Reproduction of the Middle Spotted WoodpeckerDendrocopos mediusin the Nerussa-Desna Woodland, SW Russia, with Particular Reference to Habitat Fragmentation, Weather Conditions and Food Supply

Abstract

The Nerussa-Desna woodland (south-western Russia) is one of the core areas for the Middle Spotted Woodpecker Dendrocopos medius, a habitat specialist restricted mainly to mature oak forests. Recent studies have shown a lower breeding density of the species in fragmented compared with continuous habitat. To evaluate reproduction as a possible mechanism underlying the fragmentation effects, we studied reproductive performance of the species in both continuous habitat and fragmented habitat in the forested landscapes of the Nerussa-Desna woodland. During 1997–2001, annual variation of reproductive indices in both continuous and fragmented habitat was insignificant. None of the indices was significantly lower in fragmented habitat though nestling losses were significantly higher in fragments. Timing of breeding in 2000 was later in fragmented than continuous habitat, probably due to differential landscape phenology. In fragments, fledging success in a year was inversely related to the mean amount of rain. On ...