Fan-delta deposits, an important sedimentary component of fan-delta complexes, are scarcely described in the ancient record because it is very difficult to distinguish them from other types of coarse-grained deposits (e.g., alluvial, fluvial, or deep marine deposits). This obstacle is overcome when a well-record of physical/biogenic sedimentary structures or fossils unambiguously marks the influence of the receiving basin. However, this is usually not the case. Here, we present the Ophiomorpha ichnofabric record as a key proxy of marine influence within a generally fluvial-dominated system for improving the paleoenvironmental understanding and reconstructing animal-substrate interplay during the evolution of a fan-delta complex. The ichnological record, together with detailed sedimentological analysis, allows us to define three ichnofabric types, distributed in four levels. The stratigraphic distribution of these features points to distal alluvial environments towards the base, dominated by debris and sheet flows deposits and locally shallow braided channels; they evolve upward into well-developed braided fluvial system whose facies arrangement is related to channel fills, locally influenced by debris flows. Subsequently, a basinward migration of the sedimentary system is recorded through the occurrence of the first level with Ophiomorpha ichnofabric A, and then ichnofabrics A and B in levels 2 and 3, respectively, suggesting marine-influenced mouth bar settings. Toward the top, another steep basinward migration is evidenced by Ophiomorpha ichnofabric C in level 4, indicating wave-dominated processes in distal mouth bars environments. The record of marine-influenced and dominated levels may indicate relative sea-level fluctuations related to local or autogenic processes at the basin scale. Furthermore, the present study demonstrates the ability of Ophiomorpha tracemaker to colonize challenging sedimentary environments.
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