Abstract

Surprisingly high levels of within-group cooperation are observed in conflict situations. Experiments confirm that external threats lead to higher cooperation. The psychological literature suggests proximate explanations in the form of group processes, but does not explain how these processes can evolve and persist. The authors provide an ultimate explanation, in which cooperation is a rational response to an external threat. In the model, groups vary in their willingness to help each other against external attackers. Attackers infer cooperativeness of groups from members' behavior under attack and may be deterred by a group that bands together against an initial attack. Then, even self-interested individuals may defend each other when threatened in order to deter future attacks. A group's reputation is a public good with a natural weakest-link structure. The model extends to cooperative and altruistic behavior in general.

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