The genus Duvaliopsis was established by Jeannel (I928) for a small group o.f endogeno.us, ano,phthalmous trechines trom the Carpathian Mountains and the Transylvanian Alps ,o.f Romania, Czechoslovakia, and Poland. Although earlier authors had classified them with Anophthalm,us Sturm, Trechus Clairville, or Duvalius Delarouz6e (formerly considered a subgenus of Trechus), Jeannel (1928) clearly demonstrated their morphological similarity to Trechoblemus Ganglbauer and to North American cavernicole trechines of the genera Pse.udanophthalmus and Neaphaenops. Duvaliopsis, Pseudanophthalmus, and Neaphaenops were placed in a s6rie phyl6tique de Trechoblemus, united by the common possession o.f certain .characters (I) the mentum is fused to. the prementum; (2) the recurrent portion of the apical groove o.f the elytro.n is usually .connected to or directed t,o.ward the 3rd longitudinal stria; (3) the copulatory sclerites (o.f which there are one or two) are placed laterally (anisotopic), rather than ventrally (isotopic), in the internal sac; and (4) the anterior tibiae are pubescent on the ,outer side. Subsequent to I928, additional genera in North America and Japan have been described which sho.uld probably be allied with this. series (Valentine 1952, Yoshida and Namura 19.52, U6no 1956 and 1958, Barr I96O). In the eastern United States, the largest and most widely distributed genus of cave beetles is Pseudanophthalmus, species .o.f which are no.w known from Indiana, Kentucky, Tennessee, Alabama, Georgia, Ohio, Virginia, West Virginia, and Pennsylvania. Although found only in caves up to the present time, a few Virginia species have rudimentary eyespots, suggesting comparative recency of adopting a wholly subterranean mode of life. The absence of epigean trechines from North America which seem to share a relatively recent ancestry with Pse.udanoDhthalmus and other cave genera has provoked considerable speculation on the history and evolution of the, group. Trechofdes fasciatus Mo.tschulsky, from the Oligocene Baltic amber, could belong either to Lasiotrech,us or Trechoblemus (Jeannel, 928). This fossil demon
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