Bobwhite Habitat Research Articles

Sericea lespedeza control with postemergence and preemergence herbicide applications in fields managed for northern bobwhite

AbstractSericea lespedeza (Lespedeza cuneata) is an invasive nonnative legume that commonly occurs on sites managed for northern bobwhite (Colinus virginianus). Sericea reduces bobwhite habitat availability by outcompeting native plants that provide food and cover. Sericea can be controlled with postemergence herbicides such as glyphosate as well as a mixture of triclopyr with fluroxypyr, but these herbicides do not provide preemergence control. Imazapic has been effective at controlling a limited number of planted sericea seeds, but the efficacy of imazapic at various application rates to control sericea seedbank response following treatment with a postemergence herbicide has not been evaluated. We designed a field experiment to evaluate sericea control and changes in the plant community as related to bobwhite habitat with 2 postemergence herbicides followed by preemergence treatment of imazapic at 3 sites in Tennessee and Alabama, USA, 2018–2022. Specifically, we treated sericea with postemergence broadcast applications of glyphosate or triclopyr with fluroxypyr in 2018 and applied spot treatment with the same herbicides in 2019–2021. We applied imazapic at 4 rates following prescribed fire in 2019 and 2022. We measured coverage of sericea, native forbs, annual grasses, perennial grasses, and bobwhite food plants before treatment in 2018 and after all treatments in 2022. Nearly all postemergence treatments reduced sericea coverage, but imazapic did not increase control compared to postemergence herbicides alone. Perennial grasses used for nesting and bobwhite foraging were maintained following treatments. We recommend managers use either glyphosate or triclopyr with fluroxypyr to control sericea postemergence and consider using a low rate of imazapic to reduce annual grasses if coverage is a problem on the site.

Effects of Native and Non-Native Invasive Grasses on Northern Bobwhite Habitat in South Texas

Usable space for northern bobwhite (Colinus virginianus) has been reduced across South Texas rangelands because of the spread of non-native, invasive grasses. A native grass, tanglehead (Heteropogon contortus), has also rapidly expanded its dominance in South Texas within the past 25 yr. It has formed high-density monocultures, similar to non-native grasses, which are associated with losses of herbaceous diversity and bare ground, both key components of bobwhite habitat. The objectives of our research were to 1) determine selection-avoidance of invasive grass cover by bobwhites and 2) determine effects of invasive grass cover on vegetation characteristics.We hypothesized the effects of tanglehead on bobwhite would be negative and similar to the effect of non-native grasses in the region. We recorded covey locations via helicopter surveys in December 2014 and 2015, with field measurements of bobwhite habitat at covey and random locations taken each spring. We measured grass and forb species richness, herbaceous vegetation height, woody-plant cover, tanglehead canopy cover, and non-native grass canopy cover at all covey detections and an equal number of random locations. We developed continuous selection ratios using Simple Saddlepoint Approximations to determine habitat selection by bobwhites. We also used quantile regression, at the 10th, 50th, and 90th quantiles, to determine relationships between invasive grass cover and vegetation factors. Bobwhites avoided areas of high canopy cover (> 20%) of all invasive grasses measured. We found negative relationships between native and non-native invasive grass cover and forb and grass species richness, as well as bare ground, and a positive relationship with herbaceous vegetation height at all quantiles modeled. Our results demonstrate the negative effects of increased tanglehead cover on native rangeland habitats, which aligns with similar non-native invasive grasses. Further expansion by tanglehead has the potential to significantly reduce usable space for bobwhites in South Texas.

A maximum entropy and GIS approach to predict potential habitat for northern bobwhites in the Black Belt prairie physiographic region of Alabama

Northern bobwhite (Colinus virginianus) populations have sustained precipitous declines throughout their range. The North American Breeding Bird Survey for Alabama indicates a rate of decline of over 6% per year since 1966. Declining bobwhite populations have been associated with large-scale habitat loss and a reduction in habitat quality due to advances in ecological succession, intensive monoculture agriculture, and intensive timber management practices. Once a very diverse region, modern land-use practices throughout the Black Belt Prairie physiographic region have created a simplified landscape that have left few places where bobwhites can live and thrive. Despite extensive research on bobwhites, there is insufficient knowledge on the spatial distribution of bobwhite habitat on a regional scale, including the Black Belt Prairie physiographic region. Knowing the spatial distribution of suitable bobwhite habitat will allow conservation planners to prioritize efforts and direct limited resources to achieve conservation goals for the region. The goals of this study were to 1) evaluate the maximum entropy modeling approach for predicting northern bobwhite habitat in the Black Belt Prairie physiographic region, and 2) develop a northern bobwhite habitat suitability map for the Black Belt Prairie physiographic region. The results showed that deciduous 1.8 km (1.8 km is the dispersal distance of bobwhites) neighborhood was the most important habitat variable, followed by water 1.8 km neighborhood. Developed 1.8 km neighborhood was the most important anthropogenic variable, ranking third in overall importance. The resultant model denotes potential habitat in the region and forms the basis for using the maximum entropy and geographic information system approach to predict suitable habitat and prioritize areas for conservation.

Examining a Habitat-Weather Threshold for Northern Bobwhite Populations in the Southwestern United States

In semiarid portions of the northern bobwhite (Colinus virginianus; hereafter, bobwhite) geographic distribution, weather is a strong driver of interannual abundance. However, the strength of this relationship may depend on habitat amount. Given this habitat–weather dependence, there is likely to be a threshold value for habitat that determines how strongly a bobwhite population responds to weather. Our objective was to evaluate the relationship between habitat amount and the relative influence of weather on bobwhite abundance in Texas and Oklahoma, USA and determine a potential land-cover threshold value. We collected bobwhite abundance and land-cover data from the Breeding Bird Survey and National Land Cover Database, respectively, for 2 time periods which corresponded to historical lows (2012–2013) and highs (2015–2016) of bobwhite abundance within this region. We reclassified land cover into grassland, shrubland, pastureland, cropland, forest, and urban cover, and combined grassland and shrubland categories to represent bobwhite habitat. We used weighted linear regression to model the difference between mean bobwhite abundance between each time period as a function of habitat amount, hypothesizing a positive, linear relationship given the theorized greater influence of weather on populations within higher habitat amounts. To evaluate a potential habitat threshold, we modeled mean bobwhite abundance, for each time period, and all individual land-cover variables using a generalized additive model with a negative binomial distribution. We detected a positive, linear relationship between habitat amount and the difference between high and low bobwhite populations (r2 = 0.30), per our hypothesis. Models of land-cover variables differed between low- and high-abundance time periods. The time period of lower abundance (2012–2013) model included all land-cover variables except cropland and showed significant positive, linear relationships with grass and hay cover, and negative, linear relationships with forest and urban cover. Shrubland cover was the only nonlinear term, increasing to approximately 50–60% cover, then decreasing sharply with increasing cover. Deviance explained for this model was 41.2%. For the period of high abundance (2015–2016) the model explained approximately 25% of the deviance and was reduced to only negative, linear relationships with forest and urban cover. Our results provide some evidence toward the habitat–weather threshold hypothesis. Additionally, while no land-cover values showed a threshold-like relationship with bobwhite abundance, we did document differences in variable selection for each time period. Stressors on bobwhite habitat seem to have a more dramatic impact on bobwhite populations than available habitat, especially in periods of higher abundance, whereas periods of lower

Home Range and Space Use of Northern Bobwhite Under Two Different Management Models in Southwestern Missouri

Northern bobwhite (Colinus virginianus; hereafter, bobwhite) management in Missouri, USA has traditionally been focused on providing an interspersion of grass, crop, old field, and woody cover juxtaposed to disked idle areas and food plots to maintain bobwhite populations. This traditional model is implemented with the goal of providing all essential habitat components within 40-acre blocks throughout a larger area used by a population. While this model can produce usable bobwhite space in agriculture-dominated landscapes, it may not be the most effective or efficient approach to producing and maintaining bobwhite in grassland-dominated landscapes. In southwestern Missouri native tallgrass prairie conservation areas are managed primarily with historical ecological processes, such as fire and grazing, to produce the desired patchy habitat mosaic. Additionally, it has been on these native tallgrass prairie conservation areas that managers have seen the most stable and productive bobwhite populations. Over a 5-year period (2014–2018) we quantified movements of northern bobwhite on 3 traditionally managed areas (n = 185) and on 3 managed native tallgrass prairie conservation areas (n = 211) to determine whether home range sizes and space use differed between these two management models. We used the 6-month (Apr–Sep) breeding period to determine core area, home range, mean movement rate, and maximum distance moved. Overall (pooled) home ranges of bobwhite did not differ significantly between traditional and grassland managed areas; however, there were significant yearly differences between management models and study areas. Males generally had larger home range sizes and had higher movement rates than females. For the 5 years of the study few birds made long-distance movements (>1.6 km; 3%), and all remained relatively close to capture locations in winter (Feb–Mar). For both traditional and grassland managed areas, bobwhite selected for areas that had disturbance (fire and grazing) in the last 2 years and for the native grassland vegetation type. These findings suggest that areas managed under the grassland management model provide preferred habitat for bobwhite and could result in significant improvement in habitat quality for tallgrass prairie wildlife.

Northern Bobwhite Response to Vegetation Management and Recovery in South Texas

The northern bobwhite (Colinus virginianus; hereafter, bobwhite) requires habitat structure and composition with grass cover for nesting, predator avoidance, and thermal refuge and forb cover for feeding on phytophagous arthropods and seeds. During the past 2 decades, many land managers with interest in promoting quail hunting opportunities have reduced or completely eliminated livestock across South Texas, USA, rangelands. Resting the land from grazing allows vegetation—especially grasses and forbs—to recover and thus provide nesting and foraging habitat for bobwhite and other birds. How bobwhite respond to postgrazing vegetation recovery is of keen interest to rangeland quail managers, but this topic is poorly known because case histories with quantitative data are lacking. Our objective was to investigate how bobwhite respond to the vegetation changes following removal of cattle grazing. Our study was conducted on a private ranch in Jim Hogg County, Texas and involved 3 different categories of postgrazing recovery and management: 1 area at 15 years post-cattle grazing where the landscape has been brush-sculpted and is actively managed since removal of cattle, 1 area at 3 years post-grazing with a previous stocking density of 7 ha/animal unit (high stocking density), and 1 area at 3 years post-grazing with a previous stocking density of 14 ha/animal unit (moderate stocking density). We trapped, radio-marked, and located bobwhites from March to September during 2015–2016 on the 3 comparative units. We estimated nest survival, adult breeding-season survival, home range size, and early winter density. We hypothesized that the 15-year postgrazing site would have higher early winter density, higher adult breeding season survival, and higher nest survival along with smaller mean home range size compared to the 2 more recently grazed sites. On average, the probability of a nest surviving the 23-day incubation period was highest on the 15-year postgrazing site at 0.61 ± 0.12 (mean nest survival ± standard error), with estimates of 0.32 ± 0.12 on the 3-year moderate postgrazing site and 0.33 ± 0.12 on the 3-year high postgrazing site. Adult breeding season survival did not differ among the 3 sites, and was instead influenced mostly by month within the season, probably a result of summer heat. An adult bobwhite had a 0.48 ± 0.04 probability of surviving the breeding season. Early winter density, after summer and fall production was complete, increased on all sites from 2015 to 2016 and was consistently highest on the 15-year postgrazing site. Home range sizes on the

Habitat Works: How Partnerships and Habitat Improvement have Restored Quail Populations in the 2C Quail Focus Area

The Missouri Department of Conservation (MDC) began establishing Quail Focus Areas (QFAs) on private lands in 2004. The goal of QFAs was to bring groups of landowners together to manage bobwhite habitat on a larger scale in a targeted landscape. Through a variety of state, federal, and other partnership programs, habitat improvement efforts have resulted in large increases in northern bobwhite (Colinus virginianus; hereafter, bobwhite) numbers in the 2C QFA. In spring 2013, MDC staff and Quail Forever biologists began monitoring bobwhite and songbirds in a portion of the 2C QFA in Carroll County, Missouri, USA and in a control area (without habitat management for bobwhite). This effort is part of the Coordinated Implementation Plan developed by the National Bobwhite Technical Committee as a part of the National Bobwhite Conservation Initiative. The goal of the monitoring plan is to document whether quail habitat management can achieve sustainable bobwhite populations within 5–10 years. We selected a 2,100-ha portion of the 2C QFA where habitat management for quail has been conducted through efforts by landowners, MDC staff, and Quail Forever volunteers. Point-transect surveys were conducted at 48 250-m radius points in spring for bobwhite and songbirds and at 12 500-m radius points in fall for bobwhite coveys. Quail densities ranged from 0.18 quail/ha (95% credible interval [CrI] = 0.09–0.32) to 0.41 quail/ha (95% CrI = 0.30–0.57) in the focus area and from 0.04 quail/ha (95% CrI = 0.01–0.11) to 0.12 quail/ha (95% CrI = 0.06–0.20) in the control area from 2013–2019. We have also documented increases in grassland songbirds through monitoring efforts. Results show that dedicated management efforts were successful in increasing bobwhite density in the focus area.

Winter Survival and Resource Use of Translocated Northern Bobwhite in the Mid-Atlantic United States

Northern bobwhite (Colinus virginianus; hereafter, bobwhite) in the Mid-Atlantic United States have been experiencing precipitous population declines due to a combination of habitat deterioration, urban and suburban sprawl, change in forest management regimes, and farming practices. In recent years, restocking of bobwhite through translocation efforts has gained interest to rebuild local populations. However, empirical studies are warranted to understand the limitations of translocation as it relates to its potential use for long-term population recovery and persistence in this region. Further, few studies nation-wide have evaluated resource use and survival during the non-breeding season on translocated sites. As such, we translocated 360 bobwhites from source populations from southern latitudes during March–April 2015–2017 into 2 different landscape types similar to the source population vegetative communities (agricultural cropland dominated in Maryland, USA versus pine forest dominated in New Jersey, USA) and tested the effects of habitat fragmentation on survival and habitat use during the subsequent non-breeding season. We found habitat fragmentation negatively affecting survival and resource use among translocated bobwhite on fragmented cropland-dominated sites as compared to larger unfragmented forested sites. Survival was lower on cropland sites compared to forested sites such that bobwhite in cropland-dominated landscapes were >125 times less likely to survive the winter than those on forested sites. In our examination of resource use, bobwhite in structurally complex forested sites used cut pine, early-successional woody, early-successional herbaceous, and thinned pine more than what was available on the landscape. On the cropland sites bobwhite used food plots, early-successional woody, and mixed woods more than what was available on the landscape and only food plots at the home range scale. While larger unfragmented forested bobwhite habitat ultimately provided a more successful translocation landscape, birds still had large home ranges and relatively low survival. Therefore, proper pine management may be necessary to optimize habitat availability during the non-breeding season. Our findings provide rare information on demographic resiliency and resource use for translocated bobwhite during the non-breeding season. Further, this research provides valuable information to improve future translocation efforts in the Mid-Atlantic.

Bobwhite Response to Cattle Grazing in South Texas

Range management practices to improve habitat for wildlife by reducing brush and increasing herbaceous plants, coupled with reduced stocking rates, can lead to dense stands of dominant grasses, such as four-flower trichloris (Trichloris pluriflora). This monoculture of trichloris creates dense vegetation unsuitable for northern bobwhite (Colinus virginianus; hereafter, bobwhite), reduces plant species diversity, and alters ecosystem functions. The objectives of this study are to 1) evaluate the effects of a proper cattle grazing regime to improve bobwhite habitat and 2) develop a management guide documenting how cattle grazing can be used as a tool to reduce the density and cover of dominant grasses and thereby allow higher plant species richness. The study is taking place in Duval County, Texas, USA, between 2 pastures with a combined area of 2,500 ha. One pasture serves as the control (1,337 ha) while the other (1,109 ha) is grazed to maintain a stubble height of 30–40 cm. We placed 10 grazing exclosures and 10 25-m transects within each treatment to determine botanical composition and cover. Double sampling is conducted monthly to determine forage standing crop. Forage standing crop, plant species richness, total ground cover, and forage utilization will be estimated. We hypothesize that 1) the grazed pasture will contain more bobwhites than the nongrazed pasture, 2) plant species richness will be greater in the grazed pasture, and 3) grazing will reduce the cover of trichloris. In 2020, plant species richness varied among sampling periods and was recorded as 7.5 species/transect and 4.8 species/transect higher in the nongrazed control compared to the grazed pasture in June and August, respectively. After the end of the first year of grazing, however, there was no statistical difference in plant species richness between the 2 pastures, a result that does not coincide with our second hypothesis. Litter cover did not vary among sampling periods in the nongrazed pasture but changed in the grazed pasture. Litter cover was 9.4%, 14.3%, 14.6%, and 8.9% higher in the grazed pasture than in the nongrazed pasture in May, August, September, and December 2020, respectively. Bare ground cover changed throughout sampling periods in the nongrazed and grazed pastures. Bare ground cover was 22%, 18%, and 22% higher in the grazed pasture than in the nongrazed pasture in May, September, and December 2020, respectively. Trichloris cover did not change throughout the sampling period in the grazed pasture but varied in the nongrazed pasture. Trichloris cover

Multisensor models for assessing recurrent fire compatibility with habitat recovery for a critically endangered species

The increased variety and availability of remotely sensed data from satellite and airborne platforms are expected to enhance data fusion approaches aimed at characterizing wildlife habitat. We investigated multisensor machine learning (ML) models to estimate desert grassland habitat suitability for a critically endangered species, the mask bobwhite quail (Colinus virginianus ridgewayi). Contemporary habitat conditions are marked by a history of intensive livestock grazing, periodic drought, and changes in arid land hydrology. Historical land use and vegetation changes were accompanied by altered disturbance regimes that once supported recurrent fire important to maintaining grassland composition and structure. Habitat assessment plots were combined with multidate and multispectral Worldview-3 (WV3) satellite imagery and airborne light detection and ranging (LiDAR) metrics to model habitat suitability on the Buenos Aires National Wildlife Refuge (BANWR). We used regression and structural equation models (SEM) to assess fire history, climate, and site biophysical effects on habitat suitability from 239 vegetation plots stratified by topography and fire frequency. We found that combined WV3/LiDAR models improved suitability predictions and consistency over WV3-only models comparing the root mean squared and mean absolute error from training data. Hyperparameter tuning for ML habitat models further improved performance. Gradient boosted regression trees and WV3/LiDAR habitat suitability models showed superior performance and correspondence to independent field validation data (F = 92.7, p ≤ 0.001, R2 = 0.57). Comparisons between predicted habitat suitability and fire history variables from 84 management units showed a significant negative relationship with increased fire frequency (F = 27.6, p ≤0.001, R2 = 0.41) and positive relationship with time since last burn (F = 41.3, p ≤0.001, R2 = 0.34). SEMs established that masked bobwhite habitat occurrence was strongly associated with site biophysical conditions that supported greater woody plant cover important for nesting, thermal, and predator protection. Our results suggest that desert grasslands associated with low primary productivity may require recovery periods in excess of 20 years to develop a desired mix of herbaceous and woody plant cover for masked bobwhite, when ≥2 fires have occurred over the last 30 years. Multiple sensor types provided a unique set of variables describing horizontal and vertical habitat structure and composition, essential to understanding fire effects on masked bobwhite habitat suitability. Combined information from active and passive remote sensing systems can likely enhance mapping and monitoring applications necessary for assessing recovery needs for numerous other wildlife species with diverse habitat requirements.

Northern Bobwhite Occupancy Patterns on Multiple Spatial Scales Across Arkansas

AbstractNorthern bobwhite Colinus virginianus populations have been rapidly declining in the eastern, central, and southern United States for decades. Land use change and an incompatibility between northern bobwhite resource needs and human land use practices have driven declines. Here, we applied occupancy analyses on two spatial scales (state level and ecoregion level) to more than 5,000 northern bobwhite surveys conducted over 6 y across the entire state of Arkansas to explore patterns in occupancy and land use variables, and to identify priority areas for management and conservation. At the state level, northern bobwhite occupied 29% of sites and northern bobwhite were most likely to occur in areas with a high percentage of early successional habitat (grassland, pasture, and shrubland). The statewide model predicted that northern bobwhite were likely to occur (≥ 75% predicted occupancy) in < 20% of the state. Arkansas is comprised of five distinct ecoregions, and analyses at the ecoregion spatial scale showed that habitat associations of northern bobwhite could vary between ecoregions. For example, early successional habitat best predicted northern bobwhite occupancy in both the Arkansas River Valley and Ozark Mountains ecoregions, and other habitat associations such as the proportion of herbaceous habitat and hay-pasture habitat, respectively, further refined predictions. Contrastingly, richness of land cover classes alone best predicted northern bobwhite occupancy in the Ouachita Mountains ecoregion. Ecoregion-level models were thus more discerning than the state-level model and should be more helpful to managers in identifying priority conservation areas. However, in two of five ecoregions, surveys too rarely encountered northern bobwhite to accurately predict their occurrence. We found that likely occupied northern bobwhite habitat lay primarily on private properties (95%), but that numerous public entities own and manage land identified as suitable or likely occupied. We conclude that management of northern bobwhite in Arkansas could benefit from cooperation among state, federal, and military partners, as well as surrounding private landowners and that ecoregion-specific models may be more useful in identifying priority areas for management. Our approach incorporates multiple landscape scales when using remote sensing technology in conjunction with monitoring data and could have important application for the management of northern bobwhite and other grassland bird species.

Effects of Prescribed Fire on Northern Bobwhite Nesting Ecology

ABSTRACTRepeated prescribed fire can create and maintain areas with sparse overstory tree cover and a dense grass‐forb‐shrub understory, providing habitat for northern bobwhite (Colinus virginianus; hereafter, bobwhite). Despite potential benefits of prescribed fires for conserving bobwhite habitat, burning during the nesting season may destroy bobwhite nests and reduce available nesting cover. We monitored radio‐transmittered bobwhite (n = 104) from 2016 to 2018 to describe nest‐site selection and determine the risk of nest destruction on a 17,000‐ha North Carolina military installation, Fort Bragg, managed with rotational growing‐season and dormant‐season prescribed fires on an approximate 3‐year return interval. We located 48 nests, of which 8 (16%) were in areas burned the same year, 9 (19%) were in one‐year post fire, 25 (52%) were in 2‐years post fire, and 6 (13%) were in ≥3‐years post fire areas. We compared vegetation composition and structure at nests to nearby random locations and determined bobwhite selected nest sites with greater woody understory and wiregrass cover, lower basal areas of pines and hardwoods, and less distance to the nearest road. Two nests (6.7%) were destroyed during prescribed fires, but success of incubated nests was high (67%). We calculated the overall risk of nest destruction by prescribed fire as the proportion of active nests in areas with ≥3 years since last fire multiplied by the proportion of the study area burned each week. Overall, 11% (weekly = 0.75%, range = 0–3%) of the study area was burned during the 2016 nesting season (3 June to 3 September), 4% (weekly = 0.31%, range 0–2%) of the study area was burned during the 2017 nesting season (5 June to 2 September), and 7.5% (weekly = 0.58%, range 0–5%) of the study area was burned during the 2018 nesting season (3 June to 31 August). We estimated that no more than 0.75% of bobwhite nests across the study site were exposed to fire annually. Most growing‐season fires occurred before the bobwhite nesting season, which limited direct effects of prescribed fire on bobwhite nest survival. However, shifting prescribed fires to later in the growing season to better match the historical lightning season (i.e., after 1 June) would increase the risk of nest destruction. Because bobwhite used older roughs (i.e. areas 2 years since fire) for nesting, shortening the fire return interval to less than 3 years would increase the proportion of nests exposed to fire. Additionally, a shortened fire return interval would decrease available nesting cover, especially in regions with low soil fertility where vegetation change following fire is less rapid than on more productive soils. © 2021 The Wildlife Society.

High-intensity short-duration grazing during spring is not an effective habitat management tool for Northern Bobwhites in Colorado

Abstract Many wildlife species, like Northern Bobwhite (Colinus virginianus), are reliant on periodic environmental disturbance to maintain heterogeneity in vegetation patterns. The Northern Bobwhite is a species of conservation concern requiring different vegetation types for nesting and brood-rearing stages, as well as for protection from predators and extreme weather. In northeastern Colorado, there are few management options to increase disturbance, other than domestic livestock grazing, to manage Northern Bobwhite habitat. As high-intensity short-duration grazing has shown promise as a mode of disturbance to manage Northern Bobwhite habitat in other regions, I used a randomized block design to test the effect of spring high-intensity short-duration grazing on Northern Bobwhites and their habitat in northeastern Colorado from 2016 to 2019. Specifically, I monitored Northern Bobwhite nest and brood survival and habitat selection in relation to grazing treatments over 3 years. I found that grazing had no effect on nest or brood survival or brood habitat selection, but Northern Bobwhites selected against grazed plots for nesting. Nest survival was negatively influenced by percent litter around the nest, and Northern Bobwhites selected nest sites with more grass cover and less bare ground. Broods selected habitat with less bare ground and more woody vegetation. Grazing affected vegetation immediately after grazing, but these effects weakened or disappeared by the end of the growing season. One exception to this overall pattern was forbs, which tended to be more abundant on grazed plots throughout the growing season. Overall, I found neutral effects of grazing on Northern Bobwhite nest and brood survival and habitat selection and neutral to positive benefits to the vegetation. Spring high-intensity short-duration grazing does not appear to be an effective tool to manage Northern Bobwhite nest or brood habitat in northeastern Colorado.

Efficacy of acclimating and releasing captive-reared and wild-translocated Northern bobwhites

Northern bobwhite populations are dwindling across their range. Accordingly, in areas with bobwhite habitat restoration and no nearby populations, effective population restoration techniques are needed. Here, we evaluated three bobwhite restoration release strategies: (1) release of captive-reared bobwhites on sites with and without resident populations, (2) translocation of wild-trapped bobwhites from a region of high population density to a region with no population, and (3) release of captive-reared and wild-translocated bobwhites acclimated on site prior to release in year 2. Wild-translocated birds survived longer than captive-reared birds. Mean survival for captive-reared bobwhites was 2.42 weeks, and 4.27 weeks in year 1, and 1.91 and 1.40 weeks in year 2, for study sites without resident and with resident birds present, respectively. Mean survival for wild-translocated birds was 8.50 weeks in year 1, and 11.54 weeks in year 2. Wild-translocated birds dispersed shorter distances than captive-reared birds. Both captive-reared and wild-translocated bobwhites only nested on study sites with conspecifics. Captive-reared birds had 0 nesting attempts on the site with no resident bobwhites, and ⩾8 nests on the site with resident bobwhites. Wild-translocated females nested six times and were subsequently observed with juveniles. On-site acclimation did not increase post-release survival for northern bobwhites. Acclimation increased site fidelity but reduced survival for captive-reared birds and had no impact on survival for wild-translocated bobwhites. Population restoration by release of captive-reared or wild-translocated birds is not irrelevant, but further investigation is needed into the relationship between captive-reared birds and predators, and methods to increase survival and reproduction of released birds.

Northern Bobwhite Non‐Breeding Habitat Selection in a Longleaf Pine Woodland

ABSTRACTEfforts to halt the decline of the northern bobwhite (Colinus virginianus; bobwhite) across its distribution have had limited success. Understanding bobwhite habitat requirements across the annual cycle and at varying scales is essential to aid efforts to conserve bobwhites. We monitored radio‐tagged bobwhites from 2016 to 2018 on a 165‐km2 portion of Fort Bragg Military Installation in the Sandhills physiographic region of North Carolina, USA, to determine factors influencing non‐breeding bobwhite habitat selection at multiple scales. We used generalized linear models (GLM) and generalized linear mixed models to assess bobwhite habitat selection at the microsite scale (the immediate vicinity of an animal) and the macrosite scale (across the study area), respectively, by comparing used points to available random points. At the microsite scale, bobwhites strongly selected areas with greater woody understory cover. Also, bobwhite selection increased with greater forb and switchcane (Arundinaria tecta) cover, but this effect plateaued at 65% forb cover and 50% switchcane cover. At the macrosite scale, bobwhites generally selected areas with greater understory cover within a 200‐m radius but avoided areas with >55% understory cover; these areas primarily were located in the core areas of drainages with extensive ericaceous vegetation. Bobwhites selected areas with 3–6 m2/ha hardwood basal area in uplands, potentially because of the availability of mast, but avoided uplands when pine (Pinus spp.) or hardwood basal area exceeded 20 m2/ha or 12 m2/ha, respectively, likely because high basal area is associated with increased shading and subsequent loss of understory cover. In addition, bobwhites selected uplands 1 growing season (≥2‐month period falling entirely between 1 Apr and 1 Oct) post‐fire regardless of burn season. Overall, managers seeking to improve habitat quality for bobwhites in longleaf pine (Pinus palustris) woodlands should employ management practices that maintain available woody understory across the landscape to provide cover during the non‐breeding season. © 2020 The Wildlife Society.

Multi‐attribute preferences for northern bobwhite habitat restoration among Texas landowners

ABSTRACTNorthern bobwhite (Colinus virginianus) abundance has declined range‐wide over the long term due to factors such as habitat loss and deterioration. Private land management is critical to bobwhite conservation because most bobwhites occur on private lands, but little research has been conducted regarding how private land managers and landowners make decisions about whether and how to restore northern bobwhite habitat. To begin addressing this deficit, we used a choice‐based conjoint approach to determine how Texas, USA, landowners and land managers (surveyed in November 2017) weigh the importance of northern bobwhite hunting opportunities, costs, and labor when making habitat restoration decisions. We also used a latent class analysis to break respondents into segments based on their valuations of these attributes. The results of our latent class analysis indicate that managers and landowners can be grouped into several segments, and most segments do not place a high value on bobwhite hunting opportunities, but instead are more interested in minimizing out‐of‐pocket costs and labor input. Bobwhite habitat restoration programs yielding cost shares ≥50%, labor inputs at ≤30 days/year, and likely coveys flushed per hunting day >4 broadly appealed to landowners; however, ensuring low labor costs was most important, particularly for pragmatic landowners. This indicates that policies minimizing labor inputs and maximizing cost shares may be preferred over increasing hunting opportunities. Finally, we discuss the management preferences and practices of the segment most interested in maximizing bobwhite hunting opportunities and potential outreach strategies for segments of opportunistic and pragmatic landowners who may be persuaded to manage for bobwhite habitat if they perceived this management as economically viable. © 2019 The Authors. Wildlife Society Bulletin Published by Wiley Periodicals, Inc.

Northern bobwhite habitat use in a food subsidized pyric landscape

ABSTRACTAnimals inhabiting disturbance‐prone systems, such as the northern bobwhites (Colinus virginianus) in pine (Pinus spp.) savanna, are adapted to certain intensities of disturbance (e.g., frequency, spatial extent, seasonality). Management practices attempt to mimic these natural conditions. Even though northern bobwhites are known as the firebird, most fire‐related management is currently based on tradition and limited peer‐reviewed literature. We studied habitat selection of northern bobwhites on private property in Georgetown County, South Carolina, USA, managed with fire, winter disking, and supplemental feeding. We radio‐tagged 338 individual bobwhites and monitored them 3–4 times weekly via very high frequency (VHF) telemetry for 2 years. We used hierarchical habitat selection functions in a Bayesian framework to model the data. We considered 2 spatial scales: the study site (second‐order) and within home ranges (third‐order). Bobwhites selected for small burns during the breeding season but had the highest selection for areas intermediately distant from burn edges during winter (i.e., 67 m). Bobwhites had the strongest selection for fire return intervals of 2–3 years during winter but 1–2 years during the breeding season. Use of supplemental feed was strong across seasons but not selected for by brooding birds within their home range. Use of fallow fields was strongest for brooding birds. Our results are useful for bobwhite managers, especially those in subtropical climates, because they provide scientifically defensible information supporting the use of prescribed fire, winter disking, and supplemental feeding in the context of habitat use. © 2017 The Wildlife Society.

A County-Based Northern Bobwhite Habitat Prioritization Model for Kentucky

Planning the management of northern bobwhite (Colinus virginianus) habitat at a statewide-scale is daunting. Native grassland restoration is difficult to manage in Kentucky because . 99% of the Commonwealth’s original grassland area has been lost to agriculture, succession, and development. We created a county prioritization model designed to target areas of grasslands and landowners likely to participate in conservation programs. Our goal was to identify 10% of the state as high priority for bobwhite habitat restoration. We created an east and west model divided by the Appalachian Mountains. The west model was designed to target production-oriented operators farming marginal lands, whereas the east model targeted reclaimed minelands. We used agricultural, landcover, and staff data to build county prioritization models in 2006 and 2011. The models targeted 16.6% and 17.6% of the state in 2006 and 2011, respectively. However, if areas of large, contiguous blocks of forests were excluded, the area total was much closer to 10%. Fifty percent of the high priority counties changed in the west model, and 33% of the counties in the east model changed over 5 years. Implementing a county prioritization model in conjunction with a finer-scale, biological targeted approach could focus conservation efforts with greater potential for success, but the models should be reconstructed at 5- to 10-year intervals to account for changes in conservation delivery potential. A modification of our technique may serve to validate or as an alternative to improve National Bobwhite Conservation Initiative 2.0.

Sustaining the 'Quail Wave' in the Southern Great Plains

The southern Great Plains (i.e., Texas and Oklahoma) historically affords some of the best, and currently most stable, northern bobwhite (Colinus virginianus) populations anywhere. However, bobwhite populations have declined in recent years over much of this area, especially east of the 98th meridian. Two subsets of the southern plains, the Rolling Plains (parts of northwestern Texas, western Oklahoma, and western Kansas) and the Rio Grande Plains (south Texas) offer the most expansive, contemporary northern bobwhite habitat throughout its range. Bobwhite habitat in the southern plains is affected primarily by rainfall and rangeland management for livestock. Range management practices (brush control, grazing management) can be prescribed to benefit bobwhite habitat, but a large part of potential bobwhite range in the southern plains suffers from overgrazing and excessive brush control. Farm Bill policies (e.g., Conservation Reserve Program) have had a major impact on dryland agriculture in this region, but their impacts on bobwhites have been only marginally positive (if at all) to date. Income generated from quail hunting in this region currently rivals or exceeds that generated from cattle grazing leases. Accordingly, more landowners are beginning to temper traditional land management goals, and incorporate more quail-friendly practices (i.e., ‘‘brush sculpting’’ and reducing stocking rates). Educational efforts aimed at landowners should strive to implement existing knowledge and develop informed decision-makers. The current demand for quail hunting affords an excellent opportunity to promote, and adopt, management practices that will hopefully sustain the heritage of quail hunting in this region of the bobwhite’s range for future generations.

Contribution of CRP to Illinois Bobwhite Habitat at the Landscape Level (Poster Abstract)

Northern bobwhite (Colinus virginianus) population declines in the midwest have been attributed to habitat degradation and loss due mainly to intensified agricultural land use and farming practices. Thus, there was initial optimism that the Conservation Reserve Program (CRP) would benefit bobwhites by converting cropland to semi-permanent grassland. However, CRP apparently has not positively impacted regional or statewide population trends in Illinois. Deficiencies at both site and landscape level may be involved. To address the latter issue, we mapped the location of each individual CRP field (> 8,800) in 11 representative counties within the bobwhite range in Illinois. We then analyzed their spatial relationship to other land cover and bobwhite habitat using Geographic Information Systems and a statewide digital land cover map based on classified satellite imagery. Existing bobwhite habitat and CRP fields tended to be spatially correlated because both are largely confined to moderately rolling terrain as opposed to very hilly or very flat areas. Nevertheless, almost 25% of all existing CRP acreage within our 11 study counties occurred in landscapes with insufficient woody edge to support high bobwhite populations. Furthermore, CRP did not always provide the habitat component most limiting for bobwhites.