Abstract

The fundamental reaction in photosynthesis of green plants is the photolysis of water into oxygen (O 2 ) and hydrogen (H) (Ruben, Randall, Kamen & Hyde 1941) (see figure 8). The over-all process is driven by the energy of visible light. The fight is absorbed by photochemically active chlorophylls. Only about 0.1% of the chlorophyll is active (Emerson & Arnold 1932). The bulk of the chlorophyll ( Chl 0 ) ( ~ 99 %) and the carotenoids are arranged in such a way that fight absorbed by them is transmitted to the active chlorophylls by energy migration (Gaffron & Wohl 1936; Förster 1947; Dutton, Mannig & Duggar 1943; French & Young 1952; Duysens 1952). The photolysis of water can also take place outside the cell in chloroplasts or chloroplast fragments using artificial (H)-aceeptors (Hill 1939). It has been demonstrated that in chloroplasts hydrogen can be accepted by triphosphorydine nucleotide ( NADP ) (Vishniac & Ochoa 1952; San Pietro & Lang 1956) and that simultaneous production of one ATP accompanies the formation of one NADP H 2 (Arnon i960). With 2 NADP H 2 and at least 3 ATP CO 2 can be reduced to sugar via the Calvin cycle (Calvin, 1962). For the generation of the third ATP molecule light energy must be used which is not accompanied by NADP H 2 -production (Arnon 1960).

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