Planktonic Foraminifera Assemblages Research Articles

ForCenS-LGM: a dataset of planktonic foraminifera species assemblage composition for the Last Glacial Maximum

Species assemblage composition of marine microfossils offers the possibility to investigate ecological and climatological change on time scales inaccessible using conventional observations. Planktonic foraminifera - calcareous zooplankton - have an excellent fossil record and are used extensively in palaeoecology and palaeoceanography. During the Last Glacial Maximum (LGM; 19,000 – 23,000 years ago), the climate was in a radically different state. This period is therefore a key target to investigate climate and biodiversity under different conditions than today. Studying LGM climate and ecosystems indeed has a long history, yet the most recent global synthesis of planktonic foraminifera assemblage composition is now nearly two decades old. Here we present the ForCenS-LGM dataset with 2,365 species assemblage samples collected using standardised methods and with harmonised taxonomy. The data originate from marine sediments from 664 sites and present a more than 50% increase in coverage compared to previous work. The taxonomy is compatible with the most recent global core top dataset, enabling direct investigation of temporal changes in foraminifera biogeography and facilitating seawater temperature reconstructions.

Coherent response of zoo‐ and phytoplankton assemblages to global warming since the Last Glacial Maximum

AbstractAimWe are using the fossil record of different marine plankton groups to determine how their biodiversity has changed during past climate warming comparable to projected future warming.LocationNorth Atlantic Ocean and adjacent seas. Time series cover a latitudinal range from 75° N to 6° S.Time periodPast 24,000 years, from the Last Glacial Maximum (LGM) to the current warm period covering the last deglaciation.Major taxa studiedPlanktonic foraminifera, dinoflagellates and coccolithophores.MethodsWe analyse time series of fossil plankton communities using principal component analysis and generalized additive models to estimate the overall trend of temporal compositional change in each plankton group and to identify periods of significant change. We further analyse local biodiversity change by analysing species richness, species gains and losses, and the effective number of species in each sample, and compare alpha diversity to the LGM mean.ResultsAll plankton groups show remarkably similar trends in the rates and spatio‐temporal dynamics of local biodiversity change and a pronounced non‐linearity with climate change in the current warm period. Assemblages of planktonic foraminifera and dinoflagellates started to change significantly with the onset of global warming around 15,500 to 17,000 years ago and continued to change at the same rate during the current warm period until at least 5000 years ago, while coccolithophore assemblages changed at a constant rate throughout the past 24,000 years, seemingly irrespective of the prevailing temperature change.Main conclusionsClimate change during the transition from the LGM to the current warm period led to a long‐lasting reshuffling of zoo‐ and phytoplankton assemblages, likely associated with the emergence of new ecological interactions and possibly a shift in the dominant drivers of plankton assemblage change from more abiotic‐dominated causes during the last deglaciation to more biotic‐dominated causes with the onset of the Holocene.

REFERENCE SECTION OF THE UPPER MAASTRICHT OF MOUNT KLEMENTYEVA (SOUTH-EASTERN CRIMEA)

The results of a comprehensive study of the reference section of the Upper Maastrichtian of Mount Klementyev (South-Eastern Crimea), composed of a terrigeneous-carbonate strata. It was found that sedimentation occurred in relatively deep-water conditions. Biostratigraphic analysis of benthic and planktonic foraminifera assemblages made it possible to identify foraminiferal zones in the section and confirm the deep-water type of sedimentation in the basin at this interval.

Middle Eocene Climatic Optimum (MECO): The first record based on planktonic foraminifera and stable isotopes from SW Anatolia/Turkiye

The Kayikoy Formation (Isparta, SW Turkiye) consisting of sandstone, claystone, and marl units contains abundant, low-moderate preserved, and diverse planktonic foraminifera assemblages. Eight planktonic foraminiferal biozones (E9-E16) were determined encompassing the Lutetian-Priabonian stages (middle-upper Eocene) by using the biostratigraphic distributions of 17 genera and 62 species of planktonic foraminifera. Stable delta 18O and delta 13C isotope values were measured for 53 individuals belonging to Morozovelloides crassatus, Acarinina bullbrooki, A. topilensis, A. echinata, Globigerinatheka curryi, G. index, G. subconglobata, G. mexicana, G. barri, G. euganea, Orbulinoides beckmanni, Turborotalia frontosa, T. pomeroli, Hantkeninamexicana, H. liebusi, H. dumblei, H. alabamensis, and Dentoglobigerina eotripartita. By integrating planktonic foraminiferal biostratigraphy, stable delta 18O and delta 13C isotopes, and paleotemperature analyses, the first detailed study of the Middle Eocene Climatic Optimum (MECO) record was performed in SW Anatolia/Turkiye. The overall signature of the MECO is similar to global observations but also involves some regional differences. The MECO paleotemperatures, determined to vary between approximately 2 degrees C and 5 degrees C, were similar to the global records, but the slight differences in values in the three sections are most likely due to the paleogeographic position of the studied section in Turkiye.

Foraminifera assemblages from Fantangisña serpentinite mud seamount in the NW Pacific Ocean during the Pleistocene (IODP Expedition 366).

The Mariana forearc system, in the northwestern Pacific, is known as the only convergent margin setting with currently active serpentine mud volcanism. The Fantangisña serpentinite mud volcano lies 62 km west of the Mariana trench, within the influence of the North Equatorial Current (NEC). Cores recovered by International Ocean Discovery Program (IODP) Expedition 366 contain pelagic sediments overlying layered serpentinite mud deposits. At the bottom of the sequence, nannofossil-rich forearc deposits were recovered from under the seamount edifice. In this study, we investigated 47 samples from Site U1498A on the southern flank of the seamount for benthic and planktonic foraminifera assemblages. Statistical analyses on planktonic assemblages differentiated two sample groups related to the ratio between thermocline/mixed layer taxa, which indicate fluctuations in the depth of the thermocline (DOT) during the Pleistocene. Variations in the DOT reflect changes in the intensity of the NEC associated with El Niño/La Niña conditions. Mudflows do not influence the ecology of planktonic foraminifera but possibly enhance their preservation against dissolution, which was instead detected in the pelagic deposit as suggested by common Globigerinoides conglobatus. Benthic foraminifers were rare in serpentinite mud deposits as they are severely affected by mudflows. Conversely, they showed high diversity pre-/post-mud-volcanism, and indicate oligotrophic bottom-water conditions.

DISTRIBUCION DE LOS FORAMINIFEROS PLANCTONICOS DEL PLIOCENO EN LA PLATAFORMA CONTINENTAL DEL SW DE ESPANA

Planktonic Foraminifera assemblages from cores of a borehole are studied. Depth interval is from 380 m to 2300 m. The drill site is located in the continental shelf of "Golfo de Cadiz" (SW Spain). The distribution of planktonic foraminiferal species is in agreement with Blow's zonation (1969) extending in age from the Messinian to the Pleistocene.

Planktonic Foraminiferal Biostratigraphy of the Kometan Formation from Kurdistan Region, NE Iraq

The Kometan Formation is widely spread in northeastern Iraq and is mostly composed of deep marine carbonates, with a predominance of planktonic foraminiferal assemblages throughout its sequences. Two outcrop sections, Smaquli and Tabin, were chosen to study the biostratigraphy of the Kometan Formation. The study identified 32 planktonic foraminiferal species belonging to 12 genera. According to stratigraphic distribution and the relative distribution of planktonic foraminifera assemblages, the study demonstrated the possibility of subdividing the sequence of the Kometan Formation into five foraminiferal biozones, including: Marginotruncana schneegansi Partial Range Zone, Dicarinella primitiva Interval Zone, Dicarinella concavata Interval Zone, Dicarinella asymetrica Total Range Zone, Globotruncanita elevata Partial Range Zone. The Planktonic zones were correlated with other zonal schemes inside and outside Iraq. They are considered to be extending from Late Turonian to Early Campanian.

Distribution of recent planktonic foraminifera in surface sediments of the Basque shelf (S Bay of Biscay): Oceanographic implications

In this work we describe for the first time the distribution of planktonic foraminifera assemblages preserved in recent surface sediments that cover the Basque shelf (southern Bay of Biscay), and their relationship with diverse oceanic currents that affect this area. In the 83 studied samples a total of 18 (morpho)species of planktonic foraminifera have been identified, being the most abundant species: Neogloboquadrina pachyderma dex., Globigerinoides ruber, Globigerina bulloides, Globoconella inflata, Orbulina universa and Turborotalita quinqueloba. These species represent the 92.8% of the total hand-picked specimens. The preferential accumulation of planktonic foraminifera shells in fine sediments deposited in the eastern margin of the Basque shelf, and the random distribution of the dominant species in this area, seem to respond to the general surface current that affect the Basque shelf, which flows eastward during most of the year. Moreover, the input of certain currents and deeper water masses in this area has an important impact on assemblage distribution. Thus, the presence of abundant individuals of Globigerinoides ruber indicates the entrance of the warm and salty Iberian Poleward Current (IPC). Likewise, the dominance of subpolar species Globigerina bulloides, Neogloboquadrina pachyderma dex. and Turborotalita quinqueloba, together with the occurrence, as accessories species, of Neogloboquadrina pachyderma sin., Globorotalia scitula, Globorotalia truncatulinoides and Neogloboquadrina dutertrei, suggest the impact of upwelling processes on the Basque shelf. In conclusion, in this work we have evidenced the useful application of planktonic foraminifera assemblages preserved in surface sediments as indicators of oceanic currents and water masses in shallow continental shelves, such as the Basque shelf.

The Loop Current Circulation Over the MIS 9 to MIS 5 Based on Planktonic Foraminifera Assemblages From the Gulf of Mexico

AbstractThe loop current (LC) in the Gulf of Mexico (GoM) is part of the western North Atlantic circulation. Recording its strength and slowdown variations can help us characterize the regional climate over the Late Pleistocene. To reconstruct the sea surface and the LC intensity in the eastern GoM, we study the distribution patterns of planktonic foraminifera in the core EN‐032‐18PC, spanning the end of Marine Isotope Stage (MIS) 9 to early MIS‐4. We reconstructed a sequence of paleoceanographic events based on stable isotopes (δ18O and δ13C) of the surface dweller Globigerinoides ruber and two faunal assemblages. The first assemblage explains most of the glacial and late interglacial periods, suggesting a subtropical environment with a deep thermocline and a reduced LC due to a moderate inflow of warm Caribbean waters. The second assemblage explains the warmest interglacial substages, dominated by tropical species, a shallow thermocline, and an extended LC, driven by summer insolation. Overall, surface ocean conditions led to more ecological successions and instability during the warmest interglacial substages than during glacial periods, as supported by the stable isotope records. Besides the GoM relationship to AMOC, as a regulator of heat transport to higher latitudes, we suggest that fluctuations in the LC rely on the migration of atmospheric circulation patterns and astronomical insolation forcing.

Planktonic foraminifera response to orbital and millennial-scale climate variability at the southern Iberian Margin (IODP Site U1385) during Marine Isotope Stages 20 and 19

The orbital configuration of Marine Isotope Stage (MIS) 19 provides a good analogue to our present interglacial, and it is marked by millennial-scale climate variability. To evaluate how orbital and millennial-scale climate changes interact during this interglacial we have conducted a high-resolution study of planktonic foraminifera assemblages at International Ocean Discovery Program (IODP) Site U1385, SW Iberian Margin, encompassing late MIS 20 and MIS 19. Estimates of sea surface temperature (SST), using transfer-functions, are compared with geochemical, XRF elemental data, biomarker and pollen records from the same site. These results are then compared with SST and ice-rafted debris records from North Atlantic IODP Site U1314 to reconstruct basin-wide palaeoceanographic conditions. During MIS 20, the large size of the ice sheets imparted a strong cooling in the subpolar gyre whereas the Iberian Margin remained under the influence of the subtropical gyre with the development of a high meridional thermal gradient in the North Atlantic. During MIS 20 terminal stadial (794–789 ka) and MIS 19b-a stadial events, the meltwater discharges related to the instability of ice sheets affected both the North Atlantic and the Iberian Margin leading to a weaker thermal gradient. Our data indicate that Termination IX (TIX, ∼795–788 ka) was punctuated by short-term warming/cooling phases in the ocean coupled with warming/cooling events on land. A short-term cooling episode, recognized in the ocean and on land, occurred on Termination IX and preceded the beginning of MIS 19c. During MIS 19c, the planktonic foraminifera SST indicates warm and almost stable conditions in the sea decoupling the cool and dry millennial-scale events on land inferred by pollen assemblages. Starting from 780 ka towards MIS 18, the planktonic foraminifera assemblages and SSTs indicate a cooling trend along the Iberian Margin related to a progressive southward migration of the polar front.

Nutrients and hydrography explain the composition of recent Mediterranean planktonic foraminiferal assemblages

The composition of planktonic foraminiferal assemblages in surface sediments of the Mediterranean Sea shows a pronounced zonal gradient. This gradient is different from what is observed in the open ocean, indicating that the assemblage composition may respond to environmental variables other than the otherwise dominant surface water temperature. Here we make use of a dataset of census counts of 18 taxonomic categories in 124 surface-sediment samples extracted from the ForCenS database to understand the main environmental factors affecting the composition of recent Mediterranean planktonic foraminiferal assemblages. The tested explanatory variables include temperature, salinity, chlorophyll, nitrate and phosphate concentrations in the surface waters, temperature, nitrate and phosphate concentrations at depth and thermal, salinity and density vertical gradients. The composition of the assemblages is aligned along two environmental gradients and redundancy analysis reveals that the tested variables explain a large portion (>70%) of the variance in the compositional data. The first environmental gradient reflects the nutrient content in the deep waters, affecting nutrient availability in the productive zone, while the second gradient is driven by higher (northern area) and lower (southern area) upward nutrient advection due to differences in density stratification, conditioned by the regional hydrography. Although sea surface temperature is the most reconstructed environmental variable from fossil assemblages of planktonic foraminifera, it seems to play a secondary role in the Mediterranean Sea. This, has implications for reconstructions of past oceanographic conditions in this region, opening up the possibility to reconstruct productivity instead, or next to, temperature.

Plankton response to global warming is characterized by non-uniform shifts in assemblage composition since the last ice age

Biodiversity is expected to change in response to future global warming. However, it is difficult to predict how species will track the ongoing climate change. Here we use the fossil record of planktonic foraminifera to assess how biodiversity responded to climate change with a magnitude comparable to future anthropogenic warming. We compiled time series of planktonic foraminifera assemblages, covering the time from the last ice age across the deglaciation to the current warm period. Planktonic foraminifera assemblages shifted immediately when temperature began to rise at the end of the last ice age and continued to change until approximately 5,000 years ago, even though global temperature remained relatively stable during the last 11,000 years. The biotic response was largest in the mid latitudes and dominated by range expansion, which resulted in the emergence of new assemblages without analogues in the glacial ocean. Our results indicate that the plankton response to global warming was spatially heterogeneous and did not track temperature change uniformly over the past 24,000 years. Climate change led to the establishment of new assemblages and possibly new ecological interactions, which suggests that current anthropogenic warming may lead to new, different plankton community composition.

Stable isotope geochemistry and microfossil assemblages of carbonate rocks in the ophiolite mélange zone of the Indo–Myanmar orogenic Belt, NE India: Implications on age and depositional environment

A comprehensive data of whole‐rock and stable isotopic geochemistry along with microfossil assemblages of the carbonate rocks of the ophiolite mélange zone of the Indo–Myanmar orogenic Belt (IMOB), north‐east India are discussed, to determine the influence of terrigenous contamination during the formation of these carbonate rocks and also to understand their depositional environment and ages. These carbonate rocks contain a diverse fauna with the dominance of foraminiferal assemblages of planktonic foraminifera (Globotruncana sp. and Heterohelix sp., etc.) which indicates they were formed during the Santonian to Maastrichtian age. Based on chemical compositions, these carbonate rocks have been identified as limestone (CaO/MgO > 50.1) to slightly dolomitic limestone (9.1 < CaO/MgO < 50.1). Total rare earth element (REE) contents in these carbonates are variable (22.39–146.05 ppm). The Post‐Archean Australian Shale (PAAS)‐normalized REE + Y patterns of these carbonates exhibit seawater‐like REE patterns with LREE depletion and relative HREE enrichment with negative Ce anomalies (Ce/Ce* = 0.32–0.79) and positive Y (4.42–27.81 ppm) and Eu anomalies (Eu/Eu* = 1.11–1.86), suggesting that they were deposited under an oxygenated environment with contamination by hydrothermal activity. They are also depleted in δ13C 0/00 (PDB) (1.02–1.570/00) and δ18O 0/00 (PDB) (−6.37 to −9.00%) values which characterize marine precipitates. Eu anomalies and spread in negative δ18O 0/00 (PDB) values to a lesser extent of δ13C 0/00 (PDB) values of these carbonates suggest their formation was altered by diagenesis in the shallow marine environment. Our new whole‐rock and stable isotope geochemical characteristics, in conjunction with microfacies, suggest that the investigated carbonate rocks might have been formed in low‐energy environments, and deposited in neritic to bathyal palaeoenvironments during the Santonian to Maastrichtian interval.

Upper Aptian–lower Albian of the southern-central Araripe Basin, Brazil: Microbiostratigraphic and paleoecological inferences

The aim of this study was to produce an integrative analysis of macroinvertebrates (decapods, gastropods, and bivalves) and microfossils collected in the Romualdo Formation, south-central Araripe Basin. Additionally, we provide the first report of calcareous nannofossils, a diverse and unique association of the marine ostracods Dicrorygma cf. minuta, Perissocytheridea spp., Dicrorygma (Orthorygma?) dimorpha, and Paracypris sp. and benthic and planktonic foraminifera. The Santo Antonio section contains assemblages of Tethyan origin that indicate an unstable paleoenvironment formed by hypersaline lagoons with two marine ingressions. Planktonic foraminifera assemblages comprised of Hedbergella aptiana and Pseudoguembelitria sp., suggest that the first marine ingression occurred in the Aptian (possibly late Aptian – planktonic foraminifera biozone Paraticinella rohri), while the second occurred during the early Albian (biozone CC8), as indicated by calcareous nannofossils (Hayesites albiensis). The fossil assemblages that characterize the Romualdo Formation, provide new biostratigraphic information and point towards the probable Aptian–Albian boundary in the Santo Antônio section.

Planktonic foraminifera from the Aptian section of the Southeastern Brazilian Atlantic margin

Upper Aptian planktonic foraminifera assemblages recorded in the post-salt section of the Santos, Campos and Espírito Santo Brazilian marginal basins are described and interpreted in terms of biostratigraphy and relative ages. The studied section represents the first marine sedimentary rocks deposited above the evaporitic sequence. Material from 26 Petrobras wells, including core, sidewalls and cuttings samples was analyzed. The upper Aptian planktonic foraminifera assemblages are composed mainly of cosmopolitan and chronostratigraphically long-ranging species, among which 22 species have been commonly recorded in the Tethyan Realm and other South Atlantic basins. Considering the biostratigraphic distribution, deposition of the studied section spans from the upper Aptian Globigerinelloides ferreolensis to Paraticinella rohri zones. Hence, the present study allowed the reassessment of the biostratigraphic framework, pointing out to an upper Aptian age for the marine section deposited immediately above the evaporitic sequence in the Santos, Campos and Espírito Santo basins. Indeed, a marine environment was established 5 to 10 Ma earlier than previously proposed. The reevaluation of the age constraints for the section provides new insight to the understanding of early South Atlantic Ocean geological evolution.

Calcareous plankton biostratigraphic fidelity and species richness during the last 10 m.y. of the Cretaceous at Blake Plateau, subtropical North Atlantic

Species distributions of well-preserved and diverse assemblages of planktonic foraminifera and calcareous nannofossils spanning the last 10 m.y. of the Cretaceous (middle Campanian through Maastrichtian) are analyzed from samples taken across a 1400 m depth transect at Blake Nose in the western subtropical North Atlantic (Ocean Drilling Program Sites 1049, 1050 and 1052). Age models constructed by integrating foraminiferal, calcareous nannofossil, and magnetic polarity datum events provide a reliable framework for temporal correlation of the sites. This framework enables comparisons of species richness and abundance among sites and evaluation of the reliability of first and last appearance datums for regional and global correlation. Among the standard primary zonal marker datums, six of nine planktonic foraminifer and six of seven calcareous nannofossil events are considered reliable for constraining the age-depth models. Secondary datum ages calculated for 17 planktonic foraminiferal events suggest correlation offsets among the three sites of <0.1 m.y. for four species (including Pseudoguembelina praehariaensis Tur and Huber n. sp.), 0.1–0.5 m.y. for six events, and 0.6–1.0 m.y. for eight species. Secondary calcareous nannofossil datum ages calculated for six species show less reliability, with offsets of <0.1 m.y. for one species, 0.7 m.y. for one species, and 1.0–1.5 m.y. for the remaining four species. The distinctly identifiable new species Trinitella suturis Tur and Huber has lowest occurrences that are diachronous by as much as 1.73 m.y. among the Blake Nose sites. Occurrence rarity is the most likely explanation for the age offsets of this and other diachronous species.Planktonic foraminiferal assemblages show no significant differences in species composition and relative abundance among the three sites, suggesting the sites were all located below a single oligotrophic surface water mass. Species richness counts pooled in 200 kyr bins for the Blake Nose sites reveals high species origination rates from the late Campanian through early Maastrichtian and highest species richness (51–63 species) during the Maastrichtian. The only significant extinction pulse during the mid-Campanian through Maastrichtian occurs between 66.2 and 66.4 Ma with loss of five species representing ∼9% of the assemblage. These extinctions occur at the same time as a globally recognized warming event correlated with a pulse of eruptions at the Deccan Traps in India.Calcareous nannofossil assemblages show no significant change in relative abundance among the three sites. Two significant extinction events are documented: one from 75.20 to 75.40 Ma with a loss of six species representing ∼6% of the assemblage and one from 66.2 to 66.4 Ma with a loss of nine species representing ∼9% of the assemblage. The former event is associated with a hiatus at the base of Zone CC24, and the latter corresponds to Deccan Trap warming.Hiatuses are identified at all three Blake Nose sites near the base of the Maastrichtian (∼71.5 Ma; lowermost Pseudoguembelina palpebra/CC24 Zone) and only at the deeper Sites 1049 and 1050 in the mid-Maastrichtian (∼67.2 Ma; Pseudoguembelina hariaensis/CC26b Zone) and the mid-Campanian (∼75.9 Ma; base of Radotruncana calcarata/CC22 Zone). Slumping from across the shelf and slope could have caused the early Maastrichtian hiatus while changes in the pattern and strength of deep-water circulation may have been responsible for the mid-Maastrichtian and mid-Campanian hiatuses.

A Preliminary Study In Vertical Distribution Of Planktonic Foraminifera And Marine Ecological Conditions Of Simeulue Sub-Basin, Aceh, Indonesia

A marine sediment core EW17-09 (03˚ 28’358” latitude and 96˚ 18’788” longitude, 870 m water depth, 390 cm core length) was retrieved from the western Sumatra, Simeulue sub-basin, Indonesia. Simeulue sub-basin are situated in eastern Indian Ocean, western part of Aceh Province, which is one of the outer islands in Indonesia. This sub-basin is influenced by adjacent lands in response to tectonic and climate dynamics. The dynamics of marine ecological conditions in the past is an urgent need for providing an analogy to the changes in the future conditions. In this study, the ecological conditions were examined by identifying the vertical distribution of planktonic foraminifera assemblages. This preliminary study demonstrated the presence of Globigerinoides ruber, Neogloboquadrina dutertrei, Pulleniatina obliquiloculata, Globigerina calida calida, Globigerinoides elongates, Globigerinoides cyclostomus and Globigerinoides sacculiferus in the samples. The assemblages indicate warm water conditions prevailed in the Simeulue sub-basin during the deposition of the samples. However, subtle ecological changes might have occurred in response to the dynamic of thermocline layer. Cluster analysis of planktonic foraminifera abundance and diversity resulted in three groups showing different ecological conditions. Warm water conditions, high salinity, deeper thermocline with moderate sedimentation disturbance prevailed during the deposition of lower part of the core. Oligotropic water conditions with higher temperature, lower salinity, shallower thermocline layer, and moderate sedimentation disturbance predominated during the deposition of the middle part of the core. The paleooceanography conditions of the upper part of the core are comparable to the lower part. Nevertheless, there are a shoaling of the thermocline in the end of the period. These conditions may indicate an increase in upwelling fluctuations and may represent a change in the IOD-like mean state of the Indian Ocean.

Upper Eocene planktonic foraminifera from northern Saudi Arabia: implications for stratigraphic ranges

Abstract. The Rashrashiyah Formation of the Sirhan Basin in northern Saudi Arabia contains diverse assemblages of planktonic foraminifera. We examined the biostratigraphy, stratigraphic range and preservation of upper Eocene planktonic foraminifera. Assemblages are well-preserved and diverse, with 40 species and 11 genera. All samples are assigned to the Priabonian Globigerinatheka semiinvoluta Highest Occurrence Zone (E14), consistent with calcareous nannofossil biostratigraphy indicating Zone CNE17. Well-preserved planktonic foraminifera assemblages from the lower part of the upper Eocene are rare worldwide. Our study provides new insights into the stratigraphic ranges of many species. We find older (Zone E14) stratigraphic occurrences of several species of Globoturborotalita previously thought to have evolved in the latest Eocene (Zone E15, E16) or Oligocene; these include G. barbula, G. cancellata, G. gnaucki, G. pseudopraebulloides, and G. paracancellata. Older stratigraphic occurrences for Dentoglobigerina taci and Subbotina projecta are also found, and Globigerinatheka kugleri occurs at a younger stratigraphic level than previously proposed. Our revisions to stratigraphic ranges indicate that the late Eocene had a higher tropical–subtropical diversity of planktonic foraminifera than hitherto reported.

Late Quaternary carbonate preservation in the Andaman Sea versus the Central Indian Basin: A test of dissolution under diverse oceanographic settings in the Indian Ocean

Late Quaternary carbonate dissolution was evaluated by using planktonic foraminifera indices perfect test ratio (PTR) of Goborotalia menardii, G. menardii fragmentation index (MFI), percentage of total resistant species (RSP) and sediment calcium carbonate (CaCO3) content in five sediment cores collected from two diverse oceanographic settings of the Indian Ocean. The cores collected from a depth range of 2000 and 4250 m of the Andaman Sea, a marginal sea in the Northern Indian Ocean and a seamount rise in the Central Indian Basin (CIB) in the abyssal Indian Ocean depict the intensity of selective carbonate dissolution in the glacial-interglacial cycles of the late Quaternary period. The assemblages from the Andaman Sea show little or no dissolution (deepest core AAS 11, water depth: ~3000 m), whereas cores collected from the CIB shows moderate to very high dissolution (SVBC 37, water depth of ~4000 m and core BC 37, water depth of ~4250 m). Excellent carbonate preservation in the Andaman Sea is reflected in the high order of species diversity consisting of thirty-five species. In contrast, selective dissolution in the CIB resulted in poor diversity with just about twenty-three species. The well-preserved planktonic foraminifera assemblages provide a better paleoceanographic record in the Andaman Sea. The carbonate dissolution proxies in the Andaman Sea point towards excellent preservation during Marine Isotope Stage 2 (MIS 2), especially during the deglacial period (Heinrich event 1) attributed to enhanced ventilation and deepening of carbonate compensation depth (CCD). Cores from the CIB demonstrate that the dissolution does not uniformly modify an assemblage due to the differential susceptibility of each species. Selective dissolution is an important issue in interpreting foraminiferal assemblages for climatic and environmental changes. The study reveals that the relationship between G. menardii abundances and its fragments in a single core is an effective parameter to understand the dissolution as its ratio is not affected by climatic or related environmental changes. Utmost care should be taken while using species variation as an evidence for climatic or related environment change from the CIB. The present investigation observes that the MFI can trace deep-sea carbonate dissolution quantitatively at both CIB and the Andaman Sea locations.

Planktonic Foraminiferal biostratigraphy of Jaddala Formation in well (K-152), Northern Iraq

Many species of planktonic foraminifera were identified from the Jaddala Formation from well (K-152)of depth (1010-900 m.) northern Iraq. The recorded planktonic foraminifera assemblages permits the recognition of four biozones these are (from top to bottom):Globigerinatheka subconglobata biozone (p11), Acarinina bullbrooki biozone (p10), Acarinina Pentacamerata biozone (p9) 1- Morozovell aragonensi biozone (p8) and On the basis of biocorrelation with previous works Early to Middle Eocene age confirmed for the Jaddala Formation at the studied section.