The Middle Permian of southern France and a new protopsyllidioidean family reveals a shared hemipteran fauna with the Permian of the Russian Federation

Abstract

Further investigation of two exposures of middle Permian (Guadalupian) strata in southern France enable us to describe new hemipteran taxa and better resolve their Paleozoic diversity. The new protopsyllidioidean family Lodevopsyllidiidae is described on the basis of the new genus and species Lodevopsyllidium jouaulti from the Guadalupian of Lodève (France). Permospyllidium pumilio sp. nov., a second species of Permopsyllidiidae from the same insect assemblage, is described. The serpentivenid Tomioscarta dominiquea sp. nov. is described from the Wordian of Gonfaron (‘red’ Permian, France). Its closest relative is Tomioscarta surijokovensis from coeval strata of the Russian Federation. The prosbolid Kaltanetta gonfaronensis sp. nov. is also described from the same insect assemblage at Gonfaron. Its closest relatives are Kaltanetta nigra and Kaltanetta pallida, both from the Roadian of the Russian Federation. These distributions suggest strong affinities in the middle Permian hemipteran faunas between Western Europe and Central Asia. We discuss the definition and taxa included in Serpentivenidae as encompassing Serpentivena and Tomioscarta and excluding Cercopidium mimas. Mathieu Boderau [mathieuboderau@gmail.com], Romain Garrouste [romain.garrouste@mnhn.fr], and André Nel* [anel@mnhn.fr], Institut de Systématique, Évolution, Biodiversité (ISYEB) Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, CP50, 57 rue Cuvier 75005, Paris, France.