Abstract
Shallow subterranean habitats (SSHs) are aphotic subterranean habitats relatively close to the surface ( milieu souterrain superficiel (MSS), and consist of the spaces between rocks, rock cracks, and the like. All of these habitats are more variable than caves, typically with a pronounced annual temperature cycle. Available organic matter is also higher in SSHs than in caves. The importance of SSHs as gateways to the subterranean realm is not known, but it is clear that they contain species both modified for subterranean life, and unique to these habitats.
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