Abstract

Abstract. Material belonging to a new fossil genus and species of caddisfly, Pulchercylindratus punctatus n. gen., n. sp., was collected from the Daohugou locality (Middle Jurassic, Jiulongshan Formation; Inner Mongolia, China). The new species is assigned to the Hydrobiosidae according to subcylindrical shape of the 2nd segment of maxillary palp, the forked R1 (in the forewing, located near apex), and long anal cells (in the forewing). In addition, we propose to transfer the genus Juraphilopotamus Wang, Zhao & Ren 2009, known from the same locality, to the family Hydrobiosidae, based on the 1st and 2nd segments of the maxillary palp being cylindrical, shorter than the 3rd segment. A Middle Jurassic origination of family Hydrobiosidae can be established based on the new discovery. doi: 10.1002/mmng.201300005

Highlights

  • The order Trichoptera, so-called “hairy wings”, contains three suborders, namely Annulipalpia, Integripalpia, and Spicipalpia, with over 13,000 known extant species

  • The order is widely distributed around the world, except for the polar regions (Morse 2012), and has an extensive fossil record, with 37 families, 193 genera, and more than 660 species reported to date (Handlirsch 1906–1908, 1939; Meunier 1918; Martynova 1958; Sukastcheva 1968, 1973, 1982, 1990; Erickson 1983; Novokshonov 1993; Botosaneanu 1995; Jarzembowski 1995; Novokshonov et al 1995; Ansorge 2002; Ivanov & Sukatsheva 2002; Ivanov & Melnitsky 2005; Wichard 2007; Wichard et al 2009, 2011)

  • The Spicipalpia consists of five extant families, namely Rhyacophilidae, Hydrobiosidae, Glossosomatidae, Hydroptilidae and Ptilocolepidae (Ross 1956; Schmid 1970; Morse 1997; Malicky 2001, 2005; Ward et al 2004)

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Summary

Introduction

The order Trichoptera, so-called “hairy wings”, contains three suborders, namely Annulipalpia, Integripalpia, and Spicipalpia, with over 13,000 known extant species. The Spicipalpia consists of five extant families, namely Rhyacophilidae, Hydrobiosidae, Glossosomatidae, Hydroptilidae and Ptilocolepidae (Ross 1956; Schmid 1970; Morse 1997; Malicky 2001, 2005; Ward et al 2004). The family Hydrobiosidae was first erected by Ulmer (1905) as a subfamily of Rhyacophilidae, and granted with family rank by Schmid (1989). This is a large family of approximately 50 genera (Holzenthal et al 2007). Only three genera and three species of the fossil Hydrobiosidae have been reported (Table 1)

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