Asymmetric Motion Research Articles

Application of Smart Insoles in Assessing Dynamic Stability in Patients with Chronic Ankle Instability: A Comparative Study

Chronic ankle instability (CAI), due to its chronic nature and biomechanical complexity, is well-suited for continuous monitoring and tele-rehabilitation using wearable sensor technology. This study assessed whether a smart insole system, equipped with 4 force-sensing resistor sensors and an inertial measurement unit, combined with functional tests and biomechanical indices, could distinguish CAI patients from healthy controls. A total of 21 CAI patients (23.8 ± 5.1 years) and 16 controls (22.62 ± 2.60 years) completed a battery of functional performance tests while wearing the smart insole system. The results showed an increased medial-lateral pressure ratio in the CAI during heel raise (p = 0.031, effect size = 0.82) and hop tests, suggesting an everted foot position. Significant deviations in center-of-pressure trajectory during double-leg heel raises (p = 0.005, effect size = 1.10) suggested asymmetric motion coordination, while compensatory fluctuations of the lifted limb during single-leg balance tests (p = 0.011, effect size = 1.03) were greater in CAI patients. These findings facilitated the development of features to characterize CAI-specific movement patterns. Together, this system shows promise as a quantitative assessment tool for CAI, supporting improved treatment outcomes through tele-rehabilitation.

Hydrodynamic Characteristics Study of Bionic Dolphin Tail Fin Based on Bidirectional Fluid-Structure Interaction Simulation.

Using bidirectional fluid-structure interaction technology, the dorsal-ventral motion of the dolphin tail fin was simulated, and the feasibility of the numerical simulation method was validated through underwater motion experiments. This study investigated the effects of structural parameters and motion modes of bionic dolphin tail fins on their propulsion performance. The results show that flexible tail fins can enhance propulsion performance. Compared to equal-thickness flexible tail fins, variable-thickness flexible tail fins that conform to the structural characteristics of real dolphin tail fins exhibit better propulsion performance. Asymmetric motion modes have a certain thrust-enhancing effect, but altering the frequency ratio F and amplitude ratio H of heaving motion leads to an increase in pitching moment, reducing swimming stability. Additionally, the greater the difference in frequency and amplitude between the up-and-down motions, the larger the pitching moment. The study results provide references for the optimized design and motion control of bionic tail fins.

Enhancing Performance and Preventing Injuries in Cricketers: The Role of Functional Movement Screening

Functional Movement Screening (FMS) has emerged as a vital tool in sports science for identifying movement inefficiencies, preventing injuries, and enhancing athletic performance. In cricket, a sport characterized by its unique combination of asymmetrical movements, repetitive actions, and high physical demands, FMS offers significant potential to optimize player outcomes. This paper investigates the application of FMS in cricket, focusing on its role in detecting biomechanical dysfunctions, designing corrective interventions, and monitoring performance improvements over time. By examining a cohort of state-level cricketers, the study highlights the effectiveness of FMS in reducing injury incidence, improving movement efficiency, and enhancing key performance metrics such as agility and accuracy. The findings underscore the importance of integrating FMS into cricket training programs and emphasize the need for a multidisciplinary approach involving coaches, physiotherapists, and sports scientists. Despite challenges in implementation, such as resource limitations and the need for specialized training, FMS proves to be a transformative tool in advancing the health and performance of cricketers. Future research directions include the establishment of sport-specific benchmarks, the integration of wearable technology for real-time assessments, and longitudinal studies to explore sustained benefits.

From Motion to Prevention: Evaluating Ergonomic Risks of Asymmetrical Movements and Worker Well-Being in anAssembly Line Work

From Motion to Prevention: Evaluating Ergonomic Risks of Asymmetrical Movements and Worker Well-Being in anAssembly Line Work

Dynamics effect on combustion of a diesel pilot ignition hydrogen vibration combustor with nonlinear asymmetric motion

Dynamics effect on combustion of a diesel pilot ignition hydrogen vibration combustor with nonlinear asymmetric motion

Pilot Study on the Relationship Between Different Lower Limb Raising Velocities and Trunk Muscle Contraction in Active Straight Leg Raise.

The active straight leg raise requires intricate coordination between the hip, knee, pelvis, and spine. Despite its complexity, limited research has explored the relationship between lower limb raising velocity and trunk muscle motor control during an active straight leg raise in healthy individuals. This study aimed to explore the potential effects of increased lower limb raising velocity on core muscle contractions during active straight leg raises. Six healthy adult men (mean age: 24.5 ± 2.5 years) participated in this study. Electromyography signals were recorded using surface electrodes placed on the rectus abdominis, external oblique, and internal oblique/transverse abdominis muscles. The participants performed active straight leg raises at three different velocities: 3 s, 2 s, and as fast as possible (max). The electromyography data were analyzed from 250 ms before to 1000 ms after movement initiation, with muscle activity expressed as a percentage of the maximal voluntary isometric contraction. Statistical analyses were conducted using non-parametric tests, including the Friedman test for overall differences, followed by pairwise Wilcoxon signed-rank tests with Bonferroni correction for multiple comparisons (p < 0.05). During the 250 ms before movement initiation, the internal oblique/transverse abdominis, external oblique, and rectus abdominis muscles showed greater activity in the max condition compared to the 3 s and 2 s conditions (Friedman test, p < 0.05), but no significant differences were found in pairwise comparisons (Wilcoxon test, p > 0.05). Similarly, during the 500 ms after movement initiation, internal oblique/transverse abdominis activity was higher in the max condition, with no significant pairwise differences observed. Faster lower limb raising velocities during active straight leg raise may enhance core stability by activating anticipatory and sustained internal oblique/transverse abdominis, external oblique, and rectus abdominis activity on the raised limb side. Training to promote this activation could improve dynamic stability in rapid or asymmetric movements.

Transient Deformation in the Tatun Volcano Group, Taiwan: A Spatiotemporal GPS Analysis

AbstractThis study analyzed time series data from six GPS stations within the Tatun Volcano Group (TVG), a long‐dormant volcanic system in northern Taiwan, using multichannel singular spectrum analysis to search for potential spatiotemporally correlated transient deformations. A notable cycle of transient deformation was identified from 2015 to 2020, characterized by ground subsidence and uplift of up to 10 mm, accompanied by asymmetric horizontal motions directed inward and outward toward Dayoukeng, the largest fumarole and hydrothermal area in TVG. Evidence from earthquake focal mechanisms and gas composition, along with preliminary source modeling, suggest that these transient phases were likely caused by the pressure change of shallow hydrothermal systems beneath Dayoukeng. Further analysis of time series data from three long‐operating GPS stations revealed similar patterns of transient motion in the area from 2006 to 2015, indicating that TVG has experienced cyclical deformation, akin to many other volcanic systems worldwide.

Long Terms Results of Temporal Facelift: 6 Years of Experience in 250 Cases.

Temporal facelift (TFL) is an innovative technique for lifting the upper and mid-face. It is characterized by a unique dissection plane above the subgaleal fascia, which seamlessly transitions into the sub-superficial muscular aponeurotic system (SMAS) layer in the mid-face. This approach enables comprehensive mid-face elevation, robust canthopexy, and a significant brow lift in various vectors. The authors present their experience with 250 TFL procedures over a period of 6 years. This retrospective study analyzed 250 of 441 patients who underwent TFL surgery. The surgical procedures, conducted under general anesthesia by a senior surgeon following the TFL method, involved a vertical-vector deep-plane mid-face lift, canthopexy, and brow-lift triad. Close monitoring of complications and detailed photographic documentation of the outcomes were performed. Postoperative care included taping the operation area to reduce swelling, with subsequent follow-up examinations and interventions such as lymphatic massage and botulinum toxin injections for asymmetric brow movements or steroid injections for excessive mid-face swelling. Among 250 patients (248 female, 2 male; mean age, 37y), unilateral neuropraxia of the frontal branch of the facial nerve occurred in 5.6% and resolved spontaneously within 2.8 months. Persistent dimples (2.8%) resolved by the fifth postoperative month. Six patients (2.4%) underwent revision surgery with no observed complications like hematoma, necrosis, infection, or seroma. The TFL technique represents a significant advancement in upper and mid-face lifting procedures and offers several advantages over the traditional methods.

Analytic insight into the physics of the standing accretion shock instability

Context. During the core collapse of a massive star, and immediately before its supernova explosion, there is amplification of asymmetric motions by the standing accretion shock instability (SASI). This imprints a frequency signature on the neutrino flux and the gravitational waves that carries direct information about the explosion process. Aims. The physical interpretation of this multi-messenger signature requires a detailed understanding of the instability mechanism. Methods. We carried out a perturbative analysis to characterise the properties of SASI and assess the effect of the region of neutronization above the surface of the proto-neutron star. We compared the eigenfrequencies of the most unstable modes to those obtained in an adiabatic approximation where neutrino interactions are neglected above the neutrinosphere. We solved the differential system analytically using a Wronskian method and approximated it asymptotically for a large shock radius. Results. The oscillation period of SASI is well fitted with a simple analytic function of the shock radius, the radius of maximum deceleration, and the mass of the proto-neutron star. The oscillation period is weakly dependent on the parameterised cooling function, but this latter does affects the SASI growth rate. We describe the general properties of SASI eigenmodes using an adiabatic model. In this approximation, the eigenvalue problem is formulated as a self-forced oscillator. The forcing agent is the radial advection of baroclinic vorticity perturbations and entropy perturbations produced by the shock oscillation. We reduced the differential system defining the eigenfrequencies to a single integral equation. Its analytical approximation sheds light on the radially extended character of the region of advective-acoustic coupling. The simplicity of this adiabatic formalism opens new perspectives for the investigation of the effect of stellar rotation and non-adiabatic processes on SASI.

Numerical study on hydroelastic responses and bending-torsion coupled loads of a ship in oblique regular waves

In this paper, the CFD-FEM two-way coupled method is extended to simulate the asymmetric motions and wave load responses of a ship in oblique regular waves considering hydroelasticity effects. The numerical results of ship motions and loads including time series and response amplitude operators are validated by comparing with the tank experimental results. Then the ship motions, vertical bending moment, horizontal bending moment and torsion moment in oblique waves are analyzed in both time domain and frequency spectra; and the influence of wave height, wave length and ship speed on the results are systematically analyzed. Moreover, the correlation between horizontal bending moment and torsion moment as well as their component loads is analyzed by cross-spectral analysis method. The effect of springing responses corresponding to the bending-torsion coupled mode on the sectional loads is also discussed.

Integration of horizontally acquired light-harvesting genes into an ancestral regulatory network in the cyanobacterium Acaryochloris marina MBIC11017.

The acquisition of new capabilities by horizontal gene transfer (HGT) shapes the distribution of traits during microbial diversification. In the Chlorophyll (Chl) d-producing cyanobacterium Acaryochloris marina, the genes involved in the production and disassembly of the light-harvesting phycobiliprotein phycocyanin (PC) were lost in the A. marina common ancestor but then subsequently regained via HGT in A. marina strain MBIC11017. However, it remains unknown how the HGT-acquired PC genes in MBIC11017 have been reintegrated into its existing regulatory network after tens of millions of years since their loss. Here, we investigated potential mechanisms of regulatory assimilation of PC genes by comparing the transcriptomes of A. marina strain MBIC11017 and a PC-lacking close relative under both low irradiance far-red light and high irradiance white light. We found that PC assembly and degradation processes have been re-assimilated into a conserved ancestral response to high light. Further, we identified putative regulatory elements that were likely co-transferred with PC genes and could be recognized by A. marina's pre-existing light response machinery. This study offers insights into how HGT-acquired genes can be reintegrated into an existing transcriptional regulatory network that has evolved in their absence.IMPORTANCEHorizontal gene transfer, the asymmetric movement of genetic information between donor and recipient organisms, is an important mechanism for acquiring new traits. In order for newly acquired gene content to be retained, it must be integrated into the genetic repertoire and regulatory networks of the recipient cell. In a strain of the Chlorophyll d-producing cyanobacterium Acaryochloris marina, the recent reacquisition of the genes required to produce the light-harvesting pigment phycocyanin offers a rare opportunity to understand the mechanisms underlying the regulatory assimilation of an acquired complex trait in bacteria. The significance in our research is in characterizing how an ancestrally lost, complex trait can be reintegrated into a conserved regulatory network, even after millions of years.

Tropical Cyclone Wind Shear-Relative Asymmetry in Reanalyses

Abstract While tropical cyclones (TCs) are axisymmetric vortices to the first order, they often exhibit noteworthy structural asymmetries. These often result from environmental vertical wind shear, which tilts the vortex and induces a wavenumber 1 pattern in the circulation and precipitation fields. Reanalyses and climate models have improved in representing the TC structure and climatology, but their relatively coarse resolution and dependence on parameterized physics cast doubt on their ability to capture the asymmetric TC structure. We perform the most comprehensive process-oriented assessment of TC asymmetry to date in reanalyses. Specifically, we analyze the composite shear-relative TC structure in ERA5 and Climate Forecast System Reanalysis (CFSR), which vary in their resolutions, physical parameterization suites, and data assimilation techniques. These structures are compared with aircraft reconnaissance radar observations. In agreement with the observations, the strongest tangential winds are usually found left-of-shear, while inner core rainfall, ascent, vortex tilt, and low-level inflow are favored directly downshear or in the downshear-left quadrant. Outer rainband convection generally peaks in the downshear-right quadrant. Thermodynamic asymmetries are also apparent, with anomalous low-level moisture right-of-shear, midlevel warmth in the upshear-right quadrant (uptilt), and cloud properties suggestive of a realistic precipitation life cycle from growth to fallout. We also decompose rainfall contributions from the convective parameterization and large-scale cloud schemes and highlight the roles of vorticity advection, buoyancy advection, and diabatic processes in driving asymmetric vertical motions in the inner core and outer rainband regions. Our results suggest that process-level studies of TC asymmetry and TC–wind shear interaction under future warming are viable using climate models. Significance Statement Asymmetries are common in tropical cyclones (TCs), influencing their intensity, track, and hazards. Vertical wind shear often plays a leading-order role in causing these asymmetries. It is uncertain how well asymmetric structures and processes are captured in reanalyses and global climate models (GCMs) with grid spacings of 0.25° and coarser. In this study, we first evaluate TC asymmetry in reanalyses, which have the benefit of being forced by observations. This helps to assess whether the resolutions associated with GCMs sufficiently capture asymmetric structures and processes and motivates upcoming work with free-running GCMs to study how TC asymmetry may change in a warming climate.

Multi-species ecological network based on asymmetric movement: Application in an urban rural fringe

Ecological network (EN) is a popular approach for biodiversity conservation, which aims to facilitate animal movement between habitats. However, asymmetric movement caused by the subjectivity of animals and environment heterogeneity is seldom considered in EN design. To design EN based on asymmetric movement, an individual-based model, PDArunner, is developed, which can identify corridors based on explicit movement paths. Since the urban–rural fringe is susceptible to landscape change, a multi-species EN based on asymmetric movement is designed for Jinnan district, Tianjin, China by simulating movement of Mustela sibirica, Spilopelia chinensis and Gallinula chloropus using PDArunner. Asymmetric movement is partitioned based on the size of departure and arrival habitats. Asymmetric corridors are thus identified based on movement in particular direction. In Jinnan, successful transfer rate of focal species from small to large habitat increases with difference in habitat size. The spatial extent of asymmetric corridors is more concentrated for G. chloropus. There are more corridors from large to small habitats than in the opposite direction, especially for S. chinensis and G. chloropus. There are more one-way corridors for S. chinensis and G. chloropus than M. sibirica. Perceptible high vegetation coverage places with large enough contrast to environment are suggested to be consecutive within 50 m for M. sibirica in Jinnan, which also benefits S. chinensi. G. chloropus can benefit from well-conserved water quality and quantity. A cross-administration collaboration is also highlighted for large scale conservation.

Competing Bifurcations Determine Symmetry Breaking During Droplet Snaps on Smooth Patterned Surfaces.

The shape and stability of a droplet in contact with a solid surface is affected by the chemical composition and topography of the solid, and crucially, by the droplet's size. During a variation in size, most often observed during evaporation, droplets on smooth patterned surfaces can undergo sudden shape and position changes. Such changes, called snaps, are prompted by the surface pattern and arise from fold and pitchfork bifurcations which respectively cause symmetric and asymmetric motions. Yet, which type of snap is likely to be observed is an open fundamental question that has relevance in the rational design of surfaces for managing droplets. Here we show that the likelihood of observing symmetric or asymmetric snaps depends on the distance between fold and pitchfork bifurcation points and on how this distance varies for droplets that grow or shrink in size on surfaces patterned with a smooth topography. Our results can help develop strategies to control droplets by exploiting smooth surface patterns but also have broader relevance in situations where different types of bifurcations compete in determining the stability of a system, for instance in snap-through instabilities observed in elastic media.

Movement Strategy Moderates the Effect of Spatially Congruent Cues on the Stability of Rhythmic Bimanual Finger Movements.

Spatially congruent cues increase the speed of bimanual reach decisions compared with abstract symbolic cues, particularly for asymmetric reaches. Asymmetric rhythmic bimanual movements are less stable than symmetric rhythmic movements, but it is not well understood if spatially congruent cues similarly increase the stability of asymmetric rhythmic bimanual movements. To address this question, in Experiment 1, participants performed symmetric and asymmetric bimanual rhythmic finger tapping movements at different movement frequencies in time with flickering spatially congruent and abstract symbolic stimuli. As expected, symmetric movements were more stable. Spatially congruent cues similarly increased the stability of symmetric and asymmetric movements compared with abstract symbolic cues. The benefits of spatial congruence and movement symmetry were restricted to high movement frequencies (>2 Hz). To better understand if the emergence of these effects at high movement frequencies was driven by a change in movement strategy, in Experiment 2, video of the hands was concurrently recorded during task performance. Markerless motion tracking software revealed that participants switched from discontinuous to continuous movement strategies with increasing movement frequency. Because discontinuous and continuous movements are thought to be controlled by distinct neurocognitive systems, this might explain why the beneficial effects of spatial congruence and response symmetry emerged only at high movement frequencies. Overall, results from the current study indicate that the perceptual quality of the stimulus use to cue movement frequency can have powerful effects on the stability of rhythmic bimanual movements, but that these effects may depend on whether discontinuous or continuous movement strategies are selected.

Load-based divergence in the dynamic allostery of two TCRs recognizing the same pMHC.

Increasing evidence suggests that mechanical load on the αβ T cell receptor (TCR) is crucial for recognizing the antigenic peptide-loaded major histocompatibility complex (pMHC) molecule. Our recent all-atom molecular dynamics (MD) simulations revealed that the inter-domain motion of the TCR is responsible for the load-induced catch bond behavior of the TCR-pMHC complex and peptide discrimination. To further examine the generality of the mechanism, we perform all-atom MD simulations of the B7 TCR under different conditions for comparison with our previous simulations of the A6 TCR. The two TCRs recognize the same pMHC and have similar interfaces with pMHC in crystal structures. We find that the B7 TCR-pMHC interface stabilizes under ~15-pN load using a conserved dynamic allostery mechanism that involves the asymmetric motion of the TCR chassis. However, despite forming comparable contacts with pMHC as A6 in the crystal structure, B7 has fewer high-occupancy contacts with pMHC during the simulation. These results suggest that the dynamic allostery common to the TCRαβ chassis can amplify slight differences in interfacial contacts into distinctive mechanical responses and potentially nuanced biological outcomes.

Propulsive efficiency of spatiotemporally asymmetric oscillating appendages at intermediate Reynolds numbers

Many organisms use flexible appendages for locomotion, feeding, and other functional behaviors. The efficacy of these behaviors is determined in large part by the fluid dynamics of the appendage interacting with its environment. For oscillating appendages at low Reynolds numbers, viscosity dominates over inertia, and appendage motion must be spatially asymmetric to generate net flow. At high Reynolds numbers, viscous forces are negligible and appendage motion is often also temporally asymmetric, with a fast power stroke and a slow recovery stroke; such temporal asymmetry does not affect the produced flow at low Reynolds numbers. At intermediate Reynolds numbers, both viscous and inertial forces play non-trivial roles---correspondingly, both spatial and temporal asymmetry can strongly affect overall propulsion. Here we perform experiments on three robotic paddles with different material flexibilities and geometries, allowing us to explore the effects of motion asymmetry (both spatial and temporal) on force production. We show how a flexible paddle's time-varying shape throughout the beat cycle can reorient the direction of the produced force, generating both thrust and lift. We also evaluate the propulsive performance of the paddle by introducing a new quantity, which we term "integrated efficiency". This new definition of propulsive efficiency can be used to directly evaluate an appendage's performance independently from full-body swimming dynamics. Use of the integrated efficiency allows for accurate performance assessment, generalization, and comparison of oscillating appendages in both robotic devices and behaving organisms. Finally, we show that a curved flexible paddle generates thrust more efficiently than a straight paddle, and produces spatially asymmetric motion---thereby improving performance---without the need for complex actuation and controls, opening new avenues for bioinspired technology development.

Enhancing underwater unmanned vehicle efficiency through asymmetric dynamics in manta-like swimming

This study explores the hydrodynamic performance of a National Advisory Committee for Aeronautics 0012 foil using computational fluid dynamics with the unsteady Reynolds-averaged Navier–Stokes (URANS) method and shear stress transport k-ω model to assess the impact of asymmetric motion parameters in manta-like swimming. The angles of attack during the mid-upstroke (αmu), mid-downstroke (αmd), and stroke duration (S) are varied to understand their effect. At low Strouhal numbers (StA = 0.2–0.35), a smaller αmd compensates for thrust loss at the start of the upstroke due to a greater αmu. At high Strouhal numbers (StA = 0.5), a greater αmd reduces negative thrust and compensates for the smaller thrust generated by a small αmu during the upstroke. Shorter stroke durations increase asymmetry, leading to more significant positive thrust peaks during the downstroke. If both the angle of attack and S are large, the slower downward speed extends negative thrust, reducing thrust peaks and lowering average thrust. A smaller stroke duration combined with a large angle of attack enhances efficiency due to a greater thrust-to-power ratio, highlighting the interplay between these parameters. A smaller S and greater αmd and StA maximize thrust and efficiency, suggesting aquatic organisms increase thrust while ensuring propulsion efficiency by using a large angle of attack and an asymmetric stroke duration. This study demonstrates how asymmetric parameters interact, providing insights into designing biomimetic underwater vehicles. The findings suggest that asymmetric dynamics enhance propulsion efficiency.

Diversity and transition of periodic motion of a periodically excited soft-impacting machinery

Dynamics of a periodically excited vibro-impact system with soft impacts is investigated. Essential features of period-one multi-impact motion group and correlated transition characteristics in low-frequency range are discussed in detail by the way of two-parameter bifurcation space providing qualitative domains for different periodic motions. The main focus is given to the effect of sensitive parameters including constraint stiffness k 0, clearance threshold b, and damping parameter ζ on the system response. The low-frequency characteristics in the finite-dimensional parameter space are particularly explored. It is found that the increase of k 0 induces multi-type bifurcation of period-one double-impact symmetrical motion, which induces a rich variety of periodic motions, and period-one multi-impact motion group orbit primarily exist in the small-clearance b and low-frequency ω zone. Based on the evolution irreversibility of adjacent period-one multi-impact orbit, the mechanism of singularies appearing in pairs and two different transition zones (hysteresis and liguliform zones) is studied, the result of which provides a theoretical reference value for the common low-frequency vibration instability phenomenon in the field of mechanical engineering. For small-damping coefficient ζ, period-one multi-impact motion has a large quantity, and the main bridge for the transition of adjacent period-one multi-impact motion is liguliform zone, which embraces period-one multi-impact asymmetrical motion and period-n multi-impact subharmonic motion and a certain chaotic zone. For large-damping coefficient ζ, the amount of period-one multi-impact motion group is reduced, and the main bridge for the transition of adjacent period-one multi-impact motion is hysteresis zone, where adjacent period-one multi-impact orbits can coexist according to initial conditions. As designing and renovating impact mechanical equipment, the reasonable matching law of dynamic parameters can be determined through two-parameter bifurcation space, which is conducive to making the system work in stable periodic motion and obtaining larger instantaneous impact velocity.

Tree Frogs Alter Their Behavioral Strategies While Landing On Vertical Perches.

As an arboreal animal, tree frogs face diverse challenges when landing on perches, including variations in substrate shape, diameter, flexibility, and angular distribution, with potentially significant consequences for failed landings. Research on tree frog landing behavior on perches, especially concerning landing on vertical substrates, remains limited. This study investigated the landing strategies (forelimb, abdomen, and hindlimb) of tree frogs on vertical perches, considering perch diameter. Although all three strategies were observed across perches of different diameters, their frequencies differed. Forelimb landing was most common across all perch diameters, with its frequency increasing with perch diameter, while abdomen and hindlimb landing strategies were more prevalent on smaller diameter perches. During the process from take-off to landing, the body axis underwent some deviation owing to the asymmetric movement of the left and right limbs; however, these deviations did not significantly differ among landing strategies. Additionally, different landing strategies led to variations in the landing forces, with abdominal landings generating significantly higher impact forces than the other two strategies. These findings provide insights into the biomechanics and biological adaptations of tree frogs when landing on challenging substrates, such as leaves or branches.