To evaluate responses of range grasses to herbage removal, removal patterns should simulate those under grazing. We compared responses of western wheatgrass (Agropyron smithii Rydb.) in mixed-grass range to no clipping, conventional clipping, and clipping which simulated continuous grazing. Two years of simulated grazing did not affect herbage production or tiller numbers, but both declined under conventional clipping. Belowground phytomass decreased as herbage removal increased. Total nonstructural carbohydrate concentration in rhizomes decreased when utilization exceeded about 40930, but that of roots and crowns decreased only when utilization exceeded 60-70%. Most clipping studies of range grasses make little effort to simulate herbage removal patterns which occur under grazing. Clipping studies usually include clipping at uniform heights, not random heights as occurs under grazing (Jameson 1963), and more severe defoliation than occurs under grazing (Heady 1961). Harris (1976) indicated that frequency, intensity, and timing of defoliation all influence plant responses. Lack of knowledge of these parameters has inhibited design of realistic clipping experiments. Hart and Balla (1982) photographed western wheatgrass (Agropyron smithii Rydb.) tillers at intervals during the grazing season and determined frequency, intensity, and timing of herbage removal. We used this information to design a clipping experiment ‘which simulated grazing more closely than ever before, and compared response of western wheatgrass under simulated grazing to that under conventional clipping and no clipping. Our objective was to provide better guides to grazing management than could be provided from responses to conventional clipping. thread (Stipa comata Trin. & Rupr.). Soils underlying the study plots were a fine-loamy Aridic Argiustoll of the Altvan Series. Stevenson et al. (1984) have given a more complete description of the range sites and vegetation of the research station. Twenty-four plots (25 X 40 cm) were established on an area of native range ungrazed since 1978 and containing450-600 tillers/m* of western wheatgrass. Each plot was bordered by aluminum lawn edging placed into the ground approximately 20 cm to eliminate rhizome spread between plots and between the plots and adjacent native range. Four replications of 6 treatments, in a randomized complete block design, were applied to the 24 plots. The 6 treatments included an unclipped control, clipping western wheatgrass to 2.5 cm 4 times during the summer, and 4 clipping treatments simulating 4 levels of continuous grazing of western wheatgrass. Frequency under simulated continuous grazing was defined as the percentage of tillers clipped 0, I, 2,3, or 4 times, while intensity was the percentage of clipping events at which I / 3,2/ 3, or all the aboveground herbage was removed from a tiller. Two frequencies (Table 1) times 2 intensities (Table 2) gave us 4 simulated grazing treatments.
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