Abstract

1 West Liberty University, Department of Natural Sciences and Mathematics, P.O. Box 295, West Liberty, WV 26074, USA 2 U.S. Geological Survey, West Virginia Cooperative Fish and Wildlife Research Unit, 322 Percival Hall, Morgantown, WV 26506, USA 3 Carnegie Museum of Natural History, Section of Invertebrate Zoology, 4400 Forbes Avenue, Pittsburgh, PA 15213-4080, USA 4 Midwest Biodiversity Institute, 4673 Northwest Parkway, Hilliard, OH 43026, USA

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