthic, alkaliphilic Fragilaria taxa reached their highest abundances in forested lakes, likely because of the more alkaline nature of these lakes. Arctic tundra lakes were characterized by higher abundances of circumneutral to acidophilic taxa. Heavily silicified Aulacoseira taxa (e.g., Aulacoseira lirata, Aulacoseira perglabra) were more common in deeper tundra lakes, likely because of the less alkaline nature of these lakes and greater wind-induced turbulence in this zone. These trends provide important information on the variability of aquatic ecosystems across this climatically sensitive vegetational gradient.Key words: arctic treeline, Canada, diatoms, paleolimnology, weighted-average optima, climate change.

Synchrony in lemming and vole populations in the Canadian Arctic

Population fluctuations may occur in synchrony among several rodent species at a given site, and they may occur in synchrony over large geographical areas. We summarize information on synchrony in lemmings and voles from the Canadian Arctic for the past 20 years. The most detailed available information is from the central Canadian Arctic, where snap-trap samples have been taken annually at several sites for periods of up to 15 years. Geographical synchrony in the same species among different sites was strong, especially for the central and eastern Canadian Arctic. Synchrony among different species at a given site was also generally high. When one species is at high density, densities of all species at that site tend to be high. These results do not easily fit the mobile-predator hypothesis proposed to explain regional synchrony, and are more consistent with the weather hypothesis, which we suggest both entrains synchrony among sites and enforces synchrony among species within a site. We tentatively support the weather hypothesis for geographical synchrony in lemmings, and recommend the establishment of a circumpolar program to monitor lemming cycles and predator movements that would advance our understanding of these large-scale patterns of cyclic synchrony.

Hierarchical habitat selection by barren-ground grizzly bears in the central Canadian Arctic.

Using resource selection functions, we examined habitat selection patterns of barren-ground grizzly bears (Ursus arctos) in the central Canadian Arctic among and within home ranges. There was no difference between the sexes with regard to habitat selection patterns at the home range level (Wilks' λ, approx. F 11,11=1.27, P=0.37). Bear home ranges contain more esker habitat, tussock/hummock successional tundra, lichen veneer, birch seep, and tall shrub riparian areas relative to the proportional availability of habitats in the study area. We observed differences in habitat selection within home ranges among levels of sex/reproductive status (Wilks' λ, approx. F 20,412=3.32, P<0.001) and by season (Wilks' λ, approx. F 30,605=2.71, P<0.001). Eskers and tall shrub riparian zones were the habitats most preferred by bears throughout the year. Tussock/hummock successional tundra was also favored by males at varying times during the year and lichen veneers were favored in spring and autumn by most bears. Females with cubs tended to avoid the highest ranked habitat for males throughout the year. This pattern of habitat selection was not observed for females without accompanying young. Results of this study underline the importance of scale dependence in habitat selection. Failure to view habitat selection as a hierarchical process may result in a narrow and possibly misleading notion of habitat selection patterns.