Upstream Extent Research Articles

Emigration of post-spawned twaite shad Alosa fallax, an anadromous and iteroparous fish, in a highly fragmented river.

Anthropogenic barriers are widely known to negatively impact the spawning migrations of anadromous fishes, by delaying or preventing passage upstream. Although the impacts of barriers on emigrating post-spawned adults are less well studied, they could potentially impact the fitness and subsequent return rates of iteroparous species. In this study, passive acoustic telemetry was used to track the emigrations of 53 twaite shad Alosa fallax in the River Severn basin in their first spawning migration a year after being tagged, giving insights into their emigration movements and the impacts of anthropogenic weirs on these movements. A. fallax began their emigrations after spending varying amounts of time and migrating various distances within the river, with late-emigrating individuals moving fastest and most directly. Emigrations became faster and more direct the further downstream individuals were from their furthest upstream extent. Downstream passage delays at weirs increased emigration times by a median of 61%, with environmental conditions (i.e., temperature, flow, and tidal influence on river level) having little influence on downstream passage at weirs with no modifications to facilitate fish passage. As weir-induced emigration delays are suggested to deplete energy reserves (when energy levels are already depleted post-spawning), limit spawning opportunities (by preventing access to downstream spawning habitat), and expose individuals to increased predation risk and suboptimal conditions (e.g., high temperatures), these delays can potentially diminish the benefits of iteroparity. The evidence presented here suggests that more consideration should be given to the potential impacts of anthropogenic barriers on the emigrations of iteroparous species when assessing river connectivity or undertaking barrier mitigation.

WURRANDERRA’S SYMBOLS: AN EXPLORATION AND CONTEXTUALISATION OF THE THURK PETROGLYPH SITE (KINGSTON-ON-MURRAY) ON THE MURRAY RIVER, SOUTH AUSTRALIA

This article describes and contextualises the rock art at the Thurk Petroglyph Site on the Murray River in South Australia using multiple methods. The Thurk Aboriginal engravings comprise at least 524 motifs made up predominantly of geometric line elements as well as a small number of other ‘simple’ geometric motifs, two ‘bird tracks’, one figurative design (a ‘fish’) and a possible anthropomorphous figure. This paper provides the first synthesis of rock art sites/complexes and motifs from other sites on the Murray River as well as visual symbols recorded from senior Aboriginal ‘knowledge carriers’. These syntheses allow us to consider the relationship of Thurk to other cultural places and to highlight and honour the traditional knowledges and beliefs which underpin the rock art. Thurk’s placement within the riverscape, its unique geological canvas, lack of observable ‘domestic’ archaeological evidence combined with it being the likely upstream extent of Murray River rock art in South Australia contribute additional dimensions to its cultural significance. That Thurk’s rock art, and the broader site, have been desecrated by gratuitous graffiti, vandalism and infrastructure brings into sharp focus Australia’s poor record of heritage protection and provides a challenge to current and future generations of non-Aboriginal people to remedy this past.

The Importance of Phenology and Thermal Exposure to Early Life History Success of Nonnative Smallmouth Bass in the Yellowstone River

AbstractKnowledge of potential spread by introduced species is critical to effective management and conservation. The Smallmouth Bass Micropterus dolomieu is an example of a fish that has been introduced globally, often spreads after introduction, and has substantial predatory impacts on fish assemblages. Nonnative Smallmouth Bass in the free‐flowing Yellowstone River, Montana, have expanded from warmer, downstream sections of river into colder, upstream sections containing socio‐economically valuable trout fisheries. We sought insight into mechanisms controlling upstream spread by evaluating whether progressively colder upstream climates physiologically constrained successful recruitment by limiting age‐0 growth and preventing overwinter survival (i.e., population establishment). We documented the phenology, growth, and overwinter survival of age‐0 Smallmouth Bass across a temperature gradient leading to their upstream extent in the Yellowstone River. The upstream extent of population establishment did not appear limited by water temperature alone. Age‐0 body size at the onset of winter did not differ significantly between colder, upstream reaches and warmer, downstream reaches. Instead, the earlier hatch timing exhibited by some age‐0 individuals in upstream sections allowed them to experience longer growing seasons than many individuals in downstream sections. This counter‐intuitive hatching phenology mediated much of the expected decreases in growth in colder, upstream climates. Furthermore, evidence of successful overwinter survival and simulations of age‐0 starvation mortality indicated that age‐0 individuals at the upstream extent of their distribution successfully recruited to the age‐1 year‐class during four consecutive years. However, age‐0 individuals were rare or absent throughout the uppermost upstream distribution of adults, suggesting that something other than temperature limits or discourages reproduction farther upstream. Taken together, our results suggest that Smallmouth Bass have not yet reached the thermal limit of their upstream distribution in the Yellowstone River and that future spread may challenge fisheries managers tasked with management of coldwater trout fisheries in this river.

A Theory for the Response of Tropical Moist Convection to Mechanical Orographic Forcing

Abstract Spatial patterns of tropical rainfall are strongly influenced by mountains. Although theories for precipitation induced by convectively stable upslope ascent exist for the midlatitudes, these do not represent the interaction of moist convection with orographic forcing. Here, we present a theory for convective precipitation produced by the mechanical interaction of a tropical ridge with a basic-state horizontal wind. Deviations from this basic state are represented as the sum of a “dry” perturbation, due to the stationary orographic gravity wave, and a “moist” perturbation that carries the convective response. The moist component dynamics are subject to the weak temperature gradient approximation; they are forced by the dry mode’s influence on lower-tropospheric moisture and temperature. Analytical solutions provide estimates of the precipitation distribution, including peak precipitation, upstream extent, and rain shadow extent. The theory can be used with several degrees of complexity depending on the technique used to compute the dry mode, which can be drawn from linear mountain wave theory or full numerical simulations. To evaluate the theory, we use a set of convection-permitting simulations with a flow-perpendicular ridge in a long channel. The theory makes a good prediction for the cross-slope precipitation profile, indicating that the organization of convective rain by orography can be quantitatively understood by considering the effect of stationary orographic gravity waves on a lower-tropospheric convective quasi-equilibrium state.

Where rivers jump course.

Rivers can abruptly shift pathways in rare events called avulsions, which cause devastating floods. The controls on avulsion locations are poorly understood as a result of sparse data on such features. We analyzed nearly 50 years of satellite imagery and documented 113 avulsions across the globe that indicate three distinct controls on avulsion location. Avulsions on fans coincide with valley-confinement change, whereas avulsions on deltas are primarily clustered within the backwater zone, indicating a control by spatial flow deceleration or acceleration during floods. However, 38% of avulsions on deltas occurred upstream of backwater effects. These events occurred in steep, sediment-rich rivers in tropical and desert environments. Our results indicate that avulsion location on deltas is set by the upstream extent of flood-driven erosion, which is typically limited to the backwater zone but can extend far upstream in steep, sediment-laden rivers. Our findings elucidate how avulsion hazards might respond to land use and climate change.

Fish assemblage response to removal of a low‐head dam in the lower reach of a tropical island river

Abstract Dams are often removed from rivers to restore habitat connectivity for biota such as fish. Removal of inland dams is well studied in temperate mainland rivers but this approach has been little studied in fish assemblages in islands, tropic systems, or for dams near the mouth of the river. In Puerto Rico, one of the most intensively dammed territories in the world, all native river fishes migrate between fresh water and the sea, and previous work shows that these movements are impeded or blocked by dams. Fish assemblages were compared before and after removal of the Cambalache dam, a porous, low‐head structure near the mouth of the Río Grande de Arecibo, as well as in two other rivers in western Puerto Rico, one with a similarly sized and positioned dam, and one reference river without artificial barriers. Fish were sampled using backpack electrofishing on 39 occasions during 2017–2019, including seven samples collected after removal of the Cambalache dam, at four to six sites per river. Fish assemblages upstream from dams were poorer in species, and species richness showed a marginal tendency (p = 0.0515) to increase upstream of the Cambalache dam 3 months after its removal. The two small lowland dams studied herein limited the upstream extent of marine species, which recolonised upstream sites of the Río Grande de Arecibo after removal of the Cambalache dam. An estimate of relative density (catch per unit effort) of common native freshwater species was higher above these two dams, and decreased at upstream sites after removal of the Cambalache dam. The estimated relative density of a native freshwater species that is of conservation concern, the American eel (Anguilla rostrata), was reduced above dams, and increased upstream of the former Cambalache dam after its removal. In extensive surveys conducted previously in Puerto Rico, sampling was concentrated higher in the catchment, and native fishes were more common and abundant below than above dams. The present work was conducted near the river mouth, and opposite results were observed. These contrasting results suggest that the effects of dams (or dam removal) on fish assemblages vary along the river gradient, although data from other systems are needed to confirm this. The present results suggest low‐head dam removal to be a viable method of restoring connectivity in fish assemblages in lower reaches of rivers in Puerto Rico and, potentially, other tropical islands. Removal of dams near the mouth of the river appears to be of particular benefit to marine fish species that use lower river reaches.

Riverscape View of Fish Assemblages, Habitat, and Stream Temperatures during Summer Low Flows in the Chehalis River, Washington

The Chehalis River is a coastal river in southwestern Washington State where decision makers are considering options to reduce flood damage and improve freshwater habitats. This study focused on the mainstem river above and below the location of a potential dam, one of the flood-reduction options being considered. Our objectives were to describe spatial patterns of fish species, physical habitat, and stream temperature, and identify associations between fish species and the physical environment at two spatial scales. Data were collected in spatially continuous reaches following a “riverscape” approach, and patterns in the data were described using three different ordination techniques. Most of the variation in fish species composition occurred at the sub-basin scale (entire study area), with additional but minor patterns at the reach (1-km) scale. At the sub-basin scale, fish species composition was organized in an upstream (salmonid) to downstream (cyprinid) replacement pattern and was best explained by the combination of river location, habitat, and temperature. At the reach scale, physical habitat and stream temperature differentiated juvenile trout versus coho salmon in the salmonid-dominated upstream extent of our study area, and dace versus shiner in the cyprinid-dominated downstream extent. We conclude that salmon and steelhead are particularly vulnerable to habitat loss upstream of the proposed dam because they disproportionately use this area as summer rearing habitat. Protection and restoration of headwater areas, such as our study area in the Chehalis River, will be critical to support salmonid populations into the future.

Individual movement variation in upstream‐migrating sea lamprey Petromyzon marinus in a highly fragmented river

Abstract Anthropogenic reductions in riverine connectivity can severely impede the migrations of anadromous species. In fragmented rivers, successful migrations depend on the ability of migrants to negotiate barriers or locate alternative passage routes. However, individual variation in the specific aspects of movement that determine migration success in fragmented rivers, is poorly characterised. Here, individual variation was investigated in the spawning migrations of 56 adult sea lamprey Petromyzon marinus, an anadromous, semelparous species that does not show fidelity to natal rivers. The variability and consequences of two key aspects of fish migration within fragmented rivers were tested: passage time (total time taken to pass a barrier) and retreats (exploratory downstream movements after unsuccessful passage attempts). These were tested using acoustic telemetry in the highly fragmented River Severn catchment, western England, U.K. Distinct unimodal, bimodal, and multimodal patterns of variation in passage times were displayed across the different barriers, potentially related to the physical characteristics of the barrier and prevailing river discharge conditions when the barriers were first approached, but were not related to lamprey body sizes. At the first three barriers encountered by upstream‐migrating sea lamprey in the study, between 30% and 46% of individuals made retreat movements, and between 5% and 100% of retreating individuals were able to locate alternative tributaries. Retreating individuals were highly variable in their distance and frequency of retreats; overall, retreat movements comprised 11% (lower‐upper quartiles 0%–52%, range 0%–76%) of the total distance moved before reaching spawning areas. Time‐to‐event analysis indicated that retreat rates reduced as river discharge increased. There was no evidence indicating that individual variation in passage time, or presence of retreat movements at barriers, influenced the subsequent upstream migration speed or final upstream extent of lampreys. While predictability in rank arrival timing was high within three unobstructed reaches, this predictability was disrupted at barriers due to individual variation in passage times. Anthropogenic barriers thus can both disrupt and reveal individual variation in the migration dynamics of anadromous species. Substantial variability in retreat behaviours can be displayed by anadromous species facing delays at barriers, with these behaviours also associated with environmental conditions and the availability of alternative migration routes. Individual variation in exploration and passage time of migrants strongly influence their eventual spawning distribution.

Invasion of Virile Crayfish Faxonius virilis (Hagen 1870) in the Lower Henrys Fork Drainage, Idaho

The invasive virile crayfish Faxonius virilis (Hagen 1870) has recently been documented in the upper Snake River drainage of Idaho, but its distribution is poorly known. Our objective was to determine the presence and distribution of F. virilis in the Henrys Fork drainage of the Snake River. Sampling was conducted during summer and autumn 2018 at 30 sites located in Henrys Fork River drainage, including the major tributaries Teton River, North Fork Teton River, South Fork Teton River, and Moody Creek. We used baited minnow traps and kick nets to determine presence of F. virilis. Absence was only reported if we unsuccessfully captured F. virilis using kick nets because this technique is more effective. Faxonius virilis was detected in all five streams. We did not detect F. virilis at sites at the upstream extent of sampling. This pattern suggests that F. virilis are invading the drainage in an upstream direction. Presence of species of conservation concern (e.g., Yellowstone cutthroat trout [Oncorhynchus clarkii bouvieri Jordan and Gilbert 1883], bluehead sucker [Catostomus discobolus Cope 1871], and western pearlshell [Margaritifera falcata Gould 1850]) warrant additional research to determine the effect of F. virilis on these species.

Cumulative impacts of habitat fragmentation and the environmental factors affecting upstream migration in the threatened sea lamprey, Petromyzon marinus

Abstract River ecosystems are often fragmented by artificial structures, such as weirs. For anadromous species, these structures can impede access to upstream spawning sites and ultimately lead to severe population declines. This study focused on the freshwater spawning migration of the sea lamprey, Petromyzon marinus, an anadromous species threatened by habitat fragmentation across its native range. To quantify the cumulative impacts of multiple weirs on upstream‐migrating adults, and to explore the environmental factors affecting migratory movements, passive acoustic telemetry was applied to 56 individuals during their spawning migration in the heavily fragmented River Severn basin, UK. While 89% of tagged sea lamprey passed the first weir upstream of the release site on the main river, only 4% passed the fifth weir. For 85% of migrants, the upstream extent of migration was immediately downstream of a weir. Individuals that passed weirs upstream of the release site (n = 50) took 21.6 ± 2.8 days to reach their most upstream location, experiencing cumulative passage times at weirs of 15.7 ± 2.8 days; these delays constituted a median of 84% of total upstream movement times. Multistate models showed that the weir passage rates of sea lamprey in tidal and non‐tidal areas increased significantly when downstream river level and discharge were elevated. Upstream‐to‐downstream changes in direction were frequent downstream of weirs, but rare in unobstructed river sections. The results provided evidence for a cumulative effect of multiple weirs on sea lamprey movements, substantially delaying upstream migrants and limiting their spawning to atypical habitat. The results also demonstrated the crucial roles of high tides and elevated discharge events in enabling weir passage. Although the Severn Estuary features conservation designations for sea lamprey, this study reveals that barriers are inhibiting their upstream migration, a problem that should be addressed to assist sea lamprey conservation.

A Small Proportion of Breeders Drive American Bullfrog Invasion of the Yellowstone River Floodplain, Montana

The American bullfrog (Lithobates catesbeianus) is a non-native invader of aquatic habitats across the northwestern United States. It recently invaded the Yellowstone River, Montana, and has spread to over 140 km of floodplain habitat. We analyzed seven microsatellites in 528 tadpoles sampled across nearly the entire Yellowstone River invasion (about 140 river km) to characterize invasion genetics, compare our results with those of a recent mtDNA study (Kamath et al. 2016), and to inform control efforts. Microsatellite variation supports the mtDNA-based hypothesis of at least two independent introductions to the floodplain from genetically divergent populations in the midwestern United States, followed by massive range expansion. One introduction is associated with the upstream extent of the invasion near Park City, Montana and the other more broadly with downstream populations. All sites were characterized by small effective numbers of breeders (Nb; harmonic mean = 9.97), and therefore, a small proportion of highly successful adults may drive the invasion by producing large families. Microsatellites and mtDNA produced discordant estimates of genetic admixture between the upstream and downstream invasions, which may reflect small effective population size. Although we observed isolation by distance using both types of markers, microsatellites appear to reflect population structure resulting from secondary contact between the two introductions, as opposed to structure resulting from equilibrium between gene flow and genetic drift. Most sites showed evidence for genetic bottlenecks, which supports the recent history of invasion. Small Nb paired with known high localized extinction rates following colonization suggests focused removal of post metamorphic life stages at sites less likely to go extinct on their own could help limit invasion by bullfrogs.

The role of glacial dynamics in the development of ice divides and the Horseshoe Intersection Zone of the northeastern Labrador Sector of the Laurentide Ice Sheet

The Labrador Sector of the Laurentide Ice Sheet is characterized by a complex network of ice divides and an extensive landform record outlining two broad and opposing ice flows that are separated by a narrow Horseshoe Intersection Zone (HIZ). This geomorphic system gave rise to contrasting reconstructions, which reflect uncertainties on the temporal evolution of ice divides, the nature and age of the main landform systems and the overall pattern of ice retreat during the last deglaciation. Here, we address these issues through systematic mapping of glacial landforms and ice-flow indicators in a large area of northeastern Quebec and Labrador. The application of cosmogenic (10Be and 26Al) dating to esker-fed glaciomarine deltas and different rock surfaces brings new constraints on the chronological framework and insights on the subglacial thermal regime. Our results outline four main landform assemblages, two of which relating to a major system of opposite ice flows that delineate the position of the eastern Ancestral Labrador ice divide. Our reconstruction shows that the HIZ occurs several tens of km to the west of this divide and that these two features are genetically distinct. The HIZ relates to the late-glacial development of ice streams showing a massive convergent ice flow into Ungava Bay. The upstream extent of this ice flow system shows a strong coupling with the outline of Ungava Bay drainage divide, suggesting that the configuration of the HIZ could be the expression of a topographic control on the late-glacial ice sheet dynamics. Landforms also indicate a significant shift in the deglaciation mode, which evolved from an overall warm-based to an areally-confined cold-based ice retreat near the center of the former ice mass – a change in dynamics that also played a role in the configuration of the HIZ. Overall, areas indicative of cold-based ice conditions are limited to highly elevated terrains, thus ruling out models arguing for an extensive cold-based ice cover over Ungava Bay throughout the last deglaciation.

Can eDNA metabarcoding offer a catchment-based approach for biodiversity monitoring?

In previous studies eDNA metabarcoding has been demonstrated as a viable tool for catchment-level biodiversity sampling in rivers (Deiner et al. 2016). However, questions still remain over the appropriate sampling protocol for large spatial scale sampling. River reaches are composed of multiple habitats with species composition varying from one to the next (Costa and Melo 2007). Therefore, how many spatial replicates are needed to reliably represent the river network? Is the previously used approach to sample at every river confluence (Deiner et al. 2016) sufficient or is more needed? These questions were addressed using a case study in the headwaters of the Cound Brook, a tributary to the River Severn in Shropshire, UK. Two sub-catchments of the Cound Brook were used. One sub-catchment had a sample taken at the most downstream point before the confluence. Additionally, a sample at the upstream extent of the same sub-catchment was taken to estimate any correlation between the species found at the beginning of the river reach and at the end. Another sub-catchment also had the same up- and downstream sample design. However, in between was a systemic sampling regime every 500 m. This is to test if increasing the spatial resolution gave significantly different results to the sparser sampling regime. At each sample location, a 1 L water sample was sequentially filtered through membranes of three different mesh sizes: 5µm, 0.45µm and 0.2µm. Sequential filtering was performed because DNA resides in two forms in the environment (Civade et al. 2017), within whole cells (cellular DNA) and outside of cells (extracellular). The theory is that the coarser filters predominantly collect cellular DNA and the finer filters collect predominantly extracellular DNA of increasingly smaller fragment lengths. Consequently, sequential filtering could represent DNA degradation in the environment (Fig. 1). Also, Turner et al. 2014 suggested that the larger particles could determine very recent or local organisms. Therefore, we hypothesised that the DNA collected by the coarser filters would represent local diversity and the DNA collected by the finer filters would reflect biodiversity further upstream. Initial results suggest sequential filtering through the 5µm and 0.45µm filters caught detectable levels of eDNA where the 0.2µm did not catch enough to show up through gel electrophoresis. The relevance of the initial finding suggests that if we only used a 5µm filter the data collected at 0.45µm could have been discarded. Further investigations of any differences in species compositions between filters and the relationships to other sampling locations is still to be determined. This ongoing research is intended to determine the appropriate sampling protocol for a large-scale biodiversity assessment regime combining eDNA metabarcoding and species distribution modelling.

Observations regarding Lake Sturgeon spawning below a hydroelectric generating station on a large river based on egg deposition studies

AbstractSpawning behaviour of the Lake Sturgeon (Acipenser fulvescens) in large rivers is poorly understood, complicating the approach to fish‐friendly hydroelectric design and mitigation. In 2006, dam safety concerns prompted the need to modernize the spillway infrastructure of the 100‐year old Pointe du Bois Generating Station on the Winnipeg River, Manitoba. The associated regulatory process provided an opportunity to study Lake Sturgeon spawning in a location characterized by water depths >5 m, high velocities, and hydraulic instability. Over six spawning periods (2007–2012), a wide range of flow conditions occurred, and spawning was documented using egg mats in both the powerhouse tailrace and downstream of the spillway rapids. Herein, we consider “….the preferential utilization by sturgeon of certain ranges of the physical variables studied, but within habitats considered favourable for the reproduction of the species” conclusion forwarded by La Haye et al. as our working hypothesis. Overall, 27,362 of 30,085 (90.1%) Lake Sturgeon eggs were captured during the primary spawning intervals, defined based on 6‐day annual periods encompassing the date of peak deposition. Of the 2,200 mats retrieved, 74.4% captured 0 eggs, 12.0% captured 1–5 eggs, 5.2% captured 6–25 eggs, 6.1% captured 26–99 eggs, and just 2.2% captured ≥100 eggs (high‐yield). Data were examined in relation to status‐quo habitat variables (depth, velocity, and substrate), as well as two additional variables hypothesized to have predictive power: distance from physical and/or energetic barriers and hydraulic complexity. Distance‐based analyses revealed upstream to downstream redistribution of eggs via flow. Spawning (as inferred based on high‐yield mats) was spatially discrete during each spawning period as opposed to well‐distributed among habitats broadly considered to be suitable for the species. We suggest that the current‐edges located proximal to physical and/or energetic barriers (i.e., at the upstream extent of usable habitat) drive Lake Sturgeon spawning site selection in large rivers, and advise caution prior to making fine‐scale inferences when only small quantities of Lake Sturgeon eggs are captured.

Threats to Rearing Juvenile Chinook Salmon from Nonnative Smallmouth Bass Inferred from Stable Isotope and Fatty Acid Biomarkers

AbstractNonnative Smallmouth BassMicropterus dolomieuare increasingly sympatric with juvenile spring Chinook SalmonOncorhynchus tshawytschain the rivers of western North America. Understanding the potential effects of introduced Smallmouth Bass is essential to efficiently direct salmon management efforts, especially given the potential for upstream range expansion of Smallmouth Bass in response to the climate‐induced warming that many rivers may experience. However, the predatory and competitive threats of Smallmouth Bass to juvenile salmonids remain largely unexamined, particularly in salmon rearing areas. To address this knowledge gap, we collected stable isotope (δ13C and δ15N) and fatty acid data to assess the trophic role of Smallmouth Bass in the upstream reaches of the North Fork John Day River, a tributary of the Columbia River where the upstream invasion extent of Smallmouth Bass co‐occurs with the native predator Northern Pikeminnow and river‐rearing juvenile Chinook Salmon. We found that Smallmouth Bass and Northern Pikeminnow occupy divergent dietary niches, and there was little evidence of a strong predation or competition threat to juvenile Chinook Salmon from either predator. In addition, we found a disproportionate reliance on autochthonous resources across the entire community and low isotopic uniqueness, suggesting potential competition among the broader fish community given the scarcity of resources. Importantly, this study focused on the overlap between the upstream‐most habitats of encroaching Smallmouth Bass and downstream‐most habitats of juvenile Chinook Salmon. Therefore, the relative scarcity of rearing salmon in comparison with other available prey in this zone may limit current predation threats. The results from this study help guide resource‐limited managers that are tasked with conserving and restoring native salmonid populations.

Impacts of Sea-level Rise on the Tidal Reach of California Coastal Rivers using the Coastal Storm Modeling System (CoSMoS)

O'Neill, A.C.; Erikson, L.H.; and Barnard, P.L., 2020. Impacts of sea-level rise on the tidal reach of California coastal rivers using the Coastal Storm Modeling System (CoSMoS). In: Malvarez, G. and Navas, F. (eds.), Global Coastal Issues of 2020. Journal of Coastal Research, Special Issue No. 95, pp. 1223-1228. Coconut Creek (Florida), ISSN 0749-0208.In coastal rivers, the interactions between tides and fluvial discharge affect local ecology, sedimentation, river dynamics, river mouth configuration, and the flooding potential in adjacent wetlands and low-lying areas. With sea-level rise, the tidal reach within coastal rivers can expand upstream, impacting river dynamics and increasing flood risk across a much greater area. Rivers along the Pacific coast of California are constrained by coastal mountain ranges, in contrast to the typical large and low-gradient river deltas and estuaries often used in tidal reach studies. Yet, the impacts of tides are ever-present and considerable to these rivers' habitats and neighboring communities. The impacts of a range of sea-level rise, discharge, and coastal storm conditions on the upstream extent of a spring tidal influence and the associated flood potential are assessed for several coastal rivers across California using coupled Delft3D FLOW-WAVE hydrodynamic simulations that explicitly account for the interactions between waves, currents, surge, and water levels in this study. Projected changes in tidal reach among the various rivers are characterized and implications for adjacent communities and stakeholders are discussed.

Beaver dam influences on streamflow hydraulic properties and thermal regimes

Beaver dams alter channel hydraulics which in turn change the geomorphic templates of streams. Variability in geomorphic units, the building blocks of stream systems, and water temperature, critical to stream ecological function, define habitat heterogeneity and availability. While prior research has shown the impact of beaver dams on stream hydraulics, geomorphic template, or temperature, the connections or feedbacks between these habitat measures are not well understood. This has left questions regarding relationships between temperature variability at different spatial scales to hydraulic properties such as flow depth and velocity that are dependent on the geomorphology. We combine detailed predicted hydraulic properties, field-based maps with an additional classification scheme of geomorphic units, and detailed water temperature observations throughout a study reach to demonstrate the relationship between these factors at different spatial scales (reach, beaver dam complexes, and geomorphic units). Over a three-week, low flow period we found temperature to vary 2 °C between the upstream and downstream extents of the reach with a net warming of 1 °C during the day and a net cooling of 0.5 °C at night. At the beaver dam complex scale, net warming of 1.15 °C occurred during the day with variable cooling at night. Regardless of limited temperature changes at these larger scales, the temperature variability within a beaver dam complex reached up to 10.5 °C due to the diversity of geomorphic units. At the geomorphic unit scale, the highly altered flow velocity and depth distributions within primary geomorphic units provide an explanation of the temperature variability within the dam complex and insight regarding increases in habitat heterogeneity.

Flows over surface-mounted bluff bodies with different spanwise widths submerged in a deep turbulent boundary layer

The spatio-temporal dynamics of separation bubbles induced by surface-mounted bluff bodies with different spanwise widths and submerged in a thick turbulent boundary layer is experimentally investigated. The streamwise extent of the bluff bodies is fixed at 2.36 body heights and the spanwise aspect ratio ($AR$), defined as the ratio between the width and height, is increased from 1 to 20. The thickness of the upstream turbulent boundary layer is 4.8 body heights, and the dimensionless shear and turbulence intensity evaluated at the body height are 0.23 % and 15.8 %, respectively, while the Reynolds number based on the body height and upstream free-stream velocity is 12 300. For these upstream conditions and limited streamwise extent of the bluff bodies, two distinct and strongly interacting separation bubbles are formed over and behind the bluff bodies. A time-resolved particle image velocimetry is used to simultaneously measure the velocity field within these separation bubbles. Based on the dynamics of the mean separation bubbles over and behind the bluff bodies, the flow fields are categorized into three-dimensional, transitional and two-dimensional regimes. The results indicate that the low-frequency flapping motions of the separation bubble on top of the bluff body with $\mathit{AR}=1$ are primarily influenced by the vortex shedding motion, while those with larger aspect ratios are modulated by the large-scale streamwise elongated structures embedded in the oncoming turbulent boundary layer. For $\mathit{AR}=1$ and 20, the flapping motions in the wake region are strongly influenced by those on top of the bluff bodies but with a time delay that is dependent on the $AR$. Moreover, an expansion of the separation bubble on the top surface tends to lead to an expansion and contraction of separation bubbles in the wake of $\mathit{AR}=20$ and 1, respectively. As for the transitional case of $\mathit{AR}=8$, the separation bubbles over and behind the body are in phase over a wide range of time difference. The dynamics of the shear layer in the wake region of the transitional case is remarkably more complex than the limiting two-dimensional and three-dimensional configurations.

An invader in salmonid rearing habitat: current and future distributions of smallmouth bass (Micropterus dolomieu) in the Columbia River Basin

Invasive species and climate change are leading threats to freshwater ecosystems. In the Columbia River Basin (CRB), nonnative fishes are a critical consideration in salmon recovery, yet managers lament a lack of distribution information. Combining a species distribution model (SDM) with environmental DNA (eDNA), we locate range boundary regions of nonnative smallmouth bass (Micropterus dolomieu) and evaluate its overlap with native salmonids. A combination of thermal, hydrological, and geomorphic variables predict that smallmouth bass is distributed across ∼18 000 river kilometres and overlaps with 3%–62% of rearing habitat of salmonids (species-dependent) in the CRB. Under a moderate climate change scenario, smallmouth bass is predicted to expand its range by two-thirds (totaling ∼30 000 river kilometres) by 2080. Basin-wide models were sufficiently accurate to identify upstream invasion extents to within 15 km of the eDNA-based boundary, and including eDNA data improved model performance at critical range boundary regions without sacrificing broadscale model performance. Our study highlights how eDNA approaches can supplement large geospatial data sets to result in more accurate SDM predictions, guiding nonnative species management.

Identification of Summer‐Run Chinook Salmon Spawning Areas in the Main‐Stem Columbia River Upstream of Wells Dam, Washington

AbstractSummer‐run Chinook Salmon Oncorhynchus tshawytscha historically spawned in the Columbia River as far upstream as British Columbia, Canada, but the upstream extent of main‐stem spawning in the contemporary Columbia River has been documented in the tailrace of Wells Dam, Washington. We utilized radiotelemetry and video monitoring equipment between 2011 and 2013 to track wild summer‐run Chinook Salmon upstream of Wells Dam and document any main‐stem spawning sites. Two small spawning areas totaling 3,389 m2 were identified within 1.76 km downstream of Chief Joseph Dam and were used by spawning salmon in each year of the study. Redd deposition totaled 70, 59, and 134 redds in 2011, 2012, and 2013, respectively. When considered with tributary redd counts upstream of Wells Dam from the Methow and Okanogan rivers, main‐stem spawning represented between 1.6% and 2.9% of the annual redd deposition upstream of Wells Dam within each spawning year. Redds were located in depths between 3.7 and 7 m, with spawning occurring between late October and mid‐November in each year. These novel results are the first to detail main‐stem spawning locations of summer‐run Chinook Salmon upstream of Wells Dam and should inform population monitoring metrics, such as annual escapement and prespawn mortality estimates.