Trace Fossil Assemblages Research Articles

Assessing Vertical Bioturbation Intensity from Bedding Planes

Bedding planes hold significant value in ichnological analyses as they improve the recognition of ichnotaxa and yield important insights into the distribution and morphology of trace fossils. However, a significant limitation lies in the challenge of accurately estimating bioturbation intensity from bedding planes. This is in part because current approaches do not establish a connection between bioturbation intensity as seen on bedding planes (plan view) and in cross-section (elevation view). This disconnect complicates the integration of bedding plane assessments into more conventional datasets collected from elevation view. Seventy-seven million Monte Carlo simulations were performed on 11 non-palimpsest Skolithos assemblages, each with varying constraints on burrow length and diameter. These simulations were used to investigate the range and frequency of cross-sectional bioturbation intensities across a spectrum of bedding plane intensities (2-50%). The results show that elevation view bioturbation intensity can be reliably approximated by multiplying plan view bioturbation intensity by the average burrow length relative to the bed thickness, independent of burrow diameter. These findings improve the ichnological analysis of bedding planes for vertical trace fossil assemblages and offer a framework for future research into more complex assemblages.

Full-Body Impressions: A category of trace fossils unique to exceptionally preserved bedding planes and indicators of true substrates

Certain biogenic sedimentary structures can be considered to be reliable indicators of true substrates. Full-body impressions, or FBIs, are trace fossils that are shallow, highly-detailed impressions of the ventral axial anatomy of low body mass animals made while living . Their mode of preservation makes FBIs distinct from undertracks, body moulds, or passive tool marks. FBIs are made at the air- or water-substrate interface during sedimentary stasis. Associated trace fossils, like those that exhibit excavations with preserved waste mounds and very near-surface burrow roofs, located on the same bedding plane, can support the interpretation of true substrate preservation. Integration of the biogenic and sedimentological data from both Carboniferous and Pleistocene sites with well-preserved bedding-plane trace fossil assemblages has confirmed the conditions under which true substrates are preserved. These include periods (as brief as days to weeks) of sedimentary stasis, followed by gentle deposition without erosion, and without the need to invoke biofilms or biomats as consolidants. The potential for these taphonomic conditions exist in both alluvial and lacustrine environments, as well as intertidal-estuarine valleys, as described by other published examples of FBI preservation. Future FBI discoveries will prove useful in identifying true substrates across a range of time periods and environments.

EVALUATING TRACE FOSSILS, FLUVIAL ARCHITECTURE, AND COLONIZATION PATTERNS IN CHANNEL AND OVERBANK DEPOSITS FROM THE MIOCENE VINCHINA FORMATION, WESTERN ARGENTINA

Abstract Trace-fossil distribution within the framework of three-dimensional fluvial architecture has been commonly overlooked. The Miocene Vinchina Formation in western Argentina preserves extensive outcrops of fluvial deposits, including architectural elements of both anastomosing and braided systems identified along the Quebrada de La Troya. Multistorey sandy channels, amalgamated sandy channels, heterolithic multistorey channels, channels with gravel bars, abandoned channels, muddy floodplains, crevasse splays, and crevasse channels have been identified. Of these, only the deposits of three elements were bioturbated, namely crevasse splays, anastomosing abandoned channels, and braided abandoned channels. Vertical simple burrows (Skolithos isp.), large-sized J burrows (Capayanichnus vinchinensis), and simple horizontal burrows (Palaeophycus tubularis) are the most common trace fossils in the Vinchina Formation. Other elements include the horizontal meniscate trace Taenidium barretti and the vertebrate footprints Tacheria troyana, Macrauchenichnus troyana, and Ardeipeda isp. The trace-fossil assemblages identified in the Vinchina Formation collectively illustrate the Scoyenia Ichnofacies. In addition, five ichnofabrics are characterized. The position of the water table, substrate consistency, flow energy, and time between depositional events under arid to semi-arid climate conditions were the main parameters controlling bioturbation. Based on detailed observation of the cross-cutting relationship among ichnotaxa, the ichnofabric distribution and the preservation features of the trace fossils studied, a colonization sequence for each of the subenvironments of the Vinchina Formation is proposed in this study. In addition to integration with conventional facies analysis, articulating ichnologic data and fluvial architecture provides further insights into the application of trace fossils to unravel the sedimentary dynamics of alluvial systems.

Geochemical characterization of trace fossil assemblages in spotted marls and limestones of the Lower Jurassic of the Western Carpathians: environmental implications

AbstractThe trace fossils of the spotted limestones and marls from upper Pliensbachian of the Skladaná Skala section, Western Carpathians (Slovakia), are characteristic of Zoophycos ichnofacies preserved in outer shelf deposits. The ichnoassociation is dominated by Lamellaeichnus and Chondrites, while Palaeophycus, Planolites, Teichichnus, Thalassinoides, Trichichnus, and Zoophycos, are less abundant. The presence of relatively large sized traces (reaching a decimetre in horizontal dimension for Thalassinoides and Zoophycos) and continuously well bioturbated deposits point to well oxygenated shallow zone (mixed layer) and deeper part of stiffed substrate of the transitional layer of bottom substrate up to the first tens of centimetres to depth. A first tier (tier I) corresponds to the shallowest sediment occupied by Bathysiphon tests and other epifaunal body fossils. The tier II is characterized by shallow infaunal burrows of deposit feeders (Planolites) and permanent dwelling, open burrows (Palaeophycus, Thalassinoides) located in a Ca-rich sediment (corresponding to CaCO3). The tier III is dominated by deposit feeders (Lamellaeichnus, Teichichnus) that burrowed stiff Ca-rich sediment below the redox boundary. Trace fossils of tier III are rich in fine siliciclastics (high content in Si, Al, K mainly associated to clays) and pyrite framboids (enrichment in Fe, S and Co). The deposit-feeders of tier III maintain connection to the sediment-water interface. Tier IV (Zoophycos, Teichichnus and large Chondrites), and deepest infaunal forms of the tier V. (Trichichnus, Pilichnus and tiny forms of Chondrites) comprises trace fossils commonly rich in pyrite framboids with relatively high content of Fe, S, and Co, congruent with chemosymbiotic behaviour commonly inferred for Chondrites, Trichichnus, and the organic-matter storage of Zoophycos.

Benthic response to event deposition and environmental disturbance in a shoreface to subaqueous delta system: Ichnology of the Silurian-Devonian Furada Formation of Asturias, Spain

The Furada Formation of Asturias, Spain, represents a clastic shallow-marine unit deposited during the middle Silurian to the Early Devonian. This formation contains abundant evidence of event deposition and non-uniform distribution of bioturbation structures representing a benthic response to multiple stressors. The 170-m-thick succession was measured and described, and the ichnotaxa were recorded and associated with five sedimentary facies. Most shallow-marine environments are characterized by periodic events of environmental disturbance, mainly by episodic deposition, which may be recorded by a change in both the degree of bioturbation and ichnodiversity. The general depositional setting for this formation has been previously interpreted as a wave-dominated and storm-influenced shallow-marine environment. However, sedimentologic features described in this study, such as anomalous heterolithic and mudstone units, representing fluid mud layers, hyperpycnal flow deposits and plume collapse accumulations, suggest the influence of fluvial discharge, making it a more complex depositional setting. The proposed model comprises an inner sandy shoreface belt flanked seawards by a muddy subaqueous delta platform. The trace-fossil assemblages of this formation reflect environmental stress factors introduced by the interplay of storms and fluvial input (e.g., high sedimentation rate, fluctuating hydrodynamic energy, decreased substrate consolidation), resulting in reduced ichnodiversity and low abundance. From an ichnofacies perspective, the shoreface complex is characterized by the Cruziana Ichnofacies, whereas the subaqueous delta platform is represented by the Phycosiphon Ichnofacies. Integration of sedimentologic and ichnologic datasets allows for a refined depositional interpretation of the formation and greater understanding of the environmental diversity of wave-and river-influenced shallow-marine clastic systems, including the responses of the middle Paleozoic shallow-marine benthos to event sedimentation and environmental disturbance. This study is one of the first detailed documentations of the ichnology of subaqueous deltas.

Small topographical variations controlling trace maker community: Combining palaeo- and neoichnological data at the Porcupine Abyssal Plain

Ichnological research has generally assumed that abyssal plains are dominated by quiescent, homogenous environmental conditions. Thus, deep-sea trace fossil assemblage changes have been usually linked to significant spatial and temporal environmental variations. Here, we conducted a comparative ichnological analysis between a small abyssal hill (50 m elevation) and the surrounding abyssal plain; this modest bathymetric variation is known to generate substantial environmental heterogeneity for the benthic fauna community of the Porcupine Abyssal Plain (c. 4850 m depth), Northeast Atlantic. Based on X-ray data from a 5 × 5 core grid emplaced in two box cores, we compared hill and plain bioturbational sedimentary structures, including trace fossil assemblages (e.g., ichnotaxonomy) and biodeformational structures (e.g., mixed-layer depth). We observed that topographically-enhanced near-bottom currents over the hill likely produce significant changes in depositional dynamics and sediment properties (e.g., grain size, organic matter content and degradation), and control specificities of bioturbational sedimentary structures (e.g., trace fossils, mixed layer attributes such as thickness, mottled background, discrete traces). Palaeoichnological data suggested that the abyssal plain had experienced consistent conditions during the last thousands of years while the abyssal hill recorded improving environmental conditions for the trace maker community. Our results highlight the complexity of the deep-sea environment, demonstrating that small changes in bioturbated sedimentary assemblages appear even within the same box core (m-scale), and that substantial changes can occur due to environmental heterogeneity (e.g., subtle topographic variations) at the local scale (km-scale). Considering the vast global extent of abyssal hill terrain, we suggest that their influence on the bioturbational sedimentary record may be significantly under-appreciated and require more attention in palaeoenvironmental reconstructions.

Are aeolian ichnofacies distinctive and useful? An analysis of trace fossil assemblages from Cretaceous aeolian facies in northern Patagonia, Argentina

In this paper, Cretaceous trace fossil assemblages are reported from aeolian facies within four units from Patagonia, Argentina, to evaluate the usefulness of the aeolian ichnofacies concept. From oldest to youngest, studied units comprise the Agrio (Valanginian-Hauterivian), Cerro Barcino (Aptian-Cenomanian), Candeleros (Cenomanian), and Allen (Campanian-Maastrichtian) formations. Twenty one ichnotaxa were recorded, including ten of invertebrate origin (Archaeonassa fossulata, Arenicolites tenuis, Beaconites isp., Digitichnus laminatus, Edaphichnium lumbricatum, cf. Octopodichnus isp., Palaeophycus tubularis, Skolithos linearis, Taenidium barretti, and an unidentified arthropod trackway), seven tetrapod footprints and burrows (Avipeda isp., Brasilichnium isp., cf. Chelonipus isp., Reniformichnus katikatii, two types of indeterminate footprints, and burrow fills) and four rhizolith types. Entradichnus meniscus (formerly used as eponym of an aeolian ichnofacies) is considered a junior synonym of Taenidium barretti. Common ichnotaxa in aeolian and fluvial sequences like Taenidium barretti, Skolithos linearis and Palaeophycus tubularis are facies-crossing ichnotaxa and should not be used to distinguish ichnofacies. Trace fossil assemblages and ichnofabrics from aeolian dune, dry interdune and wet interdune facies are interpreted in terms of the potential producers and their palaeoecological and palaeoenvironmental constraints. This procedure is more informative than the ichnofacies approach in aeolian deposits. Based on a literature review of 32 examples of trace fossil assemblages from Cretaceous aeolian sequences worldwide, we conclude that the most distinctive ichnotaxa occur in aeolian dunes, including Brasilichnium, Farlowichnus, lacertoid footprints, Octopodichnus and Paleohelcura. It is recommended to abandon the usage of the Chelichnus, Entradichnus and Entradichnus-Octopodichnus ichnofacies, which are poorly defined in terms of recurrence and environmental significance or based on uncharacteristic or unavailable ichnotaxa. We propose a new definition of the Octopodichnus ichnofacies, diagnosed by the presence of subequant tetrapod footprints in quadrupedal trackways and/or selected arachnid /insect trackways. This ichnofacies is a taphofacies recorded from the Permian to the Cretaceous, and probably extends to the Cenozoic.

Paleoenvironmental insights from a middle Pennsylvanian trace-fossil assemblage during the late Paleozoic ice age in southern Brazil

Trace fossils play a fundamental role as biotic records in glacial deposits associated with the Late Paleozoic Ice Age, especially in the upper units of the Itararé Group in the Paraná Basin, where they are widely described and studied. The absence of ichnological studies for the lower interval of the Itararé Group, represented by the Campo do Tenente Formation, creates a significant gap in understanding the interaction between organisms and glacial environment during the middle Pennsylvanian period in the Paraná Basin. Addressing this gap provides new insights into adaptive strategies and paleoecological aspects related to extreme climatic events. Through macro and microscopic descriptions, we documented for the first time the occurrence of eight ichnotaxa in a siltstone interval of the Campo do Tenente Formation: Dimorphicnus isp., Diplichnites gouldi, Helminthoidichnites tenuis, Hormosiroidea meandrica, Irichnus saltatorius, Merostomichnites narragansettensis, Monomorphicnus lineatus, and Umfolozia sinuosa. This trace fossil association, dominated by arthropod trackways and burrows of wormlike organisms, indicates the coexistence of the Mermia and Scoyenia ichnofacies. Combined with the sedimentological data, these results reflect marginal continental conditions of shallow and restricted environments, with occasional input of glacial freshwater. These contributions not only provide new insights into the understudied ichnological content of this interval of the Itararé Group but also offer important paleoenvironmental and paleoecological perspectives on the middle Pennsylvanian during the most significant glaciation of the Phanerozoic.

Ordovician-Silurian trace fossils from the Takche Formation, Parahio Valley, Spiti Himalaya, India: biostratigraphic and paleoenvironmental significance

ABSTRACT The Ordovician-Silurian trace fossils from the Takche Formation exposed near the Gechang locality, Parahio Valley, Spiti region, Himalaya are described. Following trace fossils were found in the Takche Formation: Arenicolites isp. Aulichnites parkerensis, Archaeonassa fossulata, Catenichnus isp. Cochlichnus isp. Cruziana furcifera, Cruziana goldfussi, Cruziana semiplicata, Chondrites targoni, Chondrites isp. A, Chondrites isp B, Helminthodoichnites isp. Lockeia siliquaria, Laevicyclus mongraensis, Planolites isp. Phycodes circinatus, Phycodes rarus, Phycodes isp. Treptichnus pedum, Thalassinoides isp. and Zoophycos isp. The lowermost part of the Takche Formation yielded significant traces of Cruziana affinity i.e. Cruziana furcifera, Cruziana goldfussi, and Cruziana semiplicata suggesting Tremadocian-Darriwilian age (Lower to Middle Ordovician). The middle part of the Takche Formation is dominated by Zoophycos isp. Chondrites targoni, Chondrites isp. A, and Chondrites isp. B suggests a deeper water offshore setting. The uppermost part yielded Archaeonassa fossulata, Aulichnites parkerensis, Laevicyclus mongraensis, Planolites isp. Helminthoidichnites isp. Phycodes rarus, Phycodes isp. and scratch marks. Overall, the trace fossil assemblages indicate a shallow marine shoreface to offshore depositional setting.

Trace fossil assemblages from the Lower Devonian (Lochkovian-Emsian) of the Touat area (Reggane Basin, Algerian Sahara) and their palaeoenvironmental significance

Trace fossil assemblages from the Lower Devonian (Lochkovian-Emsian) of the Touat area (Reggane Basin, Algerian Sahara) and their palaeoenvironmental significance

Sedimentary facies analysis of a thin-bed-dominated turbidite succession in the Miocene Temburong Formation, Kampung Bebuloh, Labuan Island, Malaysia

Thin-bedded turbidites are important hydrocarbon-bearing reservoirs in many mature fields throughout the world. The type of turbidite depositional setting strongly influences vertical and horizontal continuity of the reservoir. However, distinguishing between lobe and levee associated thin-bedded turbidites remains challenging due to their generally similar facies characteristics. This study aims to contribute some insights to this problem by conducting a detailed bed-scale facies analysis on the Early Miocene Temburong Formation exposed at Kampung Bebuloh, Labuan Island, Malaysia, which may uncover features that can help in differentiating between both depositional types. Six facies are recognized in the Temburong Formation, which are interpreted as low density turbidites (F1–F4), hybrid event beds (F5), and sustained turbidites (F6). Detailed facies and ichnology analysis reveal features which are consistent with a lobe fringe deposit rather than levee-associated environment interpretation, including tabular bed geometries, presence of hybrid event beds, and the absence of thick-bedded channel-fill sandstones. Five facies associations are identified and interpreted as representing sub-environments within an overall lobe depositional setting. A diverse trace fossil assemblage, comprising the Nereites ichnofacies is consistent with a deep marine environment. Further identification of the Paleodictyon and Nereites sub-ichnofacies indicates a distal turbidite system setting, most likely lobe fringe. Based on the thick accumulation of FA2 and FA3 and no distinct observable vertical trend, the Early Miocene Temburong Formation in Labuan is interpreted as turbidites deposited at the fringes of lobe complexes rather than a single lobe fringe. Earlier works into the Temburong Formation in SW Labuan proposed a middle slope to proximal basin floor setting where the thin-bedded intervals were interpreted as potentially representing either levee deposits or the fringes of confined lobe deposits. Conversely, the Temburong Formation at Kampung Bebuloh exhibits characteristics of an unconfined lobe setting, suggesting an evolutionary change in depositional conditions over time.

Evaluating tracemaker recovery after the Cretaceous–Paleogene (K–Pg) boundary event: different biotic responses at the Caravaca section

AbstractTrace fossils are an aid to characterize major extinction events, enriching evolutionary paleoecology research. Ichnological analysis at the Cretaceous–Paleogene (K–Pg) marine boundary interval, worldwide, points to a relatively minor disruption in the macrobenthic tracemaker community; that is, trace-fossil assemblages do not change significantly across the K–Pg boundary, showing relatively rapid recovery, locally in just a few years post-impact. To evaluate the incidence of this event and the rapid recovery of the tracemaker community afterwards, the evolution of planktic and benthic groups at the K–Pg boundary interval was analysed in the Caravaca section (Betic Cordillera) based on the integration of available data. In general terms, planktic foraminifera and calcareous nannoplankton dropped in abundance and diversity at the K–Pg boundary, whereas benthic foraminifera did not show significant extinction, but rather a sudden and dramatic decrease in diversity and reorganization. After the K–Pg boundary event, planktic communities exhibit a prolonged delay in recovery—mainly occurring above the dark boundary layer—with respect to benthic foraminifera. The K–Pg boundary event did not induce a severe crisis for the burrowing macroinfauna, as revealed by the similarity between pre- and post-event ichnotaxa, showing a comparatively rapid first colonization phase, less than 2 ky after the event. The record of Zoophycos and Chondrites at the base of the dark boundary layer evidences an opportunistic behaviour of tracemakers and a high independence from oxygen in pore waters and food in the sediment, allowing for the colonization of an overall unfavorable environment.

AVIAN FORAGING ON AN INTERTIDAL MUDFLAT SUCCESSION IN THE EOCENE TANJUNG FORMATION, ASEM ASEM BASIN, SOUTH KALIMANTAN, INDONESIAN BORNEO

Abstract Moderately diverse trace fossil assemblages occur in the Eocene Tambak Member of the Tanjung Formation, in the Asem Asem Basin on the southern coast of South Kalimantan. These assemblages are fundamental for establishing depositional models and paleoecological reconstructions for southern Kalimantan during the Eocene and contribute substantially to the otherwise poorly documented fossil record of birds in Island Southeast Asia. Extensive forest cover has precluded previous ichnological analyses in the study area. The traces discussed herein were discovered in newly exposed outcrops in the basal part of the Wahana Baratama coal mine, on the Kalimantan coast of the Java Sea. The Tambak assemblage includes both vertebrate and invertebrate trace fossils. Invertebrate traces observed in this study include Arenicolites, Cylindrichnus, Diplocraterion, Palaeophycus, Planolites, Psilonichnus, Siphonichnus, Skolithos, Thalassinoides, Taenidium, and Trichichnus. Vertebrate-derived trace fossils include nine avian footprint ichnogenera (Aquatilavipes, Archaeornithipus, Ardeipeda, Aviadactyla, cf. Avipeda, cf. Fuscinapeda, cf. Ludicharadripodiscus, and two unnamed forms). A variety of shallow, circular to cylindrical pits and horizontal, singular to paired horizontal grooves preserved in concave epirelief are interpreted as avian feeding and foraging traces. These traces likely represent the activities of small to medium-sized shorebirds and waterbirds like those of living sandpipers, plovers, cranes, egrets, and herons. The pits and grooves are interpreted as foraging traces and occur interspersed with both avian trackways and invertebrate traces. The trace fossils occur preferentially in heterolithic successions with lenticular to flaser bedding, herringbone ripple stratification, and common reactivation surfaces, indicating that the study interval was deposited in a tidally influenced setting. Avian trackways, desiccation cracks, and common rooting indicate that the succession was prone to both subaqueous inundation and periodic subaerial exposure. We infer that the Tambak mixed vertebrate-invertebrate trace fossil association occurred on channel-margin intertidal flats in a tide-influenced estuarine setting. The occurrence of a moderately diverse avian footprint and foraging trace assemblage in the Tambak Member of the Tanjung Formation illustrates that shorebirds and waterbirds have been using wetlands in what is now Kalimantan for their food resources since at least the late Eocene.

The Lower Cambrian deposits of the Le Rozel Formation (Normandy, NW France): Insight into a newly described ichnofossil assemblage

The rise of morphological complexity and taxonomic diversity of trace fossils during the Cambrian Period has been a prominent focus to track the evolution and diversification of early animals. Here, we present a newly described trace fossil assemblage from the lower Cambrian Le Rozel Formation in Normandy (Northwestern France) composed of the following ichnogenera: Archaeonassa, Bergaueria, Helminthoidichnites, Helminthopsis, Nereites, Psammichnites, and Treptichnus. Trace fossils are exquisitely preserved on top of strata (i.e., epirelief), and are distributed through the entire thickness of the siliciclastic succession of about 200 m. This thick sedimentary succession shows sedimentary environments with ripples generated by unidirectional current processes or by oscillatory processes, such as hummocky cross-stratification (HCS) storm facies, tidal facies with flaser stratification, and unidirectional current ripples. In addition, syneresis cracks and microbially induced sedimentary structures (MISS) suggest that the Le Rozel Formation was deposited in a shallow marine shelf environment. New U-Pb dating of detrital zircon grains suggests a late Ediacaran maximum deposition age of 549 ± 3 Ma. This dating is supplemented by the substantial preservation of the three-dimensional burrow systems (Treptichnus pedum) and the large sediment bulldozers (Nereites, Psammichnites). Collectively, our findings suggest that the trace fossils assemblage was deposited during the Terreneuvian Epoch, beneath the first archaeocyaths and the first trilobite-bearing facies from both other lower Cambrian Carteret and Saint-Jean-de-la-Rivière Formation. The ichnofauna described from the Le Rozel Formation complements the worldwide trace fossil record around this critical time of life history.

Quantifying a tide-dominated, wave-, and river-influenced delta in Miocene facies of the Niger Delta basin

ABSTRACT The modern Niger Delta has long been classified as a mixed tide-, wave-, and river-influenced delta. Still, no detailed studies have quantified the relative proportions of formative processes and facies. This work presents the first quantitative estimate of the relative influence of formative processes in Miocene deposits based on core data from the Greater Ughelli, Central Swamp, and Coastal Swamp depositional belts of the Niger Delta basin. Facies analysis of 288.2 m of core from four wells shows approximately 53% tidal facies, 33% fluvial facies, and 14% wave-formed facies, indicating deposition in a prograding tide-dominated, wave-, and river-influenced delta. Cores from the Greater Ughelli and Central Swamp depobelts exhibit coarsening-upwards, prograding deltaic facies successions overlain by fluvial mudstone. Tidal deposits in the Coastal Swamp depobelt show coarsening-upwards prograding deltaic facies successions with well-developed tidal bundles indicating seasonal deposition. The basal part of these facies successions also reveals repeated floods characterized by slump and load structures and dewatering features. Delta-plain and delta-front–prodeltaic facies associations are identified based on physical sedimentary and biological structures. The delta-plain facies association consists of weakly bioturbated mudstone, fissile mudstone, and coarse-pebbly stratified sandstone facies with sparse trace fossils. The delta-front–prodeltaic facies association contains muddy and sandy heteroliths, stratified, medium and coarse-grained, cross-bedded sandstone, and convoluted mudstone facies. The facies vary but characteristically contain trace-fossil assemblages of the recently proposed Rosselia and Phycosiphon Ichnofacies, indicating delta-front and prodeltaic settings, respectively. Variability in the facies is determined by the relative influence of hydrodynamic processes (tide, wave, and river), variations in physicochemical stress, and the episodic character of deposition. In addition, based on the relative influence of each hydrodynamic processes, the facies differ through the successions, sometimes subtly.

Interpreted depositional conditions of balanced-fill lake basin strata incorporating vertebrate and invertebrate trace fossils, Triassic Santa Clara sub-basin, Cuyana rift basin, Argentina

ABSTRACT The Santa Clara Abajo and Santa Clara Arriba formations host a diverse assemblage of trace fossils that record a wide range of behaviors and a broad array of ecological niches during the Middle Triassic—a critical period in the evolution of continental fauna with the diversification of both synapsids (cynodont and dicynodont) and archosauromorphs (dinosaurs, pterosaurs, and crocodilians) that represent post-Permian faunal recovery. The Santa Clara formations are part of the continental infill of the Cuyana rift basin in Argentina and represent a lacustrine system with fluvial input and delta development. Sedimentological characteristics of these units as well as their stacking patterns characterize a “fluctuating profundal” facies association typical of a balanced-fill lake basin. The lacustrine and associated terrestrial environments preserve a rich record of invertebrate traces with 26 ichnogenera from ethological classes of fodichnia, domichnia, repichnia, pascichnia, and cubichnia occupying all continental tiers (subaerial and subaqueous, surficial, and/or very shallow, shallow, mid, and deeper) and ecological niches (epiterraphilic, terraphilic, hygrophilic, and hydrophilic). In association with invertebrate traces, two taphonomic modes of tetrapod footprints have been found: a moderate-fidelity mode and a high-fidelity mode. Physical sedimentary features, burrows, trails, and tracks, and their stratigraphic positions are integrated to interpret the main factors involved in footprint preservation in these subsettings. The most significant and variable preservational factor found is water-table fluctuation controlled by the paleohydrology of a balanced-fill lake system. These data show that in balanced-fill lake systems, diverse trace assemblages occur in the lake and associated subsettings such as delta plains and lake-margin settings, whereas trace fossils can be totally absent in coeval lake-center strata, particularly if anoxic lake-bottom conditions occur, as probably occurred in the meromictic Santa Clara lake system.

Shelf and Lower-Shoreface Deposits in the Upper Midway Group and the Transition into Fluvial-Dominated Deltaic Deposits in the Hooper Formation (Lower Wilcox Group) in the Southeastern Texas Gulf Coast

The Paleocene Upper Midway Group in the southeastern Texas Gulf Coast records a stratigraphic succession of shelf and lower-shoreface systems. Four (4) recognition criteria for these systems include (1) thin (<2 ft [<0.6 m]), erosion-based and lenticular, very fine-grained and fine-grained sandstone beds with swaley cross-stratification, (2) a diverse Cruziana trace-fossil assemblage consisting of Teichichnus, Palaeophycus, Planolites, and Schaubcylindrichnus, (3) net-sandstone trends consisting of strike-elongate ribbons and pods that merge updip (north and northwest) into broad, sheet-like patterns, and (4) an absence of northwestern and northern feeder systems inferred from net-sandstone maps. Net-sandstone trends in Upper Midway shelf deposits in southeastern Texas are irregular and narrow, strike-elongate pods and ribbons ranging in thickness from 20 to 35 ft (6 to 11 m). They grade southward and southeastward into extensive, muddy areas with <10 ft (<3 m) of net sandstone. In contrast, lower-shoreface deposits at the top of the Upper Midway Group contain wide (>10 mi [>16 km]) strike-elongate belts of >40 ft (>12 m) of net sandstone. These belts extend northeastward >50 mi (>80 km) along depositional strike. Sandstone-body continuity, inferred from wireline-log cross-sections, differs greatly among these shelf and lower-shoreface deposits. Sandstone-body continuity in Upper Midway lower-shoreface deposits at 10 mi (16 km) well spacing is approximately 60%, whereas it is only 20% at the same well spacing in shelf deposits. At 20 mi (32 km) well spacing, lower-shoreface sandstone-body continuity is 37% but only 4% in shelf deposits. Microscale heterogeneity in the Upper Midway Group is related to sedimentary structures and accessory features that disrupt vertical sandstone-bed continuity over scales of millimeters to centimeters. These include mudstone drapes, scour surfaces, shell debris, mudstone clasts, and trace fossils. Mesoscale heterogeneity, inferred from cores and wireline-log cross-sections, is defined by lateral sandstone-body pinchouts and vertical stacking patterns. In contrast, macroscale heterogeneities in the Upper Midway Group, inferred from net-sandstone maps, compose facies architecture at a regionally-mappable scale. They include paleogeographic elements such as shelf sand-shoal complexes, strike-fed shoreface systems, and abandoned-deltaic headlands. These paleogeographic elements extend for tens of miles or kilometers along depositional strike from southwest to northeast. The transition in depositional systems in the Upper Midway Group to the Lower Wilcox Group is abrupt, recording a change from wave-dominated shelf and shoreface systems to fluvial-dominated deltaic depositional systems. Dip-elongate, fluvial-dominated deltaic deposits at the base of the Wilcox Group (Hooper 1 depositional unit) have depositional axes oriented orthogonally with respect to those in the underlying Midway Group. Hooper 1 depositional axes are narrow (commonly <4 mi [<6.4 km] across. They are erosionally-based, net upward-fining, and composed of fine- to medium-grained sandstone with abundant siltstone and mudstone rip-up clasts. Distributary-channel trends in the Hooper 1 depositional unit are connected southward and southwestward to downdip-bifurcating trends representing upward-coarsening sub-delta deposits that pinch out into interdistributary mudstones. Microscale heterogeneity in fluvial-dominated deltaic deposits in the lower part of the Hooper Formation occurs at the scale of large (>2 in [>5.1 cm]) diameter siltstone and mudstone clasts in distributary-channel deposits. Mesoscale heterogeneity is controlled by lateral pinchouts of distributary-channel sandstone bodies into floodplain and interdistributary mudstones. Macroscale heterogeneity is represented by numerous dip-elongate distributary-channel depositional axes that segment lower-coastal-plain systems into discrete sandy depocenters. Six Hooper 1 deltaic depocenters occur along a 60 mi (96 km) span along the Angelina-Caldwell Flexure. Sandstone-body continuity along depositional strike (southwest to northeast) in the Hooper 1 depositional unit is poor, being only approximately 25% at 10 mi (16 km) well spacing. These sandstone-body continuity values record the presence of multiple, locally occurring deltaic depocenters within a fluvial-dominated deltaic system, in contrast to the wave-dominated shelf and lower-shoreface deposits in the underlying Upper Midway Group.

Diverse Lophoctenium burrows from the Upper Devonian (Famennian V and VI) of Algeria

Three ichnospecies of Lophoctenium (L. comosum, L. richteri and L. cf. haudimmineri) are reported from the Upper Devonian Argiles de Marhouma Formation (southwestern Algeria) for the first time. This formation contains a diverse assemblage of trace fossils belonging to the Nereites ichnofacies. We describe three ichnospecies of Lophoctenium that correspond to three different feeding strategies among the diverse ichnofauna of the Argiles de Marhouma Formation. The strategy of L. comosum evidences food-rich sediments as it maximises the amount of collected food per foraged distance. The strategy of L. richteri is probably most efficient for feeding in slightly less nutrient rich sediment, but gives the opportunity to discover more prolific feeding sites by moving on longer distances. The strategy with a straight central burrow and probes on both sides of the main burrow with long interspaces (L. cf. haudimmineri) was presumably used in the least food rich sediment. It seems that the Lophoctenium was widespread in the Devonian, but restricted to seas surrounding Gondwana.

Ichnology of the Cenomanian Buda Formation, USA: assessing environmental and taphonomic controls on carbonate trace-fossil assemblages

Ichnology of the Cenomanian Buda Formation, USA: assessing environmental and taphonomic controls on carbonate trace-fossil assemblages

Controls on bioturbation and sediment distribution in carbonate shoreface deposits: insights from heterogeneity in Pleistocene and Recent strata

ABSTRACT Burrowing organisms alter sedimentary textures, influence cement distribution, and affect petrophysical characteristics of carbonate strata. Although many descriptions of carbonate successions reference bioturbation, quantitative data on spatial variability of trace fossils is rare, and fewer studies address trace-fossil influence on postdepositional modification of sedimentary deposits, which can affect petrophysical properties. To address these unknowns and determine the controls on ichnology in carbonate shoreface successions, this study evaluates the along- and across-strike distribution of sediment and bioturbation in recent, Holocene, and Pleistocene shoreface deposits on the leeward margin of Crooked–Acklins Platform (CAP), southern Bahamas. To the north, the extant margin shelf is characterized by poorly to moderately sorted, very fine–fine, skeletal–peloid–ooid sand with an average of 16% mud (< 62.5 µm) that is moderately to intensely bioturbated (ii3–6). Trace assemblages are diverse, and include horizontal tracks and trails, abundant horizontal deposit-feeding and locomotion traces, as well as dwelling and resting burrows attributable to the proximal Cruziana Ichnofacies. In marked contrast, shelf margin deposits to the south are well-sorted, medium ooid–peloid sand with < 1% mud, and display a range of bioturbation, from nonbioturbated to moderately intense bioturbation (ii1–4). Trace-fossil assemblages exhibit low ichnodiversity, dominated by vertical dwelling burrows with reinforced wall lining attributable to the Skolithos Ichnofacies. Holocene and Pleistocene strata show similar proximal-to-distal and along-strike variations in sediment attributes, ichnodiversity, and bioturbation. These trends reflect a progressive, north-to-south increase in energy reflecting the change in margin orientation relative to the direction of dominant wave energy, analogous to the recent system. This study provides data for an integrated sedimentologic–ichnologic conceptual model for processes and patterns of sediment accumulation on carbonate shorefaces, and are distinct from siliciclastic analogs.