-A fossil roadrunner from interstadial deposits (ca. 33,590 BP) in New Mexico is identified as Geococcyx californianus. Fossil remains from California, dating from the subsequent late Wisconsinan stadial, have been assigned to the same taxon. However, approximately contemporaneous populations of larger-sized individuals occurring inland have been named G. conklingi. This taxon also has been recognized as surviving into the early Holocene before becoming extinct. This geographic and temporal pattern is hypothesized as being the result of local adaptations for temperature regulation: hot summer temperatures result in heat stress, resulting in selection against large individuals that otherwise would be best adapted for winter cold; absence of hot temperatures allows selection for large-sized individuals. The inland population of large-sized roadrunners, then, was a geographic and temporal subspecies, G. californianus conklingi. Conkling's Roadrunner (Geococcyx conklingi) was described by Howard (1931) from late Pleistocene deposits of Conkling Cavern, Dona Ana Co., New Mexico (Fig. 1). The species was characterized by size greater than that of the living roadrunner, Geococcyx californianus. Specimens from a nearby site (Shelter Cave) also were referred to this species (G. californianus also was reported, but some specimens definitely were modern and the remainder probably so). Since that time, G. conklingi has been reported from the late Pleistocene sites of San Josecito Cave, in southern Nuevo Leon, Mexico (L. Miller, 1943); and Dark Canyon Cave (Howard, 1971), southwest of Carlsbad, Eddy Co., New Mexico. Gehlbach and Holman (1974) reported its presence in Holocene deposits (<6,000 BP) of Pratt Cave in the Guadalupe Mountains of Culberson Co., Texas; Rea (1980) confirmed the identification, though it is slightly larger and somewhat different in characters from the type. Fossil G. californianus have been reported only from the late Pleistocene asphalt deposits of California: Rancho La Brea, Los Angeles Co.; Carpinteria, Santa Barbara Co.; and McKittrick, Kern Co. (Larson, 1930). Thus the literature indicates the presence of G. californianus in California at the same time as G. conklingi occurred inland, with the latter persisting well into the Holocene. In this paper, we add two pertinent records of fossil roadrunners and attempt to clarify the taxonomy and paleoecology of the