Abstract
The Timpanoga complex is a holophyletic group of mayflies in the subfamily Ephemerellinae consisting of 21, mostly North American species, distinguished by the absence of lamellate gills on abdominal segment 3 in the larvae, and associated absence of the gill socket remnants in subimagos and adults. A phylogenetic reclassification is proposed, based on a cladistic analysis, and is an example of reclassification necessary in much of the Ephemeroptera. Of the options available for generic classification, the one proposed is both informative in terms of expressing diversity within the complex and practical in terms of allowing morphological differentiation in all life stages, including both sexes of alate stages. Three main sequential lineages are recognized as the genera Attenella, Eurylophella, and Timpanoga [=Dannella n. syn.]. The two main phylogenetic branches in Eurylophella are treated as subgenera Dentatella and Eurylophella s.s. The two main phylogenetic branches of Timpanoga are treated as subgener...
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