Patterns of size variation over time in ponderosa pine stands established at different initial densities

Abstract

We used six metrics of size and growth variation (standard deviation (SD), coefficient of variation (CV), skewness coefficient (S), Gini coefficient (G), Lorenz asymmetry coefficient (LAC), and growth dominance coefficient (GD)) to describe changes in two long-term ponderosa pine (Pinus ponderosa Douglas ex P. Lawson & C. Lawson) initial spacing trials in Oregon and Washington, USA. Trends were examined over a 35-year time period and across a range of initial stand densities (from 154 to 2470 trees·ha−1) for four measures of tree size: diameter at breast height (dbh, 1.37 m), basal area (BA), height, and volume. Unlike many previous studies of size variation in monospecific stands, our results suggest that variation declined or remained relatively stable for all treatments at both study areas. This suggests that these stands are experiencing size symmetric competition for belowground resources. We found that a combination of metrics is necessary to provide a complete picture of size variability and differentiation in developing stands. We recommend using the CV or G, as there were clear trends with increasing density for all size variables. If the objective of the assessment was to track changes in absolute size within an individual stand, we would recommend using the SD, as there were consistent trends with time for all size variables. S, LAC, and GD may be less suited for comparing differentiation during the early stages of stand development because of a lack of clear trends with stand density and time.