Ripples on tidal flats significantly influence bedform roughness and near-bed turbulence, yet their dynamics in sandy and muddy environments remain incompletely understood. While laboratory studies have elucidated the effects of mud content and extracellular polymeric substances (EPSs) on ripple formation and stability, the interactions between ripple characteristics, EPS, and mud content in natural settings are more complex and not thoroughly explored. To address this gap, we conducted field studies on sand ripples at the central Jiangsu coast, China, using high-resolution drone photogrammetry to accurately measure ripple dimensions. We performed bedload sampling at both ripple crests and troughs to analyze median grain size, mud content, and EPS concentrations. Our results show that the investigated ripple wavelengths range from 34 to 46 mm, and heights vary between 2.7 and 5.3 mm. Ripples developed near tidal creeks show pronounced asymmetry. Significantly, EPS concentrations are markedly higher at the ripple crests than at the troughs, and it follows a power-law relationship with the median grain size. These results highlight the complex intricate relationship between the ripple morphology, environmental forcing conditions, and EPS content. Our findings enrich the understanding of ripple development and the factors influencing current-dominated ripples in natural environments. This research also adds to the better prediction of the bedform roughness and quantification of the near bed sediment transport.
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