Abstract
Photosystem I (PSI) is one of the key players in the process of oxygenic photosynthesis. This large membrane protein complex utilizes light energy to transfer electrons from the lumenal electron carriers plastocyanin or cytochrome c6 across the photosynthetic membrane to the stromal/cytosolic electron carriers ferredoxin or flavodoxin. The resulting proton gradient is used for adenosine triphosphate (ATP) production by the ATP synthase, while the electrons end up in carbon fixation. With a molecular weight of 1 million Da, trimeric cyanobacterial PSI is one of the largest membrane protein complexes with known structure. About one-third of its molecular weight comes from cofactors, mainly chlorophylls, and also carotenoids, phylloqinones, and iron–sulfur clusters.
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