Abstract
Owing to the intensity of game management in Britain, managers of grouse moors have illegally killed raptors to increase the numbers of Red GrouseLagopus l. scoticaavailable for shooting. This paper summarizes evidence for the recent scale of illegal raptor killing on grouse moors and its effects on populations. It provides insights into how raptors themselves respond demographically to different levels of killing. Over Britain as a whole, most raptors have increased and expanded considerably since the 1970s, in response to reduced killing and nest destruction, and the diminished impacts of organo‐chlorine pesticides; however, in recent decades the populations of some species have declined on and around grouse moors. This is widely evident in Hen HarrierCircus cyaneus, Peregrine FalconFalco peregrinusand Golden EagleAquila chrysaetospopulations and in more restricted areas also in Northern GoshawkAccipiter gentilisand Red KiteMilvus milvuspopulations, in all of which illegal killing has been sufficient to affect numbers over wider areas. The evidence consists mainly of: (1) greater disappearance of nesting pairs, lower breeding densities or reduced occupancy of apparently suitable traditional territories on grouse moors compared with other areas; (2) reduced nest success compared with other areas; (3) reduced adult survival compared with other areas; (4) reduced age of first breeding, reflecting the removal of adults from nesting territories and their replacement by birds in immature plumage; (5) greater levels of disappearance of satellite‐tracked birds on grouse moors than elsewhere; and (6) the finding of poisoned baits and traps, and shot or poisoned carcasses of raptors. Not all these types of evidence are available for every species, and other types of evidence are available for some. The Common BuzzardButeo buteois currently the most numerous raptor in Britain and also seems to be killed in the greatest numbers. Other raptor species, including MerlinFalco columbarius, Common KestrelFalco tinnunculusand Eurasian SparrowhawkAccipiter nisuswhich nest on or near grouse moors, have little or no significant impact on grouse and are killed less often or not at all. In the absence of illegal killing, some raptor species breed as well or better on grouse moors than in other habitats. Merlins, in particular, seem to thrive on grouse moors, benefiting from the management involved (including predator control). Other aspects of illegal raptor killing are discussed, including suggestions for ways in which it might be reduced.
Highlights
Owing to the intensity of game management in Britain, managers of grouse moors have illegally killed raptors to increase the numbers of Red Grouse Lagopus l. scotica available for shooting
Since the mid-20th century, following declines in illegal killing and organo-chlorine pesticide use, population recoveries became apparent in most raptor species in Britain, and those associated with grouse moors
White-tailed Eagles and Goshawks which had been eradicated altogether were reintroduced, and Red Kites were obtained from various parts of Europe for release in several different areas
Summary
IAN NEWTON* Centre for Ecology & Hydrology, Benson Lane, Crowmarsh Gifford, Wallingford, OX10 8BB, UK. In upland areas unsuitable for agriculture, the main game species is the Red Grouse Lagopus lagopus scotica (the subspecies of Willow Ptarmigan Lagopus lagopus that is endemic to Britain and Ireland), which cannot be raised in captivity For this species, areas of open moorland are managed to provide a plant community dominated by heather ( Calluna vulgaris, the main food plant of the species), parasites are controlled mainly by chemical means, and predators are systematically killed (Hudson 1992, Watson & Moss 2008, Werritty 2019, Newton 2020). This paper, based primarily on the scientific literature, summarizes evidence on the scale of this killing on grouse moors and its effects on raptor populations in Britain It does not discuss the impact of raptors on grouse numbers (for which see Redpath & Thirgood 1997, Park et al 2008, Newton 2013, 2020, Roos et al 2018, Francksen et al 2019). The ways in which illicit raptor killing has been studied scientifically, and its various impacts on the demography of raptor populations, are of wider interest
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