Asymmetry In Response Research Articles

Global sounding of F region irregularities by COSMIC during a geomagnetic storm

Abstract. We analyse reprocessed electron density profiles and total electron content (TEC) profiles of the ionosphere in September 2008 (around solar minimum) and September 2013 (around solar maximum) obtained by the Constellation Observing System for Meteorology, Ionosphere, and Climate (COSMIC/FORMOSAT-3). The TEC profiles describe the total electron content along the ray path from the GPS satellite to the low Earth orbit as function of the tangent point of the ray. Some of the profiles in the magnetic polar regions show small-scale fluctuations on spatial scales <50 km. Possibly the trajectory of the tangent point intersects spatial electron density irregularities in the magnetic polar region. For derivation of the morphology of the electron density and TEC fluctuations, a 50 km high-pass filter is applied in the s domain, where s is the distance between a reference point (bottom tangent point) and the tangent point. For each profile, the mean of the fluctuations is calculated for tangent point altitudes between 400 and 500 km. At first glance, the global maps of ΔNe and ΔTEC are quite similar. However, ΔTEC might be more reliable since it is based on fewer retrieval assumptions. We find a significant difference if the arithmetic mean or the median is applied to the global map of September 2013. In agreement with literature, ΔTEC is enhanced during the post-sunset rise of the equatorial ionosphere in September 2013, which is associated with spread F and equatorial plasma bubbles. The global map of ΔTEC at solar maximum (September 2013) has stronger fluctuations than those at solar minimum (September 2008). We obtained new results when we compare the global maps of the quiet phase and the storm phase of the geomagnetic storm of 15 July 2012. It is evident that the TEC fluctuations are increased and extended over the southern magnetic polar region at the day of the geomagnetic storm. The north–south asymmetry of the storm response is more pronounced in the upper ionosphere (ray tangent points h = 400–500 km) than in the lower ionosphere (ray tangent points h = 200–300 km).

THE EFFECT OF LOSS-OFFSET PROVISIONS ON THE ASYMMETRIC BEHAVIOR OF CORPORATE TAX REVENUES IN THE BUSINESS CYCLE

This paper estimates the response of corporate tax revenues to the business cycle fluctuations and calculates a heterogeneous asymmetry in the tax revenue responses between booms and recessions using a new index of loss-offset generosity provisions. I find strong short-run contemporaneous impact of business cycles on corporate tax revenues and a persistent asymmetric response of corporate tax revenues to booms and recession. Loss-offset generosity provisions enhance this asymmetry. Countries with more generous loss-offset provisions experience much more volatile response of corporate tax revenues to business cycles during recessions, magnifying the asymmetry of the cyclicality.

Dynamic leaf energy balance: deriving stomatal conductance from thermal imaging in a dynamic environment.

In spite of the significant progress made in recent years, the use of thermography to derive biologically relevant traits remains a challenge under fluctuating conditions. The aim of this study was to rethink the current method to process thermograms and derive temporal responses of stomatal conductance (gsw) using dynamic energy balance equations. Time-series thermograms provided the basis for a spatial and temporal characterization of gsw responses in wheat (Triticum aestivum). A leaf replica with a known conductance was used to validate the approach and to test the ability of our model to be used with any material and under any environmental conditions. The results highlighted the importance of the co-ordinated stomatal responses that run parallel to the leaf blade despite their patchy distribution. The diversity and asymmetry of the temporal response of gsw observed after a step increase and step decrease in light intensity can be interpreted as a strategy to maximize photosynthesis per unit of water loss and avoid heat stress in response to light flecks in a natural environment. This study removes a major bottleneck for plant phenotyping platforms and will pave the way to further developments in our understanding of stomatal behaviour.

Buzzing Sympathetic Nerves: A New Test to Enhance Anisocoria in Horner's Syndrome.

Introduction: Patients with suspected Horner's syndrome having equivocal pupil dilation lag and pharmacologic testing may undergo unnecessary MR imaging and work up in the case of false positive pupil test results. Our goal was to increase the diagnostic accuracy of pupillometry by accentuating the inter-ocular asymmetry of sympathetic innervation to the iris dilator with surface electrical stimulation of the median nerve using a standard electromyography machine. We hypothesized that an accentuated difference in sympathetic response between the two eyes would facilitate the diagnosis of Horner's syndrome.Methods: Eighteen patients with pharmacologically proven Horner's syndrome were compared to ten healthy volunteers tested before and after monocular instillation of 0.2% brimonidine tartrate ophthalmic solution to induce pharmacological Horner's syndrome. Pupillary responses were measured with binocular pupillometry in response to sympathetic activation by electrical stimulation of the median nerve in darkness and at various times after extinction of a light stimulus. Sudomotor sympathetic responses from the palm of the stimulated arm were recorded simultaneously.Results: In subjects with Horner's syndrome and pharmacologically induced unilateral sympathetic deficit, electrical stimulation in combination with the extinction of light greatly enhanced the anisocoria during the evoked pupil dilation, while there was no significant increase in anisocoria in healthy subjects. The asymmetry of the sympathetic response was greatest when the electrical stimulus was given 2 s after termination of the light or under constant low light conditions. When given 2 s after termination of light, the electrical stimulation increased the mean anisocoria from 1.0 to 1.2 mm in Horner's syndrome (p = 0.01) compared to 0.22–0.26 mm in healthy subjects (p = 0.1). In all subjects, the maximal anisocoria induced by the electrical stimulation appeared within a 2 s interval after the stimulus. Correspondingly, the largest change in anisocoria between light and dark without electrical stimulation was seen between 3 and 4 s after light-off. While stronger triple stimulation further enhanced the anisocoria, it was less well tolerated.Conclusions: Electrical stimulation 2 s after light-off greatly enhances the sensitivity of pupillometry for diagnosing Horner's syndrome. This new method may help to rule in or rule out a questionable Horner's syndrome, especially if the results of topical pharmacological testing are inconclusive.

Role of Olfactorily Responsive Neurons in the Right Dorsal Habenula–Ventral Interpeduncular Nucleus Pathway in Food-Seeking Behaviors of Larval Zebrafish

The habenula (Hb) plays important roles in emotion-related behaviors. Besides receiving inputs from the limbic system and basal ganglia, Hb also gets inputs from multiple sensory modalities. Sensory responses of Hb neurons in zebrafish are asymmetrical: the left dorsal Hb and right dorsal Hb (dHb) preferentially respond to visual and olfactory stimuli, respectively, implying different functions of the left and right dHb. While visual responses of the left dHb (L-dHb) have been implicated in light-preference behavior, the significance of olfactory responses of the right dHb (R-dHb) remains under-examined. It was reported that the R-dHb can gate innate attraction to a bile salt. However, considering a broad range of odors that R-dHb respond to, it is of interest to examine the role of R-dHb in other olfactory behaviors, especially food seeking, which is essential for animals' survival. Here, using in vivo whole-cell recording and calcium imaging, we first characterized food extract-evoked responses of Hb neurons. Responsive neurons preferentially locate in the R- but not L-dHb and exhibit either ON- (~87%) or OFF-type responses (~13%). Interestingly, this right-to-left asymmetry of olfactory responses converts into a ventral-to-dorsal pattern in the interpeduncular nucleus (IPN), a main downstream target of Hb. Combining behavior assay, we further found that genetic dysfunction or lesion of the R-dHb and its corresponding downstream ventral IPN (V-IPN) impair the food seeking-associated increase of swimming activity. Thus, our study indicates that the asymmetrical olfactory response in the R-dHb to V-IPN pathway plays an important role in food-seeking behavior of zebrafish larvae.

Investigation on EMG Profiles of the Superior Laryngeal Nerve in a In Vivo Porcine Model

Background: The electromyographic (EMG) profiles of external branch of the superior laryngeal nerve (EBSLN) have been defined and the optimal intensity of the stimulation of EBSLN in an in vivo porcine model has been explored. Materials: EMG was simultaneously registered by the surface of endotracheal tube and needle electrodes by applying a monopolar stimulation probe in 12 piglets (22 EBSLNs). Vagal nerve (VN), RLN and EBSLN were excited to record the EMG tracings and cricothyroid muscle twitch (CTM). VN, RLN and EBSLN were stimulated from 0.1 to 1.0 mA. Cmin and Cmax have been defined as the minimum and maximal stimulation to evoke an EMG response. Results: The stimulation resulted in a dose-response curve. Cmin were 0.19 mA (0.04–0.4), 0.19 mA (0.08–0.3) and 0.21 mA (0.1-0.4) for EBSLN, RLN and VN (p > 0.05) respectively. Cmax were 0.6 mA along with an amplitude value of 396 ± 330 μV, 0.5 mA including 1058 ± 382 μV, 0.8 mA coupled with 870 ± 382 μV, equally for EBSLN, RLN and VN (p > 0.05) respectively. No asymmetry of amplitude responses each side for EBSLN, RLN and VN (p = 0.317, p = 0.203 and p = 0.468, respectively) was noted. The amplitudes of EBSLN were significantly lower than RLNs and VN (42% of RLN and 50% of VN amplitude rates). Also, CTM twitch was always detectable with the stimulation of EBSLN. Conclusions: Cmin and Cmax of EBSLN were comparable to RLN and VN standards. The amplitude stimulus-response curves of RLN, VN and EBSLN were highly variable. It has been suggested to apply a stimulation of 1.0 mA and a visual appreciation of CTM twitch for the identification of EBSLN.

Climatology of the Quasi‐6‐Day Wave in the Mesopause Region and Its Modulations on Total Electron Content During 2003–2017

AbstractThe westward quasi‐6‐day planetary wave (Q6DW) with zonal wave number 1 is a prominent oscillation in the mesosphere and lower thermosphere region, which causes significant variabilities in the ionosphere through E region wind dynamo. In this paper, the neutral wind observations from the Thermosphere Ionosphere and Mesosphere Electric Dynamics Doppler Imager experiment, the kinetic temperature observations from the Sounding of the Atmosphere using Broadband Emission Radiometry instrument, and the total electron content (TEC) maps from globally distributed Global Positioning System (GPS) receivers during 2003–2017 are utilized to statistically study the Q6DW in both the neutral atmosphere and the ionosphere. Our statistical results show that (1) the Q6DW is most likely to occur during May and September, exhibiting a quasi‐semiannual oscillation. (2) The periods of the Q6DW show two statistical crests at ~6 and ~7 days separately. (3) The ionospheric oscillations due to the Q6DW are positively related to both the 10.7‐cm solar flux index and the neutral wave amplitudes in the mesosphere and lower thermosphere region. The stronger oscillations in TEC under higher solar activity conditions are due to the larger ionization rate and thus larger background TEC. A stronger Q6DW results in stronger wind dynamos in the E region, which leads to stronger F region responses indirectly. (4) The interhemispheric asymmetries of the ionospheric responses to Q6DW show weak negative correlations with both solar activities and neutral wave perturbations, the mechanism of which needs our further investigation.

Investigating the asymmetry in the EDL response of C60/graphene supercapacitors.

Development of efficient electrodes is one of the main ways to increase the performance of an electrochemical energy storage device. It is known that such performance is associated with the electrode specific area, which allows a much larger interfacial interaction with the electrolyte. In this work, molecular dynamics is employed to model C60/graphene composite electrodes that can expand the effective area by approximately 70% relative to a pure graphene electrode. Our simulations indicate that the performance of supercapacitors of C60/graphene electrodes is superior to those made of planar graphene, in some cases up to 150%. The inherent electrolyte asymmetry in the investigated supercapacitors has a negative effect on the total capacitance, indicating that even better results could be obtained after rational design of the fullerene density on the surface of the graphene as well as the choice of the ions in the liquid ionic composition.

A Dynamical Bifurcation Model of Bipolar Disorder Based on Learned Expectation and Asymmetry in Mood Sensitivity.

Bipolar disorder is a common psychiatric dysfunction characterized by recurring episodes of mania and depression. Despite its prevalence, the causes and mechanisms of bipolar disorder remain largely unknown. Recently, theories focusing on the interaction between emotion and behavior, including those based on dysregulation of the so-called behavioral approach system (BAS), have gained popularity. Mathematical models built on this principle predict bistability in mood and do not invoke intrinsic biological rhythms that may arise from interactions between mood and expectation. Here we develop and analyze a model with clinically meaningful and modifiable parameters that incorporates the interaction between mood and expectation. Our nonlinear model exhibits a transition to limit cycle behavior when a mood-sensitivity parameter exceeds a threshold value, signaling a transition to a bipolar state. The model also predicts that asymmetry in response to positive and negative events can induce unipolar depression/mania, consistent with clinical observations. We analyze the model with asymmetric mood sensitivities and show that large unidirectional mood sensitivity can lead to bipolar disorder. Finally, we show how observed effects of lithium- and antidepressant-induced mania can be explained within the framework of our proposed model.

Domain-wall-assisted asymmetric magnetoimpedance in ferromagnetic nanostripes

With micromagnetic simulations, we study the dynamical magnetic response of a ferromagnetic nanostripe deposited on an AC-current-carrying nonmagnetic layer (that induces an AC Oersted field inside the nanostripe), in the presence of a static axial field. Included into a circuit, the structure induces the magnetoimpedance (MI) whose value is proportional to the time derivative of the transverse magnetization of the nanostripe, hence, it is directly related to the dynamical magnetic response of the stripe. Paying especial attention to the regime of low values of the axial field, we study in detail the mechanism of the dynamical response, thus, MI which is based on the oscillations of the positions of 90° (head-to-tail) domain walls (DWs). Such DWs are present in nanostripes of uniaxial-transverse or four-fold (in-plane) magnetic anisotropies of a small (up to 10 nm) thickness, whose combination of the structural and shape anisotropies stabilizes the structure of domains of diagonal (to stripe axes) magnetization, with a non-zero axial component of the overall magnetization of the nanostripe. For the nanostripes of specific materials (uniaxial Co, cubic Fe3O4), that ordering results in a significant asymmetry of the DW-based dynamical response with respect to the reversal of the axial field. For the nanostripes of other materials (cubic: Fe, Fe3Pt), the DW-based magnetic response is almost insensitive to the axial field, thus, the low-field MI is inefficient. We explain different dynamical behaviors of the nanostripes within the analytical (Walker-like) model of the DW motion combined with a phenomenological model of the DW-based MI, including the DW interactions. The maximum relative shift of the DW-based impedance of the nanostripes ranges from about 50% for Fe3O4 to above 200% for Co nanostripes, whereas the (low-)field regime of the DW-based response (the anisotropic MI regime) is as wide as 30 kA m−1.

Chirality-Induced Electron Spin Polarization and Enantiospecific Response in Solid-State Cross-Polarization Nuclear Magnetic Resonance.

NMR-based techniques are supposed to be incapable of distinguishing pure crystalline chemical enantiomers. However, through systematic studies of cross-polarization magic angle spinning (CP-MAS) NMR in a series of amino acids, we have found a rather unexpected behavior in the intensity pattern of optical isomers in hydrogen/nitrogen nuclear polarization transfer that would allow the use of CP NMR as a nondestructive enantioselective detection technique. In all molecules considered, the d isomer yields higher intensity than the l form, while the chemical shift for all nuclei involved remains unchanged. We attribute this striking result to the onset of electron spin polarization, accompanying bond charge polarization through a chiral center, a secondary mechanism for polarization transfer that is triggered only in the CP experimental setup. Electron spin polarization is due to the chiral-induced spin selectivity effect (CISS), which creates an enantioselective response, analogous to the one involved in molecular recognition and enantiospecific separation with achiral magnetic substrates. This polarization influences the molecular magnetic environment, modifying the longitudinal relaxation time T1 of 1H, and ultimately provoking the observed asymmetry in the enantiomeric response.

Ionospheric and Thermospheric Responses to the Recent Strong Solar Flares on 6 September 2017

AbstractTwo solar flares X2.2 and X9.3 erupted over the active region 2673 on 6 September 2017, and the second flare is the strongest since 2005. In order to investigate the ionospheric and thermospheric responses to the two solar flares, the global total electron content and the critical frequency of F2 layer obtained from GPS stations and ionosondes are used. The results indicate that the ionosphere in the sunlit hemisphere increased significantly with magnitudes of 0.1 and 0.5 total electron content units for the X2.2 and X9.3 solar flares, respectively. The electron density, thermospheric neutral density, and neutral temperature simulated by the Thermosphere‐Ionosphere Electrodynamics Global Circulation Model show that the behavior of ionospheric and thermospheric responses is different. The ionospheric disturbances occurred at the altitude ranges of 150−300 km, and the thermospheric responses occurred at the altitudes of 250−400 km are caused by solar extreme ultraviolet and ultraviolet photons, respectively. Both ionospheric and thermospheric responses are proportional to the height within their corresponding altitude ranges. Observations and simulations reveal that the ionospheric and thermospheric responses are nonlinearly dependent on the solar zenith angle. The disturbances caused by the X2.2 solar flare are symmetric, but the X9.3 solar flare are not. The O/N2 density ratio simulated by Thermosphere‐Ionosphere Electrodynamics Global Circulation Model increases from lev0 to lev5.0 pressure surface with a magnitude of 0.1–1.8, while the ratio decreases in the American sector with a magnitude of −0.6 to −0.3. The longitudinal asymmetry of O/N2 density ratio is a major contributor to the longitudinal asymmetry of ionospheric and thermospheric responses.

Learning mechanisms in memristor networks based on GaN nanomembranes

We demonstrate experimentally that single crystalline GaN nanomembranes arranged in simple networks exhibit learning mechanisms such as habituation and dishabituation followed by storage of the response to a certain electrical stimulus. These artificial learning mechanisms are analogous to non-associative learning processes which are identical in simple animals and human beings. We found that the learning time depends on the number of GaN membranes in parallel, and this parameter decreases by 30% when three memristors are connected in parallel compared to the learning time of a single memristor. Moreover, an increased number of parallel memristors reduces the eventual asymmetry in the temporal response of the circuit at positive and negative step voltages.

Asymmetric neural responses for facial expressions and anti-expressions

Face recognition requires identifying both the invariant characteristics that distinguish one individual from another and the variations within the individual that correspond to emotional expressions. Both have been postulated to be represented via a norm-based code, in which identity or expression are represented as deviations from an average or neutral prototype. We used Fast Periodic Visual Stimulation (FPVS) with electroencephalography (EEG) to compare neural responses for neutral faces, expressions and anti-expressions. Anti-expressions are created by projecting an expression (e.g. a happy face) through the neutral face to form the opposite facial shape (anti-happy). Thus expressions and anti-expressions differ from the norm by the same “configural” amount and thus have equivalent but opposite status with regard to their shape, but differ in their ecological validity. We examined whether neural responses to these complementary stimulus pairs were equivalent or asymmetric, and also tested for norm-based coding by comparing whether stronger responses are elicited by expressions and anti-expressions than neutral faces. Observers viewed 20 s sequences of 6 Hz alternations of neutral faces and expressions, neutral faces and anti-expressions, and expressions and anti-expressions. Responses were analyzed in the frequency domain. Significant responses at half the frequency of the presentation rate (3 Hz), indicating asymmetries in responses, were observed for all conditions. Inversion of the images reduced the size of this signal, indicating these asymmetries are not solely due to differences in the low-level properties of the images. While our results do not preclude a norm-based code for expressions, similar to identity, this representation (as measured by the FPVS EEG responses) may also include components sensitive to which configural distortions form meaningful expressions.

Satellite-Observed Precipitation Response to Ocean Mesoscale Eddies

Among various forms of atmospheric response to ocean mesoscale eddies, the rainfall response is the most difficult to quantify and is subject to considerable uncertainty. Here the robustness of the rainfall response is examined by comparing three different satellite-derived rainfall datasets: the Tropical Rainfall Measuring Mission (TRMM) Multisatellite Precipitation Analysis (TMPA), NOAA Climate Prediction Center (CPC) morphing technique (CMORPH) global precipitation, and the newly available Integrated Multisatellite Retrievals for Global Precipitation Measurement (IMERG) that is based on the latest remote sensing technology with finer spatial and temporal resolution. Results show that all datasets exhibit a similar rainfall response to ocean eddies, but the amplitude of the rainfall response is much stronger in IMERG than in the other two, despite the fact that IMERG provides the weakest time-mean rainfall estimate. In situ validation against the NOAA’s Ocean Climate Stations Project (OCS) Kuroshio Extension Observatory (KEO) buoy rainfall measurement shows that IMERG is more accurate in estimating both the mean value of rainfall and its intensity distribution than the other two products, at least in the Kuroshio Extension region. Further analysis reveals that 1) eddy-induced precipitation response is significantly stronger in winter than in summer, and 2) warm-eddy-induced rainfall response is considerably stronger than cold-eddy-induced response, and these asymmetries in rainfall response are more robust in IMERG than in the other two datasets. Documenting and analyzing these asymmetric rainfall responses is important for understanding the potential role of ocean eddies in forcing the large-scale atmospheric circulation and climate.

Dayside Magnetospheric and Ionospheric Responses to a Foreshock Transient on 25 June 2008: 1. FLR Observed by Satellite and Ground‐Based Magnetometers

AbstractAs one type of driver of magnetospheric Alfvén waves, foreshock transients have received less attention than, for example, the Kelvin‐Helmholtz instability, discrete and broadband frequency solar wind dynamic pressure oscillations, and interplanetary shocks. Previous works show that foreshock transients can induce both Alfvén mode and compressional mode Pc 3–5 ULF waves inside the magnetosphere. However, to our knowledge, none of these reported Pc 3–5 waves, induced by foreshock transients, are proved to be localized in the magnetosphere. In this paper, using in situ and ground‐based observations, we report the generation of localized magnetospheric compressional waves and field line resonances (FLRs) by a foreshock transient. Both the foreshock transient and Pc 5 ULF waves were found on the duskside; while on the morning side of the magnetosphere, no clear wave signatures were captured. Our results demonstrate that in addition to the global effects of foreshock transients on the magnetosphere reported earlier, foreshock transients can also generate localized magnetospheric responses in the Pc 5 range with clear dawn‐dusk asymmetry. A suite of eight dayside spacecraft plus ground magnetometer measurements make possible the determination of the foreshock transient driver and dawn‐dusk asymmetry of the magnetospheric response not previously reported with such a complete data set.

Experimental and computational study on phase transformations in superelastic NiTi snake-like spring

One of the major drawbacks of mechanically loaded superelastic NiTi structures is their short fatigue life, which is believed to be related to cyclic variation of relatively large strains due to martensitic phase transformation. Thus, determination of spatial and temporal evolution of martensite within such structures may help to avoid critical loading scenarios in practical applications. However, required data are often difficult to obtain and interpret. In this study, we present a possible approach to this issue and illustrate it on a comprehensive study on isothermal stretching of a planar superelastic NiTi snake-like spring, a prospective versatile superelastic actuator element. Specifically, we complement mechanical, electrical resistivity and x-ray diffraction methods with finite element simulations employing a progressive macroscopic constitutive model and reconstruct the spatial distribution in the most strained parts of the spring. We reveal that martensite tends to form in a localized manner there, which affects distribution of phases as well as strains along the spring. Results also exemplify the substantial influence of tension-compression asymmetry of the superelastic response to distribution of phases within cross-section of a bent structure.

Modeling the Interactions between Volatility and Returns using EGARCH‐M

An EGARCH‐M model, in which the logarithm of scale is driven by the score of the conditional distribution, is shown to be theoretically tractable as well as practically useful. A two‐component extension makes it possible to distinguish between the short‐ and long‐run effects of returns on volatility, and the resulting short‐ and long‐run volatility components are then allowed to have different effects on returns, with the long‐run component yielding the equity risk premium. The EGARCH formulation allows for more flexibility in the asymmetry of the volatility response (leverage) than standard GARCH models and suggests that, for weekly observations on two major stock market indices, the short‐term response is close to being anti‐symmetric.

Physiological responses to leptin levels in lipodystrophy: a model for other hypoleptinemias?

Brown et al. report that two weeks of exogenous leptin administration to leptin-naive individuals with lipodystrophy resulted in increased energy expenditure and lipolysis, decreased ectopic liver fat, improved hepatic and peripheral insulin sensitivity, and attenuated dyslipidemia. Leptin withdrawal in individuals with lipodystrophy did not produce reciprocal effects on these phenotypes and resulted in significant improvements only in hepatic insulin sensitivity. This asymmetry in responses to leptin initiation and cessation is consistent with the other aspects of leptin biology that are dependent on the metabolic context in which this adipocyte-derived hormone functions.

Shade and drought growth conditions strongly impact dynamic responses of stomata to variations in irradiance in Nicotiana tabacum

Recent research has made progress in describing stomatal dynamics in terms of speed, amplitude of response, lag time and response time. However, little is known about the impact of growth conditions on the rapidity of stomatal movements, and their relationship with stomatal morphology within a species. We measured stomatal dynamics during opening and closing in response to changes in irradiance in tobacco plants (Nicotiana tabacum) grown under “Control”, “Drought” and “Shade” treatments. Growth conditions strongly changed the rapidity of stomatal responses to irradiance. The “Drought” treatment considerably accelerated the response and “Shade” treatment slowed it down when compared to “Control”. We confirmed for the “Control” treatment the known asymmetry of response, with closing faster than opening, but interestingly the asymmetry disappeared under both treatments. Only stomatal density and index were affected by the growth conditions, not stomatal size and form. Thus, the observed variation in stomatal closing and opening dynamic parameters (speed, amplitude, lag time, response time) was not due to a variation in the size of the stomata, and only a marginal relationship between speed of the stomatal response and stomatal density was observed. These results suggest that physiological factors might be the main driver of variations in stomatal conductance dynamics within a species grown under different environmental conditions.