Abstract

One of the central puzzles in the study of sociocultural evolution is how and why transitions from small-scale human groups to large-scale, hierarchically more complex ones occurred. Here we develop a spatially explicit agent-based model as a first step towards understanding the ecological dynamics of small and large-scale human groups. By analogy with the interactions between single-celled and multicellular organisms, we build a theory of group lifecycles as an emergent property of single cell demographic and expansion behaviours. We find that once the transition from small-scale to large-scale groups occurs, a few large-scale groups continue expanding while small-scale groups gradually become scarcer, and large-scale groups become larger in size and fewer in number over time. Demographic and expansion behaviours of groups are largely influenced by the distribution and availability of resources. Our results conform to a pattern of human political change in which religions and nation states come to be represented by a few large units and many smaller ones. Future enhancements of the model should include decision-making rules and probabilities of fragmentation for large-scale societies. We suggest that the synthesis of population ecology and social evolution will generate increasingly plausible models of human group dynamics.

Highlights

  • Human societies exhibit different social and cultural organizations and technologies [1]

  • Under the baseline conditions with an initial random patch distribution, multicellular group (MCG) emerged after about 100 years and the cells belonging to MCGs continued increasing in number, whereas Single-celled groups (SCG) initially increased in number, attaining constant populations before declining (Fig 3A)

  • Based on our numerical experiments we suggest that given enough time, a single MCG will eventually come to dominate more than 50% of the landscape

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Summary

Introduction

Human societies exhibit different social and cultural organizations and technologies [1]. Social or religious groups furnish examples of adaptations to different niches by cultural differentiation. They have heritable characteristics and are subject to selection, such that those social and/or cultural features providing greater competitive fitness to their groups tend to prevail [3,4]. In this respect, sociocultural groups resemble living organisms: they emerge, grow, interact, reproduce, and eventually dissolve or die. We develop a lifecycle theory for the emergence and evolution

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