Rib Density Research Articles

Experimental investigations on the collapse of spiral-stiffened cylindrical shells under external hydrostatic pressure

This paper presents a comprehensive experimental investigation into the collapse behavior of spiral stiffened cylindrical shells under external hydrostatic pressure. Spiral stiffened cylindrical shells with two different pitches and cylindrical shells were designed and fabricated. The fabricated specimens were divided into three groups. Each group has two identical specimens. Test specimens were welded from stainless steel. The shell was formed by cold bending, and the spiral ribs were welded to the surface of the shell to form a spiral stiffened cylindrical shell. Measurements of initial geometric imperfections and shell thickness were performed on the fabricated specimens to verify the accuracy of fabricating. Subsequently, the specimens were placed in a pressure chamber and pressurized until collapsed to obtain the ultimate collapse pressure and collapse modes. In addition, a uniaxial tensile test was conducted on the shell material to obtain its material properties. Experimental results indicated that the fabricated specimens had excellent repeatability and reasonable precision. The presence of a spiral rib effectively boosted the ultimate collapse pressure of the cylindrical shell with a more pronounced strengthening effect observed as the rib density increased.

Optimization of Wing Structure Based on Fluid-solid Coupling and Vibration Model

With the development of ages, human beings have made more efforts to develop the sky, and aircraft is widely used as the main tool for human beings to set foot in the sky. Therefore, optimizing wing structure is positive for saving resources and improving efficiency. In this paper, by establishing the conventional wing structure model and the wing model after changing its rib density, we compared results based on fluid-solid coupling and vibration mode analysis. It is found that: The wing structure optimization should appropriately increase or decrease the wing-rib aperture (i.e., increase/decrease by 10%) based on ensuring the flight performance of the wing, and it can effectively reduce the overall deformation of the wing and ensure the flight performance of the wing, which provides a valuable reference for the design of the wing structure.

Reappraisal of the latest Albian (Mortoniceras fallax Zone) cephalopod fauna from the classical Salazac locality (Gard, southeastern France)

The Salazac locality (Gard, southeastern France) is renowned for the richness of its cephalopod fauna (especially ammonites) from the Mortoniceras fallax Zone (uppermost Albian, Lower Cretaceous). However, most ammonite species have paradoxically been scarcely illustrated up to now. Furthermore, the rare assessments of ammonite taxonomic diversity are presumably inaccurate, as they either result from a drastic typological approach or could not benefit from the most recent advances in modern concepts of ammonoid intraspecific and ontogenetic variations. In this work, we document an original cephalopod assemblage from Salazac represented by abundant, well-preserved specimens (564 studied ammonites, including ca. 50% of heteromorphs) constituting a high diversity. The sample size and quality of this new material allowed some extensive morphological and biometric analyses whenever possible (i.e., for subdatasets in which a significant number of measurements is available) and to thoroughly assess intraspecific and ontogenetic variations for a few species. This led to determine the diagnostic value of various shell characters commonly used to characterize both well-documented and poorly-defined species. In particular, we show that a significant part of the variation in the number of ribs per whorl within the heteromorphic genera Mariella and Turrilitoides is directly related to shell size and that the strength of tubercles covaries negatively with the number of ribs in Mariella, therefore challenging the diagnostic value of these characters. We further suggest that rib density covaries with whorl shape in Mariella, following Buckman’s first rule of covariation. In total, one species of nautilid and 38 taxa (25 genera) of ammonites are described, among which five species are documented from Salazac for the first time. In turn, this study provides valuable data for future phylogenetic, biostratigraphical, biogeographical and broad-scale diversity studies.

Testing the New Parameters affecting The Outcome of Extracorporeal Shockwave Lithotripsy for Upper Ureteric Stones

Background: Extracorporeal Shock wave lithotripsy (ESWL) is widely used in treating patients with ureteralstones because it is effective, safe, and noninvasive. Based on factors such as size and the location of stones,there is a significant variation in the overall stone-free rate (SFR).Aim of the study: To evaluate the effect of ureteral wall thickness (UWT), stone attenuation, the time fromfirst attack of pain till first session of ESWL and stone/ rib density on the outcome of SWL in the treatmentof upper ureteral stones (UUS).Patient and methods: A prospective study when 127 patients with radio-opaque UUS ranging from 7 to 20mm and treated by ESWL were included in this study. The effect of (stone/ 12th rib) density by KUB, ureteralwall thickness by NCCT and the time from first attack of pain till first ESWL session was studied.Results: The overall successful fragmentation was 75.5%, with the overall success rates in the low density(LD) and high density (HD) groups were 83.8% and 52.94%, respectively. The average number of SWLsessions needed in the two groups for success was 1.9 compared with 2.7 sessions (p<0.05). For stones <10 mm; those with ureteral wall thickness <3.25 mm have success rate about 90.3% VS 69.7% with uretericwall thickness > 3.25 mm which is highly significant. Early ESWL within the first 24 hours of acute attackof first pain has successful fragmentation of 85.45%. With significant effect on number of ESWL sessions.The stone free rate reaches 91.1% for stones <10 mm.Conclusions: The stone free rate is inversely affected by stone /12th rib density ; ureteral wall thickness andthe time from first attack of pain till first session of ESWL, were important predictors of successful ESWL.

AMMONITES OF THE SUBFAMILY ZAPALIINAE FROM THE LOWER TITHONIAN OF ESTANCIA MARÍA JUANA, VACA MUERTA FORMATION (PORTADA COVUNCO MEMBER), NEUQUÉN BASIN, ARGENTINA

A new collection of ammonites from the Picunleufuense Zone (standard base of the Andean Tithonian) of the locality Estancia Maria Juana (southern Neuquen Basin, Argentina) has provided abundant and well preserved material of new transients of Indansites picunleufuense and Choicensisphinctes platyconus. These transients are described and a detailed review of the early evolution of the Indansites lineage is presented. The meaning of changes in rib density, variations within the microconchs (males) and the length of the bodychamber in Indansites picunleufuense are discussed. It is concluded that the changes in rib density which characterize the successive macroconchs of the transients indicate changes in the rate of shell-growth. The more densely ribbed phragmocone of later transients would have had lower growth rates, which would also explain their smaller adult size. The microconchs show high variation in adult size suggesting high plasticity in the size-age maturation, most likely caused by influence of seasonal environmental conditions. A large small-macroconch-like lapetted microconch, coming from the close locality Picun Leufu, is interpreted as a new case of sex-change.

Research on the thermal performance of matrix cooling channel with response surface methodology

In this investigation, three structural parameters are defined for matrix cooling channel: rib angle, rib-to-subchannel ratio and rib density, in a certain typical rectangular cooling space. A carefully designed wind tunnel experiment using transient liquid crystal technique is carried out to obtain surface Nusselt number distribution and pressure drop. The measured data is used to validate the computational method. Turbulence SST model is adopted to study the performance of 13 different matrix channels in Reynolds number range of 5000–90,000. A response surface method is used to fit the CFD solutions and polynomial expressions are established to calculate friction factor ratio (f/f0) and heat transfer enhancement (Nu/Nu0). Variation of thermal performance factor (TPF=(Nu/Nu0)/(f/f0)1/3) versus rib angle, rib-to subchannel ratio and rib density is plotted. The result shows that different structural parameters have to be considered simultaneously in order to improve matrix thermal performance. The comparison of different cases indicates that although smaller rib angle or narrower subchannel can produce very strong local heat transfer, the uniformity is sacrificed.

New record of Diplomoceras cylindraceum (Defrance, 1816) from the Maastrichtian of the Western Fore-Balkan (Bulgaria)

New data on the heteromorph ammonite species Diplomoceras cylindraceum (Defrance, 1816) have been obtained from three localities of the Western Fore-Balkan (West Bulgaria). A retrospection of the earlier Bulgarian records is also presented. This characteristic species is relatively common in Bulgaria and, based on both previously and the newly collected data, it is confined to the Maastrichtian. Owing to the difficulty in collecting enough specimens, the amount of variation in ribbing and whorl-section shape, which have been used by other authors for determination of more species within Diplomoceras, is difficult to judge. Nevertheless, the Bulgarian material reveals that rib density varies in broad limits and the rib index does not show a sufficient degree of variance to warrant the distinction of more than one species. Therefore, it is doubtful that all specific names that have been proposed in the literature for rib differences represent other species than Diplomoceras cylindraceum.

Shell and Genital Morphology Fails to SeparateAncotrema hybridum(Ancey, 1888) andA. sportella(Gould, 1846) (Gastropoda: Haplotrematidae)

ABSTRACT The obsolescence of radial sculpture on the last whorl of the land snail Ancotrema hybridum (Ancey, 1888) has been used to separate it from A. sportella (Gould, 1846), but shells are sometimes difficult to assign to one species or the other, prompting questions about their species status. Factor analysis of shell characters of 311 specimens of Ancotrema Baker, 1931, including characters of surface sculpture strength, shell size and rib density, revealed continuous variation of these characters without bimodality. Radial sculpture begins obsolescing at about whorl five, so shells that stopped growing near whorl five have A. sportella-like morphology, while those that continued growing after whorl five have A. hybridum-like morphology. We saw no meaningful differences in the genital anatomy that would suggest different species. The obsolescence of sculpture beginning about whorl five regardless of ultimate whorl number, lack of bimodality of shell characters, and lack of anatomical differences lead...

Association between shell morphology of micro-land snails (genusPlectostoma) and their predator’s predatory behaviour

Predator–prey interactions are among the main ecological interactions that shape the diversity of biological form. In many cases, the evolution of the mollusc shell form is presumably driven by predation. However, the adaptive significance of several uncommon, yet striking, shell traits of land snails are still poorly known. These include the distorted coiled “tuba” and the protruded radial ribs that can be found in micro-landsnails of the genus Plectostoma. Here, we experimentally tested whether these shell traits may act as defensive adaptations against predators. We characterised and quantified the possible anti-predation behaviour and shell traits of Plectostoma snails both in terms of their properties and efficiencies in defending against the Atopos slug predatory strategies, namely, shell-apertural entry and shell-drilling. The results showed that Atopos slugs would first attack the snail by shell-apertural entry, and, should this fail, shift to the energetically more costly shell-drilling strategy. We found that the shell tuba of Plectostoma snails is an effective defensive trait against shell-apertural entry attack. None of the snail traits, such as resting behaviour, shell thickness, shell tuba shape, shell rib density and intensity can fully protect the snail from the slug’s shell-drilling attack. However, these traits could increase the predation costs to the slug. Further analysis on the shell traits revealed that the lack of effectiveness in these anti-predation shell traits may be caused by a functional trade-off between shell traits under selection of two different predatory strategies.

Sexual dimorphism in shells of Cochlostoma septemspirale (Caenogastropoda, Cyclophoroidea, Diplommatinidae, Cochlostomatinae)

Sexual dimorphisms in shell-bearing snails expressed by characteristic traits of their respective shells would offer the possibility for a lot of studies about gender distribution in populations, species, etc. In this study, the seven main shell characters of the snail Cochlostoma septemspirale were measured in both sexes: (1) height and (2) width of the shell, (3) height and (4) width of the aperture, (5) width of the last whorl, (6) rib density on the last whorl, and (7) intensity of the reddish or brown pigments forming three bands over the shell. The variation of size and shape was explored with statistical methods adapted to principal components analysis (PCA) and linear discriminant analysis (LDA). In particular, we applied some multivariate morphometric tools for the analysis of ratios that have been developed only recently, that is, the PCA ratio spectrum, allometry ratio spectrum, and LDA ratio extractor. The overall separation of the two sexes was tested with LDA cross validation.The results show that there is a sexual dimorphism in the size and shape of shells. Females are more slender than males and are characterised by larger size, a slightly reduced aperture height but larger shell height and whorl width. Therefore they have a considerable larger shell volume (about one fifth) in the part above the aperture. Furthermore, the last whorl of females is slightly less strongly pigmented and mean rib density slightly higher. All characters overlap quite considerably between sexes. However, by using cross validation based on the 5 continuous shell characters more than 90% of the shells can be correctly assigned to each sex.

Pseudoskirroceras, a remarkable but poorly known Early Pliensbachian Tethyan ammonite genus: New data from the High Atlas (Morocco)

The discovery of new Early Pliensbachian ammonite faunas in Central High Atlas (Morocco) allows the re-examination of the taxonomic, stratigraphical and palaeogeographical framework of the genus Pseudoskirroceras, an intriguing but until now poorly known Tethyan taxon. For the first time, several specimens of Pseudoskirroceras mastodon (Fucini, 1935) the type species of the genus, have been collected in a well-known stratigraphical context. This material allows the evaluation of intra-specific variability and consequently the assessment of the taxonomical relevance of various geometrical and ornamental features. The best diagnostic features are the clearly evolute sub-serpenticone shell, associated with the very distinctive low location of the latero-ventral tubercles. In contrast, features such as rib density or the extent of the juvenile Coeloceras-like stage vary greatly with no taxonomic implications. These observations considerably reduce the scope of Pseudoskirroceras, which is in fact a rare taxon, probably localised in the western Mediterranean Tethys, and not a pantropical genus. Thus, the presence of Pseudoskirroceras in North and/or South America is unconvincing. These results confirm that Mediterranean (West Tethyan) ammonite palaeobiodiversity includes more rare and localised species than can be found in NW Europe.

Conchometrics, systematics and distribution of Melanopsis (Mollusca: Gastropoda) in the Levant

In this conchometric study, the systematics and distribution of the freshwater gastropod Melanopsis in the Levant are described. Of the ten species found, three are widespread, two have narrow distributions and five are known only from their type locality. Five smooth-shelled species are recognized (buccinoidea, ammonis, dircaena, khabourensis and meiostoma). Within M. buccinoidea, Jordan Valley populations may belong to a separate subspecies. M. ammonis is clearly differentiated from buccinoidea of the nearby Jordan Valley, but less so from buccinoidea of more distant sites; differences between ammonis and meiostoma are significant but not diagnostic. Five species have ribbed shells. Within costata, four subspecies are recognized (in the northern Orontes, upper Jordan, Sea of Galilee, and in southernmost parts of the Levant). Throughout the Levant, M. buccinoidea frequently hybridizes with M. costata and fossil evidence suggests that these species have been hybridizing for the last 1.5 Myr. M. saulcyi differs from M. costata in its narrower shell and shorter, bumpier ribs and frequently hybridizes with M. buccinoidea throughout the Levant. Specimens from Homs differ from those of the Jordan Valley in their higher figurativity index, fewer ribs and lower rib density. M. germaini differs from M. costata in its more numerous ribs, M. pachya in its shorter ribs, and M. infracincta in its bumpy shell in which each rib has huge tubercles, with a pronounced ridge flanking the columella. Our conclusion that there are ten species in the Levant differs from previous studies that suggested only two subspecies of one species (or superspecies). This difference could stem from (1) our use of nonstandard as well as standard conchometrics, (2) a reappraisal of the importance of the shell vs. the radula in intrageneric systematics, and (3) differences of opinion on the subspecies concept. © 2005 The Linnean Society of London, Zoological Journal of the Linnean Society, 2005, 144, 229−260.

Ontogeny and covariation in the Toarcian genus Osperleioceras (Ammonoidea)

Abstract Starting from embryonic (protoconch-ammonitella) and early juvenile shells, which are indistinguishable at the species level, growth curves of Osperleioceras from the Reynesi Subzone (Upper Toarcian) of the Causses Basin (Aveyron, France) show a continuous radiating range of correlated variation in dimensional and ornamental characters, such as involution, whorl compression, rib strength and rib density. This covariation pattern can be observed among single-horizon assemblages, as well as during individual ontogenetic development. The existence of a continuous intergradational series of shells, ranging from stout coarsely ribbed to smooth suboxycone morphologies, rules out functional or ecological selectivity to explain this non-random variability pattern. The complex interdependence of shape and sculpture can be simulated by a model in which sculpture intensity depends on mantle curvature [Guex, 1999]. The expression of covariation in subadult specimens since the base of Upper Toarcian reveals a rise in variability, concomitant with a size decrease, both contemporaneous with environmental instability. It developed in successive bursts from a fairly long low variability period spanning the whole Middle Toarcian.

Correlations between scale structure and pigmentation in butterfly wings.

We examined the correlation between color and structure of wing scales in the nymphalid butterflies Bicyclus anynana and Heliconius melpomene. All scales in B. anynana are rather similar in comparison to the clear structural differences of differently pigmented scales in H. melpomene. Where scale structural differences in H. melpomene are qualitative, they seem to be quantitative in B. anynana. There is a "gradient" in the density of some structural elements, the cross ribs, in the scales of B. anynana: black, gold, and brown scales show progressively lower cross rib density within an individual. There is, however, high individual variation in the absolute cross rib densities (i.e., scales with a particular color and cross rib density in one individual may have a different color but similar density in another individual). By ectopically inducing color pattern during early pupal development, we examined whether a scale's color and its microstructure could be uncoupled. The effect of these manipulations appears to be different in B. anynana and H. melpomene. In Bicyclus, "black" scales induced by wing damage at an ectopic location normally containing brown scales acquire both an intermediate structure and color between that of brown and normal black scales. In Heliconius, however, intermediate colors or scale structure were never observed, and scales with an altered color (due to damage) always have the same structure as normal scales with that color. The results are discussed on the basis of gene expression patterns, variability in rates of scale development and pigment, and scale sclerotization pathways.

The pattern of geographical and altitudinal variation in the land snail Albinaria idaea from Crete (Gastropoda: Clausiliidae)

Certain morphological characters in the Mediterranean land snail genus Albinaria arc believed not to be closely related to ecological conditions. Recently this view has been challenged by a study on A. idaea from central Crete (Greece) claiming that shell characters such as rib density, apcrtural lamella sizes and shell size parameters depend on ecological variables related to altitude. The present study suggests that variation in apertural lamellae, shell size parameters and shell shape were not related to altitude. Neither were they closely related to expected temperature gradients, nor to Drilits beetle predation pressure, nor to vegetational cover (forests/open land). The rib density was the only factor responding to altitude, but after accounting for altitude, altitude-independent regional variation was also of importance. The correlation with altitude was possibly caused by a response to temperature gradients.

The pattern of geographical and altitudinal variation in the land snail Albinaria idaea from Crete (Gastropoda: Clausiliidae)

Certain morphological characters in the Mediterranean land snail genus Albinaria are believed not to be closely related to ecological conditions. Recently this view has been challenged by a study on A. idaea from central Crete (Greece) claiming that shell characters such as rib density, apertural lamella sizes and shell size parameters depend on ecological variables related to altitude. The present study suggests that variation in apertural lamellae, shell size parameters and shell shape were not related to altitude. Neither were they closely related to expected temperature gradients, nor toDrilus beetle predation pressure, nor to vegetational cover (forests/open land). The rib density was the only factor responding to altitude, but after accounting for altitude, altitude-independent regional variation was also of importance. The correlation with altitude was possibly caused by a response to temperature gradients.

Large heteromorph ammonites from the Upper Cretaceous of Seymour Island, Antarctica

Two species of the heteromorph ammonite genus Diplomoceras Hyatt are described from the Upper Cretaceous of the James Ross–Seymour Islands area, Antarctica. The late Campanian–early Maastrichtian D. lambi Spath has a relatively high rib density, whereas D. maximum n. sp. has a lower rib density and is known only from the uppermost Maastrichtian of the Lopez de Bertodano Formation on Seymour Island. Both species attain an exceptionally large size with the body chamber of D. maximum n. sp. attaining a length in excess of one meter. The structure of the shell wall in Diplomoceras is characterized by the thickening of the nacreous layer below the ribs. The shell thickening results in an inner flat surface and a smooth phragmocone. Reconstruction of the shell suggests at least four parallel shafts and three U-connectives. Estimates of the total density, center of buoyancy, and center of mass in this reconstructed shell indicate a slightly positive buoyant shell with a relatively unstable floating position.