A taxonomic review of the genus Joryma Bowman and Tareen, 1983 (Crustacea: Isopoda: Cymothoidae) parasitizing the marine fishes from Indian waters, with a description of a new species

Abstract

The genus Joryma is here revised, with a description of a new species collected from the host Escualosa thoracata (Clupeidae) off the Malabar Coast of the Arabian Sea. Joryma malabaricus sp. nov. is described from female, male, transitional, pre-manca, and manca stages and the female stage is distinguished from other Joryma species by the following female characteristics: cephalon conspicuous dorsally and extending beyond pereonite 1 expansion, pereonite 1 anterolateral expansion unilateral and not bilobed, laterally overlapped pleonites 1 and 2, triangular pleotelson, uropod rami unequal in length; exopod slightly longer than endopod, mandibular palp not segmented in female (three segments in male), four recurved spines in maxilliped palp. A neotype was designated and redescribed for J. engraulidis, with an additional description from fresh material. Joryma sawayah and Joryma hilsae are redescribed from the female stage and described from the male stage respectively. The type material of Joryma tartoor and Joryma brachysoma is lost and after their original description, both species have not been reported. A key to the species of Joryma is also presented.