In Africa south of the Zambezi River, archaeologists and other experts have long explored the impact of climate and environmental changes to the development of ancient civilizations during the Iron Age (CE 200–1900). Some of the prevailing thought is however still rooted in environmental deterministic models informed by selected ethnographies, stable isotopes and archaeological evidence. For instance, the drought brought by the medieval Little Ice Age is assumed to have collapsed the civilisation at Mapungubwe in the Shashi-Limpopo valley around 1300 CE. And yet, within the wider region, and in similar ecological settings, upstream (Shashi and Upper Limpopo) and downstream civilisations (Lower Limpopo), persisted and thrived through the same climatic challenges. We draw on African cosmologies, resilience theory and archaeological evidence from Mapela and Little Mapela to spotlight adaptation strategies utilised by their inhabitants to build resilience through time. The main conclusion is that even in cases of climatic extremes, humans responded to opportunities and constraints in context specific ways.