Evolution Of Behavior Research Articles

Evolution of hydrogen isotopes retention behavior of in-situ boronization films in EAST

Effective management of hydrogen isotopes retention in plasma-facing materials (PFMs) is crucial, particularly when utilizing tritium (T) as fuel, for the success of burning plasma operations. Boronization, a widely employed technique for controlling fuel recycling and mitigating impurity influx from plasma-surface interactions into the core of burning plasma, significantly influences hydrogen isotopes retention in PFMs. In this study, boronization films were generated by ion cyclotron range of frequency (ICRF) discharge assisted boronization with C2B10H12 as boron source on tungsten substrates at Experimental Advanced Superconducting Tokamak (EAST) which employing ITER-like water-cooled W monoblock PFMs and components (PFCs), followed by in-situ glow discharge (GD) cleaning for 2 h and 20 shots (180 s) edge-plasma exposure. Employing the Material and Plasma Evaluation System (MAPES), representative samples were analyzed after each process. The resultant carbon–boron films, dense and continuous, exhibited thickness up to 120 nm and were identified as amorphous in structure. It was observed that the D2-GD cleaning resulted in a significant isotopic exchange effect, effectively reducing the hydrogen (H) retention in the carbon–boron films. This hydrogen isotope replacement efficiency was found to be influenced by the thickness of the films. Notably, after boundary plasma exposure, samples with thicker films demonstrated an enhanced capacity to capture deuterium (D), adsorbing 10 times more D than bare tungsten (W). Our findings offer transformative insights for T recycling analysis and the plasma operation of devices like International Thermonuclear Experimental Reactor (ITER), highlighting the impact of boronization and subsequent treatments on hydrogen isotope retention behavior in PFMs.

Evolutionarily stable payoff matrix in hawk–dove games

BackgroundClassical matrix game models aim to find the endpoint of behavioural evolution for a set of fixed possible interaction outcomes. Here, we introduce an evolutionary model in which not only the players’ strategies but also the payoff matrix evolves according to natural selection.ResultsWe start out from the hawk–dove matrix game and, in a way that is consistent with the monomorphic model setup of Maynard Smith and Price, introduce an evolving phenotypic trait that quantifies fighting ability and determines the probability of winning and the cost of losing escalated hawk–hawk fights. We define evolutionarily stable phenotypes as consisting of an evolutionarily stable strategy and an evolutionarily stable trait, which in turn describes a corresponding evolutionarily stable payoff matrix.ConclusionsWe find that the maximal possible cost of escalating fights remains constant during evolution assuming a separation in the time scales of fast behavioural and slow trait selection, despite the fact that the final evolutionarily stable phenotype maximizes the payoff of hawk–hawk fights. Our results mirror the dual nature of Darwinian evolution whereby the criteria of evolutionary success, as well as the successful phenotypes themselves, are a product of natural selection.

The Jojosi Dongas: An interdisciplinary project to study the evolution of human behaviour and landscapes in open-air contexts

Hunter-gatherer groups conduct most of their activities in open landscapes as they provide drinking water, food, and raw materials, and offer spaces for social gatherings. The remains left behind at such sites allow for unique archaeological insights into the spatial patterning of prehistoric behaviour. The Stone Age record in southern Africa remains best-known from sheltered sites. A paucity of stratified open-air localities precludes understanding the full spectrum of past hominin activities. Here we introduce an interdisciplinary project in KwaZulu-Natal to study the evolution of human behaviour and landscape dynamics during the Pleistocene in the open-air context of a stratified hillslope with sediments exposed in so-called dongas, a landform created by gully erosion. The project encompasses field and laboratory approaches, combining archaeological, geographical, geological, chronometric, and palaeoenvironmental data. After reviewing relevant open-air research in archaeology and geography, we identify the Jojosi Dongas as a promising research area. Our fieldwork results from 2022-2023 demonstrate the high archaeological, geographical, and geological potential of this landscape. Foot surveys found abundant MSA artefacts spread throughout the dongas, one area with ESA tools, but little to no traces of LSA or later material. Most finds lie on the surface with rarer stratified material occurring in exposed profiles. The surface MSA material is characterised by almost exclusive hornfels use, frequent cortical pieces, many large blanks and cores, and rare retouched tools. These features differ markedly from lithic trends observed in well-known shelter sites, likely the result of differences in site function. Based on the nature of the surface assemblages and the presence of a large outcrop of high-quality hornfels, we hypothesise that the Jojosi Dongas may have been a specialised quarry and workshop area. The stratified MSA occupations are the focus of ongoing excavations and further studies will aim to test our preliminary interpretations provided here.

Hot deformation behavior of a layered heterogeneous microstructure TiAl alloy prepared by selective electron beam melting

TiAl-based alloys are known for their exceptional high-temperature properties but suffer from low hot workability. The emergence of additive manufacturing (AM) technology significantly decreases the workability requirement, while the microstructure heterogeneities in AMed TiAl are troublesome. This paper explores the hot deformation behavior and microstructure evolution of a layered heterogeneous microstructure TiAl alloy prepared by selective electron beam melting. The Arrhenius constitutive model with strain compensation was established, and hot processing maps were constructed. The microstructure showed that at the initial stage of deformation, a large number of dislocation slips occur preferentially in the soft coarse-grained layers. With the increase of strain, dislocation accumulation in heterogeneous interfaces, and dynamic recrystallization (DRX) occurs. Eventually, the coarse-grained region is constantly occupied by fine DRX grains. A high density of deformation twins at high strain rates further promotes the formation of tiny DRX grains. When the hot compression temperature reaches 1200 °C, the initial alternative fine-grained and coarse-grained hetero-structure transforms into duplex (DP) and near-lamellar (NL) microstructure, and microstructure heterogeneity is eliminated. The results enhance understanding of the hot deformation behavior of heterogeneous microstructure TiAl alloys.

Evolution of a novel adrenal cell type that promotes parental care.

Cell types with specialized functions fundamentally regulate animal behaviour, and yet the genetic mechanisms that underlie the emergence of novel cell types and their consequences for behaviour are not well understood1. Here we show that the monogamous oldfield mouse (Peromyscus polionotus) has recently evolved a novel cell type in the adrenal gland that expresses the enzyme AKR1C18, which converts progesterone into 20α-hydroxyprogesterone. We then demonstrate that 20α-hydroxyprogesterone is more abundant in oldfield mice, where it induces monogamous-typical parental behaviours, than in the closely related promiscuous deer mice (Peromyscus maniculatus). Using quantitative trait locus mapping in a cross between these species, we ultimately find interspecific genetic variation that drives expression of the nuclear protein GADD45A and the glycoprotein tenascin N, which contribute to the emergence and function of this cell type in oldfield mice. Our results provide an example by which the recent evolution of a new cell type in a gland outside the brain contributes to the evolution of social behaviour.

Effect of SiC particle content on the mechanical behavior and deformation mechanism of SiCp/Al composite under high-frequency dynamic loading

SiCp/Al composites exhibit excellent structural properties but suffer from inherent brittleness, which has brought rigorous challenges on the materials processing. Ultrasonic-vibration-assisted approach offers a promising solution to improve the processability. However, the underlying mechanisms of mechanical response and defect suppression under ultrasonic action have been rarely studied for SiCp/Al composites, hindering the development of high-efficiency processing. This work aims to investigate the mechanical behavior and microstructural evolution in this class of composites with varying SiC volume fractions up to 45% under high-frequency dynamic load induced by ultrasonic vibration. The results showed that the deformation stress of SiCp/Al composites under ultrasonic compression was significantly reduced to only about 1/10 of that under conventional quasi-static loading. Moreover, as the SiC fractions increasing, this ultrasonic softening effect of SiCp/Al became more pronounced. The plastic limit of SiCp/Al with 45% SiC fraction nearly doubled under ultrasonic compression without any macroscopic cracks. The subsequent nanoindentation tests revealed a transition in work-hardening behavior of SiCp/Al with different SiC fractions under ultrasonic loading. Microstructure characterization was conducted to analyze the grain refinement, recrystallization, and deformation textures within the deformed areas, revealing the deformation mechanism of SiCp/Al under different loading mode. TEM observations confirmed the distribution of dislocations in Al matrix and dense stacking faults in SiC particles after ultrasonic compression, indicating that the plastic behaviors were enhanced in both Al matrix and SiC phase under the high-frequency cyclic stress. This research revealed the distinct deformation mechanisms of SiCp/Al composites under high-frequency dynamic loading, exhibiting extremely low deformation stress accompanied with significantly enhanced plasticity. The results provide theoretical insights for the high-efficiency processing and rapid property regulation of particle-reinforced-composites based on ultrasonic-vibration-assisted technologies.

Extreme elevational migration spurred cryptic speciation in giant hummingbirds

The ecoevolutionary drivers of species niche expansion or contraction are critical for biodiversity but challenging to infer. Niche expansion may be promoted by local adaptation or constrained by physiological performance trade-offs. For birds, evolutionary shifts in migratory behavior permit the broadening of the climatic niche by expansion into varied, seasonal environments. Broader niches can be short-lived if diversifying selection and geography promote speciation and niche subdivision across climatic gradients. To illuminate niche breadth dynamics, we can ask how "outlier" species defy constraints. Of the 363 hummingbird species, the giant hummingbird (Patagona gigas) has the broadest climatic niche by a large margin. To test the roles of migratory behavior, performance trade-offs, and genetic structure in maintaining its exceptional niche breadth, we studied its movements, respiratory traits, and population genomics. Satellite and light-level geolocator tracks revealed an >8,300-km loop migration over the Central Andean Plateau. This migration included a 3-wk, ~4,100-m ascent punctuated by upward bursts and pauses, resembling the acclimatization routines of human mountain climbers, and accompanied by surging blood-hemoglobin concentrations. Extreme migration was accompanied by deep genomic divergence from high-elevation resident populations, with decisive postzygotic barriers to gene flow. The two forms occur side-by-side but differ almost imperceptibly in size, plumage, and respiratory traits. The high-elevation resident taxon is the world's largest hummingbird, a previously undiscovered species that we describe and name here. The giant hummingbirds demonstrate evolutionary limits on niche breadth: when the ancestral niche expanded due to evolution (or loss) of an extreme migratory behavior, speciation followed.

Mechanical behavior and microstructure evolution of slag-fly ash based geopolymer stabilized sand

Geopolymers made from ground granulated blast-furnace slag and fly ash could effectively address the issues of high energy consumption and emissions associated with Portland cement, which is commonly used as a soil curing agent. In this study, the unconfined compressive strength and microstructure of geopolymer-solidified sand were presented and analyzed. The study focused on effect of the type and ratio of activator, the ratio of slag to fly ash, and the ratio of water to (activator and cementitious materials) (W/(A + C)) on the behavior of geopolymer stabilized sand. The results of computed tomography (CT), and scanning electron microscope (SEM) indicate that the variation in initial water content significantly affects the porosity of the specimen, which increased as initial water content increased. The cracks were mainly found at the joints between the individual aggregates and at the edges of the individual aggregates. The pore size distribution of the specimens was concentrated in the range 0.0761–5 μm, while the polymer particle distribution was concentrated in the range 0.0761–10 μm, with a concentration of over 90 %. A significant amount of cementing material was densely distributed around both the aggregates and the pores. CT and SEM testing techniques enable a thorough analysis and mutual verification of the self-compacting process, pore distribution, and size, as well as the distribution of cementing material within the specimens. The optimum ratio for B-S19 has been determined.

Multi-scale study on strength development, hydration behavior and phase evolution of recycled powder multi component cementitious materials

Recycled powder (RP) is considered to be a successful low-carbon alternative to ordinary Portland cement (OPC). However, the microstructure and synergistic mechanism of RP multi-component cementitious materials (RP-MCCM) after physical activation are still unclear. The purpose of this study is to study the strength development, hydration and phase structure evolution of MCCM by constructing RP-MCCM. When the content of RP after grinding activation is 30% (C70RG30), its 28 days compressive strength is increased by 7.6% on the basis of non-activation. Compared with the ternary system of RP and FA mixed with 15% (C70RG15F15), its 28 days compressive strength increased by 8.08%. After RP incorporation, the Al-OH is enhanced, the C-O and CO32- bonds peaks become narrower, and the S is evenly distributed, which is beneficial to the increase of the AFt and the CaCO3 content, but weakens the C-S-H and C-A-H phases. In the C70RG15F15 ternary system, CaCO3, Ca(OH)2, and SiO2 nanostructures are tightly bonded together without obvious faults, which improves the structural compactness and strength. RP grinding activation is mainly manifested in surface improvement, internal C-S-H activation and increase of active sites.

On the aeroelastic bifurcation of a flexible panel subjected to cavity pressure and inviscid oblique shock

The aeroelasticity of a panel in the presence of a shock is a fundamental issue of great significance in the development of hypersonic vehicles. In practical engineering, cavity pressure emerges as a crucial factor that influences the nonlinear dynamical characteristics of the panel. This study focuses on the aeroelastic bifurcation of a flexible panel subjected to both cavity pressure and oblique shock. To this end, a computational method is devised, coupling a high-fidelity reduced-order model for unsteady aerodynamic loads with nonlinear structural equations. The solution is meticulously tracked by continuous calculations. The obtained results indicate that cavity pressure plays a pivotal role in determining the bifurcation and stability characteristics of the system. First, the system exhibits hysteresis behaviour in response to the ascending and descending dynamic pressures. The evolution of hysteresis behaviour originates from the phenomenon of cusp catastrophe. Second, variations in cavity pressure induce three types of bifurcation phenomena, exhibiting characteristics akin to supercritical Hopf bifurcation, subcritical Hopf bifurcation and saddle-node bifurcation of cycles. The system's response at the critical points of these bifurcations manifests as long-period asymptotic flutter or explosive flutter. Lastly, the evolution of the dynamical system among these three types of bifurcations is an important factor contributing to the discrepancies observed in certain research results. This study enhances the understanding of the nonlinear dynamical behaviour of panel aeroelasticity in complex practical environments and provides new explanations for the discrepancies observed in certain research results.

Two-dimensional rheo-optical measurement system to study dynamics and structure of complex fluids

Abstract We have developed a novel rheo-optical measurement system based on two-dimensional polarization analysis, which can evaluate the rheological properties and structure of a complex fluid simultaneously. To assess the utility of the system, we used it to investigate the relationship between yield behavior and structural evolution in a TEMPO-oxidized cellulose nanofiber (T-CNF) suspension, which is a yield-stress fluid that has been actively studied in recent years. To analyze the structural evolution of a T-CNF suspension, stress-ramp tests were conducted. A two-step yield behavior was observed, and distributions of retardation and orientation axis varied dramatically with increasing shear stress. In particular, different distributions were observed in the three regions: after the first yield point, before the second yield point, and after the second yield point. In experiments with a low-concentration T-CNF suspension that exhibits no yield behavior, the retardation increased monotonically with increasing shear stress, and its distribution was uniform. It was demonstrated that the yield behavior and related structures can be analyzed from these results. More detailed structural mechanisms require various rheological tests using the developed system. However, the present insights demonstrate the valuable information provided by the developed rheo-optical measurement system, contributing essential knowledge for applications in various fields.

High-Temperature Cyclic Oxidation Behavior and Microstructure Evolution of W- and Ce-Containing 18Cr-Mo Type Ferritic Stainless Steel.

Due to the recurrent starting and stopping operations of automobiles during service, their engines' hot ends are continually subjected to high-temperature cyclic oxidation. Therefore, it is crucial to develop ferritic stainless steels with better high-temperature oxidation resistance. This study focuses on improving the high-temperature cyclic oxidation performance of 18Cr-Mo (444-type) ferritic stainless steel by alloying with high-melting-point metal W and the rare earth element Ce. For this purpose, a high-temperature cyclic oxidation experiment was designed to simulate the actual service environment and investigate the high-temperature cyclic oxidation behavior and microstructure evolution of 444-type ferritic stainless steel alloyed with W and Ce. The oxide structure and composition formed during this process were analyzed and characterized using scanning electron microscopy/energy dispersive spectroscopy (SEM-EDS) and electron probe X-ray micro-analyzer (EPMA), in order to reveal the mechanism of action of W and Ce in the cyclic oxidation process. The results show that 18Cr-Mo ferritic stainless steel alloyed with W and Ce exhibits an excellent resistance to high-temperature cyclic oxidation. The element W can promote the precipitation of the Laves phase between the matrix and the oxide film, and the small-sized Laves phase can inhibit the interfacial diffusion of oxidation reaction elements and prevent the inward growth of the oxide film. The element Ce can refine oxide particles and reduce the thickness of the oxide film. CeO2 particles within the oxide film can serve as nucleation sites for the formation of oxide particles from reactive elements, and they also contribute to pinning the oxide film, thereby enhancing its adhesion.

Effects of Er addition on the hot deformation behavior and microstructure evolution of AZ31 alloy

AZ31 magnesium alloys typically have poor room temperature ductility after initial deformation. To address this disadvantage, rare earth element Er is introduced to regulate the structure, weaken texture, and promote non-basal slip to achieve plasticization. In this study, the deformation behavior and microstructure evolution during the hot compression process of Er added AZ31 alloy were systematically investigated. Hot compression stress-strain curves showed that the addition of Er reduced the deformation resistance. The constitutive equation and processing map were established. By combining the processing map with the microstructure analysis and texture evolution, an optimal processing area was determined to be 350–400 °C at a strain rate of 0.01 s−1. The influence of Er addition on the twinning behavior, recrystallization mechanism, and slip system of the AZ31 alloy were discussed. During the deformation process, the twinning behavior violates the maximum SF criterion, which is related to severe local stress. Continuous dynamic recrystallization, discontinuous dynamic recrystallization, and twinning induced recrystallization are all detected in the alloy. Through the in-grain misorientation axis and transmission electron microscopy, it can be confirmed that the addition of Er promoted the occurrence of pyramidal <c+a> slip. This is related to the modification of stacking fault energy by Er and the hindrance effect of the second phase containing Er on the basal dislocation.

Tension and Shear Behaviour of Basalt Fiber Bio-Composites with Digital Image Correlation and Acoustic Emission Monitoring.

This research investigates the mechanical behavior and damage evolution in cross-ply basalt fiber composites subjected to different loading modes. A modified Arcan rig for simultaneous acoustic emission (AE) monitoring was designed and manufactured to apply quasi-isotropic shear, combined tensile and shear loading, and pure tensile loading on specimens with a central notch. Digital image correlation (DIC) was applied for high-resolution strain measurements. The measured failure strengths of the bio-composite specimens under different loading angles are presented. The different competing failure mechanisms that contribute to the local reduction in stress concentration are described. Different damage mechanisms trigger elastic waves in the composite, with distinct AE signatures that closely follow the sequence of fracture mechanisms. AE monitoring is employed to capture signals associated with structural damage initiation and progression. The characteristic parameters of AE signals are correlated with crack modes and damage mechanisms. The evolution of AE parameters during the peak load transition is presented, which enables the timely AE detection of the maximum load transition. The combination of DIC and AE monitoring improves understanding of the mechanical response and failure mechanisms in cross-ply basalt fiber composites, offering valuable insights for possible performance monitoring and structural reliability in diverse engineering applications.

Research on the evolutionary control of unsafe behavior of construction personnel based on multi-field coupled-homogeneous analysis model.

Unsafe behavior among construction personnel poses significant risks in petroleum engineering construction projects. This study addresses this issue through the application of a multi-field coupled homogeneous analysis model. By conducting case analyses of petroleum engineering construction accidents and utilizing the WSR methodology, the influencing factors of unsafe behaviors among construction personnel are systematically categorized into organizational system factors, equipment management factors, and construction personnel factors. Subsequently, employing Risk coupling theory, the study delves into the analysis of these influencing factors, discussing their coupling mechanisms and classifications, and utilizing the N-K model to elucidate the coupling effect among them. Furthermore, a novel approach integrating coupling analysis and multi-agent modeling is employed to establish an evolutionary model of construction personnel's unsafe behavior. The findings reveal that a two-factor control method, concurrently reinforcing equipment and construction personnel management, significantly mitigates unsafe behavior. This study provides valuable insights into the evolution of unsafe behavior among construction personnel and offers a robust theoretical framework for targeted interventions. Significantly, it bears practical implications for guiding safety management practices within petroleum engineering construction enterprises. By effectively controlling unsafe behaviors and implementing targeted safety interventions, it contributes to fostering sustainable development within the petroleum engineering construction industry.

Effects of H2/CO ratio and CO2 dilution on the explosion behavior and flame evolution of syngas/air mixtures

Syngas, which is used as both a fuel and a raw material, presents numerous safety risks during its utilization. In this paper, the effects of H2/CO ratio (R = 0%, 30%, 50%, 70% and 100%) and CO2 dilution ([CO2] = 0%–20%) on the explosion behaviors of syngas/air mixtures with different equivalence ratios (φ = 0.8, 1.0, 1.6, and 2.5) are examined via experiments and numerical calculations. The results show that the peaks of explosion pressure and OH* spectral intensity are significantly influenced by the H2 percentage in the syngas. The introduction of CO2 notably decreases these parameters in oxygen-rich and severe oxygen-poor states. Additionally, the flame propagation speed noticeably increases with increasing H2 proportion in the syngas. The influence of CO2 on the buoyancy instability of the spherical flame of syngas is considered negligible when compared to that of other combustible gases, such as CH4. Nevertheless, the presence of CO2 decreases the density ratio between the unburned gas and burned gas and increases the flame thickness of syngas/air mixtures. This trend reduces the cellular instability of the spherical flame, particularly in severe oxygen-poor states. Notably, there is a definite correlation between the formation of cellular flames and the rise rate of explosion pressure. The cellular structure on the flame surface becomes more pronounced when the rise rate of explosion pressure increases. Conversely, it develops later or is noticeably inhibited. Furthermore, the main reaction pathways are H2 → ∙OH → H2O2 → H2O and H2 → ∙OH → CO → CO2 during the syngas explosion process. ∙OH is critical in the entire chain reaction because it promotes the conversion of H2 to H2O and acts as a vital connection between the reactants H2 and CO in the explosion process.

The Deformation Behavior and Microstructure Evolution of Cladding Tube during Spinning Composite Process of Mg/Al Tube

The Deformation Behavior and Microstructure Evolution of Cladding Tube during Spinning Composite Process of Mg/Al Tube

From microstructural changes to electrochemical behavior: decoding the corrosion mechanism in Zr-Sn-Nb-Fe-Cr-Mo-O-V alloys

The oxide formed on a Zr-0.4Sn-0.7Nb-0.3Fe-0.1Cr-0.15Mo-0.12O-0.05V, wt% alloy, after exposure for 300 d in 360°C/20 MPa (water with 0 ppb, 300 ppb, and 1000 ppb), was characterized by transmission electron microscope (TEM), precession electron diffraction (PED) and electrochemical impedance spectroscopy (EIS). Results showed that DO accelerated the corrosion of the alloy, and this effect was enhanced with the increase of the DO concentrations. A mechanism of the correlation between electrochemical behavior evolution and microstructural changes of oxide film was proposed, which provided theoretical guidance for the selection of fuel cladding materials for small nuclear reactors.

Buoyancy control and air breathing in royal knifefish (Chitala blanci) and a new hypothesis for the early evolution of vertebrate air-breathing behaviors.

We present the first description of inspiration-first air breaths in royal knifefish, Chitala blanci, a ray-finned fish known to use four-stroke air breaths. Four-stroke breaths are used by nearly all ray-finned fish species that use their gas bladder to breathe air and are the ancestral breath type of ray-finned fishes. Interestingly, one such species, Amia calva, is known to perform two distinct breath types. Amia use four-stroke breaths when they need more oxygen and performs inspiration-first breaths to restore buoyancy. We observed that C. blanci also performs inspiration-first breaths and tested whether the two breath types are performed for the same functions in C. blanci as they are in Amia. We recorded the frequency of each breath type when exposed to aquatic hypoxia and two conditions of oxygen availability. We found that C. blanci performed more four-stroke breaths (81% ± 15% of total breaths) than inspiration-first breaths when exposed to aerial normoxia but performed more inspiration-first breaths (72% ± 40%) than four-stroke breaths when exposed to aerial hyperoxia. These patterns match those described for Amia and indicate that C. blanci performs four-stroke breaths in response to oxygen depletion and performs inspiration-first breaths to maintain buoyancy. Few studies have examined the role of air-breathing in buoyancy regulation. Decreasing buoyancy, rather than oxygen availability, to stimulate air breaths may reveal that inspiration-first breaths are more common among fishes than we are aware. We consider this possibility and present a new hypothesis for the origin and early evolution of air breathing in vertebrates.

Specialized ovipositor sensilla of Cretaceous wasps (Insecta: Hymenoptera) possibly reveal a unique way of host detection.

Insects have evolved complex sensory systems that are important for feeding, defence and reproduction. Parasitoid wasps often spend much time and effort in searching for concealed hosts with the help of specialized sensilla. However, the early evolution of such behaviour and sensilla is poorly known. We describe two fossil female wasps, †Tichostephanus kachinensis sp. nov. and †Tichostephanus longus sp. nov., from mid-Cretaceous Kachin amber. Phylogenetic analyses based on morphological data retrieved †Tichostephanus as deeply nested within Evanioidea and closely related to extant Gasteruptiidae and Evaniidae. Both of these Cretaceous wasps possess features, e.g. coronal tubercles and flexible ovipositor sheaths, that indicate that they might have laid eggs in wood where their larvae possibly parasitized insect larvae. They have a peculiar and unique 'bottle brush' of sensilla close to the apex of their ovipositor sheaths, which has not been observed in any extant parasitoid wasps. These sensilla comprise many regularly arranged plate-shaped setae, attached in relatively large sockets and with rows of longitudinal ridges. Such specialized sensilla perhaps served to enhance the ability to detect hosts inside wood.