Abstract
In 1897, Salvador Calderón y Arana published a study of the first vertebrate footprint from Spain. It was found in the Buntsandstein of Rillo de Gallo (Guadalajara), probably in the Rillo Mudstone and Sandstone Formation (middle Anisian-Landian). This isolated left pes imprint was classified as “Cheirotherium” sp. We conducted a historical study of this specimen, mentioned by several authors for more than a century, but never analyzed in detail. This well-preserved footprint, in which skin marks are present, is described here. Its characters are compared with those of valid chirotheriid ichnogenera. This work is complemented with a bibliographic analysis of the chirotheriid footprints recorded in Spain. This record is remarkable for its abundance and ichnodiversity.
Published Version
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