Abstract

1. The premeiotic protoplast shrinks away from its angular sporogenous walls. Within the space created it secretes a special external membrane that functions as the wall of the spore mother cell. 2. A marked contraction of the nuclear network is preliminary to the reduction division. This synizetic knot displays a large, orbicular, deeply staining nucleolus that rests at the polar border of the nucleus. 3. The nucleolus retreats from the polarized nuclear border toward an opposite antipolar region in the nucleus. Anastomosing freely with the nuclear membrane, the spireme recovers as coarse, looped strands centered in the nucleolus or in its disassociated parts. 4. There is no evidence of the pairing of spiremes, nor that the post-synizetic spireme is a continuous thread. Suffering constant elimination of chromatin, it is reduced to slender irregular strands. 5. The chromosomes are short thick rods. They are not derived by the segmenting of a visibly synaptic spireme, but arise as localized masses in the l...

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