Petrified Forest Member Research Articles

A New Look at Landslides of the Vermilion and Echo Cliffs, Northern Arizona

ABSTRACT The Vermilion and Echo Cliffs form a nearly continuous escarpment more than 160 km long within the Colorado Plateau physiographic province of North America. The cliffs overlie the Marble Platform in northern Arizona and are located along the Colorado River, just upstream of the Grand Canyon. Large rotational block landslides mantle the erosional escarpment along most of its extent. Although these landslides have been noted for over 100 years, their likely origin has never been explained. Landslide failure surfaces appear to be influenced by the Petrified Forest Member of the Triassic Chinle Formation, a shale layer containing smectite clay weathered from volcanic ash. Although landslides are common along the majority of escarpments comprising the Colorado Plateau where the Petrified Forest Member and other shales outcrop, most appear to have been inactive since the early Holocene. Multiple generations of landslides and remnants of previous slides exist up to 3 km from the present cliff face. Multiple working hypotheses explaining these landslides are explored in this article, including past landslides and/or lava dams along the Colorado River within the Grand Canyon, periods of wetter climate with higher groundwater levels, and earthquakes related to nearby faulting and volcanism. Various sliding modes along these cliffs are described along with potential triggering mechanisms. Back-analysis of these landslides has been conducted using mechanical properties of the formations involved as well as varying groundwater levels. Calculated factors of safety for existing slides under present conditions are greater than unity, consistent with their apparent stability.

New dinosauromorph specimens from Petrified Forest National Park and a global biostratigraphic review of Triassic dinosauromorph body fossils

Dinosauromorph specimens from Petrified Forest National Park have been recovered from four major collecting efforts since 1982, including the most recent paleontological inventory of new park lands acquired in 2011. Additionally, an emphasis on understanding the stepwise acquisition of character traits along the dinosaurian lineage has helped identify previously collected specimens in museum collections. Here we briefly describe and use apomorphies to identify 32 additional dinosauromorph specimens found at Petrified Forest National Park, bringing the total number of dinosauromorph specimens presently known from the park to 50, a 600% increase since the year 2000. These specimens are all Norian in age and come from the Blue Mesa Member, Sonsela Member, and Petrified Forest Member of the Chinle Formation. These include the proximal end of a tibia that represents the oldest unambiguous dinosaur specimen from the Chinle Formation. We then contextualize these specimens with the dinosauromorph assemblages from the Norian of Utah, Colorado, New Mexico, and Texas, as well as the Carnian and Norian dinosauromorph assemblages from South America, Africa, and Europe. Despite increased sampling we still find no evidence for sauropodomorph and ornithischian dinosaurs in Western North America. An increase in sampling, combined with the use of apomorphies to identify collected specimens, will continue to improve the global dinosauromorph fossil record that can be used to answer questions on biochronology and the evolutionary history of the avian lineage.

LA-ICPMS U–Pb geochronology of detrital zircon grains from the Coconino, Moenkopi, and Chinle formations in the Petrified Forest National Park (Arizona)

Abstract. Uranium–lead (U–Pb) geochronology was conducted by laser ablation – inductively coupled plasma mass spectrometry (LA-ICPMS) on 7175 detrital zircon grains from 29 samples from the Coconino Sandstone, Moenkopi Formation, and Chinle Formation. These samples were recovered from ∼ 520 m of drill core that was acquired during the Colorado Plateau Coring Project (CPCP), located in Petrified Forest National Park (Arizona). A sample from the lower Permian Coconino Sandstone yields a broad distribution of Proterozoic and Paleozoic ages that are consistent with derivation from the Appalachian and Ouachita orogens, with little input from local basement or Ancestral Rocky Mountain sources. Four samples from the Holbrook Member of the Moenkopi Formation yield a different set of Precambrian and Paleozoic age groups, indicating derivation from the Ouachita orogen, the East Mexico arc, and the Permo-Triassic arc built along the Cordilleran margin. A total of 23 samples from the Chinle Formation contain variable proportions of Proterozoic and Paleozoic zircon grains but are dominated by Late Triassic grains. LA-ICPMS ages of these grains belong to five main groups that correspond to the Mesa Redondo Member, Blue Mesa Member and lower part of the Sonsela Member, upper part of the Sonsela Member, middle part of the Petrified Forest Member, and upper part of the Petrified Forest Member. The ages of pre-Triassic grains also correspond to these chronostratigraphic units and are interpreted to reflect varying contributions from the Appalachian orogen to the east, Ouachita orogen to the southeast, Precambrian basement exposed in the ancestral Mogollon Highlands to the south, East Mexico arc, and Permian–Triassic arc built along the southern Cordilleran margin. Triassic grains in each chronostratigraphic unit also have distinct U and thorium (Th) concentrations, which are interpreted to reflect temporal changes in the chemistry of arc magmatism. Comparison of our LA-ICPMS ages with available chemical abrasion thermal ionization mass spectrometry (CA-TIMS) ages and new magnetostratigraphic data provides new insights into the depositional history of the Chinle Formation, as well as methods utilized to determine depositional ages of fluvial strata. For parts of the Chinle Formation that are dominated by fine-grained clastic strata (e.g., mudstone and siltstone), such as the Blue Mesa Member and Petrified Forest Member, all three chronometers agree (to within ∼ 1 Myr), and robust depositional chronologies have been determined. In contrast, for stratigraphic intervals dominated by coarse-grained clastic strata (e.g., sandstone), such as most of the Sonsela Member, the three chronologic records disagree due to recycling of older zircon grains and variable dilution of syn-depositional-age grains. This results in LA-ICPMS ages that significantly predate deposition and CA-TIMS ages that range between the other two chronometers. These complications challenge attempts to establish a well-defined chronostratigraphic age model for the Chinle Formation.

Ichnofossil assemblages and palaeosols of the Upper Triassic Chinle Formation, south-eastern Utah (USA): Implications for depositional controls and palaeoclimate

The Upper Triassic Chinle Formation in the Stevens Canyon area in south-eastern Utah represents fluvial, palustrine, and lacustrine strata deposited in a continental back-arc basin on the western edge of Pangea. Previous investigations interpreted a megamonsoonal climate with increasing aridity for the Colorado Plateau towards the end of the Triassic. In this study, we systematically integrate ichnological and pedological features of the Chinle Formation into ichnopedofacies to interpret palaeoenvironmental and palaeoclimatic variations in the north-eastern part of the Chinle Basin. Seventeen ichnofossil morphotypes and six palaeosol orders are combined into twelve ichnopedofacies, whose development was controlled by autocyclic and allocyclic processes and hydrology. Ichnopedofacies are used to estimate palaeoprecipitation in conjunction with appropriate modern analogue latitudinal and geographic settings. In the north-east Chinle Basin, annual precipitation was ~1100–1300 mm in the Petrified Forest Member. Precipitation levels were >1300 mm/yr at the base of the lower Owl Rock Member, decreased to ~700–1100 mm/yr, and then to ~400–700 mm/yr. Two drying upward cycles from ~1100 mm/yr to ~700 mm/yr occurred in the middle and upper part of the Owl Rock Member. In the overlying Church Rock Member, precipitation decreased from ~400 mm/yr at the base of the unit to ~25–325 mm/yr at the end of Chinle Formation deposition. Ichnopedofacies indicate monsoonal conditions persisted until the end of the Triassic with decreasing precipitation that resulted from the northward migration of Pangea. Ichnopedofacies in the northeast Chinle Basin indicate both long-term drying of climate and short-term, wet-dry fluctuations.

Upper Triassic lithostratigraphy, depositional systems, and vertebrate paleontology across southern Utah

The Chinle Formation and the lower part of the overlying Wingate Sandstone and Moenave Formation were deposited in fluvial, lacustrine, paludal, and eolian environments during the Norian and Rhaetian stages of the Late Triassic (~230 to 201.3 Ma), during which time the climate shifted from subtropical to increasingly arid. In southern Utah, the Shinarump Member was largely confined to pre-Chinle paleovalleys and usually overprinted by mottled strata. From southeastern to southwestern Utah, the lower members of the Chinle Formation (Cameron Member and correlative Monitor Butte Member) thicken dramatically whereas the upper members of the Chinle Formation (the Moss Back, Petrified Forest, Owl Rock, and Church Rock Members) become erosionally truncated; south of Moab, the Kane Springs beds are laterally correlative with the Owl Rock Member and uppermost Petrified Forest Member. Prior to the erosional truncation of the upper members, the Chinle Formation was probably thickest in a southeast to northwest trend between Petrified Forest National Park and the Zion National Park, and thinned to the northeast due to the lower Chinle Formation lensing out against the flanks of the Ancestral Rocky Mountains, where the thickness of the Chinle is largely controlled by syndepositional salt tectonism. The Gartra and Stanaker Members of the Ankareh Formation are poorly understood Chinle Formation correlatives north of the San Rafael Swell. Osteichthyan fish, metoposaurid temnospondyls, phytosaurids, and crocodylomorphs are known throughout the Chinle Formation, although most remains are fragmentary. In the Cameron and Monitor Butte Members, metoposaurids are abundant and non-pseudopalatine phytosaurs are known, as is excellent material of the paracrocodylomorph Poposaurus; fragmentary specimens of the aetosaurs Calyptosuchus, Desmatosuchus, and indeterminate paratypothoracisins were probably also recovered from these beds. Osteichthyans, pseudopalatine phytosaurs, and the aetosaur Typothorax are especially abundant in the Kane Springs beds and Church Rock Member of Lisbon Valley, and Typothorax is also known from the Petrified Forest Member in Capitol Reef National Park. Procolophonids, doswelliids, and dinosaurs are known but extremely rare in the Chinle Formation of Utah. Body fossils and tracks of osteichthyans, therapsids, crocodylomorphs, and theropods are well known from the lowermost Wingate Sandstone and Moenave Formation, especially from the St. George Dinosaur Discovery Site at Johnson Farm.

A juvenile Koskinonodon perfectus (Temnospondyli, Metoposauridae) from the Upper Triassic of Arizona and its implications for the taxonomy of North American metoposaurids

AbstractMetoposaurids are temnospondyl amphibians that are well known from Upper Triassic deposits in North America, Europe, India, and Africa. Two species of metoposaurids, Koskinonodon perfectus and Apachesaurus gregorii, are among the most common fossils found in the Petrified Forest National Park (PEFO) of the southwestern United States. The two are distinguished on the basis of several cranial traits and the morphology of their intercentra, and are more informally distinguished by their overall size and biostratigraphic range. While material of adult metoposaurids is extremely common in Upper Triassic deposits, described material of juveniles is very rare, which limits the study of ontogenetic trajectories that would assist in assessing whether diagnostic traits of the smaller A. gregorii are also ontogenetically influenced traits in juveniles of K. perfectus. Here we report on material from a juvenile specimen of K. perfectus from the Petrified Forest Member (Norian) of the Chinle Formation in the PEFO. This specimen is significant because it represents the highest known occurrence of the species within the PEFO, documents a rare occurrence of a juvenile specimen of a North American metoposaurid, and records a rare North American metoposaurid specimen that includes associated cranial and postcranial material. Additionally, the combination of cranial features diagnostic of K. perfectus and intercentra of proportions intermediate between K. perfectus and A. gregorii in this specimen raises questions about the utility of several characters for species discrimination in North American metoposaurids.

Upper Triassic lithostratigraphy, depositional systems, and vertebrate paleontology across southern Utah

The Chinle Formation and the lower part of the overlying Wingate Sandstone and Moenave Formation were deposited in fluvial, lacustrine, paludal, and eolian environments during the Norian and Rhaetian stages of the Late Triassic (~230 to 201.3 Ma), during which time the climate shifted from subtropical to increasingly arid. In southern Utah, the Shinarump Member was largely confined to pre-Chinle paleovalleys and usually overprinted by mottled strata. From southeastern to southwestern Utah, the lower members of the Chinle Formation (Cameron Member and correlative Monitor Butte Member) thicken dramatically whereas the upper members of the Chinle Formation (the Moss Back, Petrified Forest, Owl Rock, and Church Rock Members) become erosionally truncated; south of Moab, the Kane Springs beds are laterally correlative with the Owl Rock Member and uppermost Petrified Forest Member. Prior to the erosional truncation of the upper members, the Chinle Formation was probably thickest in a southeast to northwest trend between Petrified Forest National Park and the Zion National Park, and thinned to the northeast due to the lower Chinle Formation lensing out against the flanks of the Ancestral Rocky Mountains, where the thickness of the Chinle is largely controlled by syndepositional salt tectonism. The Gartra and Stanaker Members of the Ankareh Formation are poorly understood Chinle Formation correlatives north of the San Rafael Swell. Osteichthyan fish, metoposaurid temnospondyls, phytosaurids, and crocodylomorphs are known throughout the Chinle Formation, although most remains are fragmentary. In the Cameron and Monitor Butte Members, metoposaurids are abundant and non-pseudopalatine phytosaurs are known, as is excellent material of the paracrocodylomorph Poposaurus; fragmentary specimens of the aetosaurs Calyptosuchus, Desmatosuchus, and indeterminate paratypothoracisins were probably also recovered from these beds. Osteichthyans, pseudopalatine phytosaurs, and the aetosaur Typothorax are especially abundant in the Kane Springs beds and Church Rock Member of Lisbon Valley, and Typothorax is also known from the Petrified Forest Member in Capitol Reef National Park. Procolophonids, doswelliids, and dinosaurs are known but extremely rare in the Chinle Formation of Utah. Body fossils and tracks of osteichthyans, therapsids, crocodylomorphs, and theropods are well known from the lowermost Wingate Sandstone and Moenave Formation, especially from the St. George Dinosaur Discovery Site at Johnson Farm.

A new rauisuchid (Archosauria, Pseudosuchia) from the Upper Triassic (Norian) of New Mexico increases the diversity and temporal range of the clade.

Rauisuchids are large (2–6 m in length), carnivorous, and quadrupedal pseudosuchian archosaurs closely related to crocodylomorphs. Though geographically widespread, fossils of this clade are relatively rare in Late Triassic assemblages. The middle Norian (∼212 Ma) Hayden Quarry of northern New Mexico, USA, in the Petrified Forest Member of the Chinle Formation, has yielded isolated postcranial elements and associated skull elements of a new species of rauisuchid. Vivaron haydeni gen. et. sp. nov. is diagnosed by the presence of two posteriorly directed prongs at the posterior end of the maxilla for articulation with the jugal. The holotype maxilla and referred elements are similar to those of the rauisuchid Postosuchus kirkpatricki from the southwestern United States, but V. haydeni shares several maxillary apomorphies (e.g., a distinct dropoff to the antorbital fossa that is not a ridge, a straight ventral margin, and a well defined dental groove) with the rauisuchid Teratosaurus suevicus from the Norian of Germany. Despite their geographic separation, this morphological evidence implies a close phylogenetic relationship between V. haydeni and T. suevicus. The morphology preserved in the new Hayden Quarry rauisuchid V. haydeni supports previously proposed and new synapomorphies for nodes within Rauisuchidae. The discovery of Vivaron haydeni reveals an increased range of morphological disparity for rauisuchids from the low-paleolatitude Chinle Formation and a clear biogeographic connection with high paleolatitude Pangea.

Sedimentological constraints on the evolution of the Cordilleran arc: New insights from the Sonsela Member, Upper Triassic Chinle Formation, Petrified Forest National Park (Arizona, USA)

The Sonsela Member of the Upper Triassic Chinle Formation in Petrified Forest National Park, Arizona, forms a distinctive part of an extensive fluviolacustrine complex that blanketed the Colorado Plateau region and surrounding areas. Upper Triassic sediments were deposited within a continental backarc setting, synchronous with the development of the Cordilleran magmatic arc. Although the Chinle Formation was derived from a broad variety of source terranes, petrographic and geochemical data clearly fingerprint the Cordilleran arc as a region important to provenance. The Sonsela Member exhibits a stratigraphically abrupt change in depositional style and sedimentary architecture that likely reflects the dynamics of the evolving backarc fluvial system and syndepositional arc magmatism. The stratigraphic boundary between the two disparate depositional styles is marked by a laterally persistent red silcrete horizon that is also associated with a biostratigraphically significant faunal turnover. The lower Sonsela Member—a coarse-grained assemblage dominated by persistent sheet sandstone and conglomerate bodies—reflects a coarse-grained progradation of sediment that suggests low rates of dynamic subsidence and high rates of sediment input. The upper Sonsela Member—characterized by an overall finer-grained assemblage and ribbon sand bodies with an increased proportion of encasing mudstone—reflects an increasing rate of basin subsidence that likely persisted throughout the deposition of the overlying Petrified Forest Member. These packages reflect a sharp change from lower- to higher-sinuosity river systems that closely coincides with the increased rate of subsidence across the regional stratigraphic marker. Previous studies have suggested the importance of the silcrete horizon as a key biostratigraphic marker associated with a significant faunal turnover. This study supports the hypothesis that this horizon may represent a regional unconformity as opposed to local channeling, perhaps the product of an abrupt change in basin geometry followed by an increase in basin subsidence. Because the Sonsela Member is the stratigraphically lowest unit of the Chinle Formation dominated by volcanic clastic detritus on the Colorado Plateau, architectural analysis of the Sonsela Member serves as the most reliable indicator of short-term fluctuations in arc dynamics of the poorly preserved and disjointed Mesozoic portion of the Cordilleran magmatic arc.

An Arizona Clam Dig, Carnelian-Replaced Clams From the Petrified Forest Area, Apache County, Arizona

10.1080/00357529.2011.636265-F0001 Figure 1. The author's son Chris looking for carnelian-replaced clams in badlands typical of the Petrified Forest Member of the Chinle Formation near Petrified Fo...

High-precision U-Pb zircon geochronology of the Late Triassic Chinle Formation, Petrified Forest National Park (Arizona, USA): Temporal constraints on the early evolution of dinosaurs

The Triassic successions of the Colorado Plateau preserve an important record of vertebrate evolution and climate change, but correlations to a global Triassic framework are hampered by a lack of geochronological control. Tuffaceous sandstones and siltstones were collected from the Upper Triassic Chinle Formation exposed in the Petrified Forest National Park, Arizona, USA, within a refined stratigraphic context of 31 detailed measured sections. U-Pb analyses by the isotope dilution–thermal ionization mass spectrometry (ID-TIMS) method constrain maximum depositional ages for nine tuffaceous beds and provide new insights into the depositional history of the Chinle fluvial system. The base of the Blue Mesa Member of the Chinle Formation is placed at ca. 225 Ma, and the top of the Petrified Forest Member is placed at 208 Ma or younger, bracketing an ∼280-m-thick section that spans nearly the entire Norian Stage of the Late Triassic. Estimated sediment accumulation rates throughout the section reflect extensive hiatuses and/or sediment removal by channel erosion. The new geochronology for the Chinle Formation underscores the potential pitfalls of correlation of fluvial units based solely on lithostratigraphic criteria. A mid-Norian age (ca. 219–213 Ma) for the distinctive Sonsela conglomeratic sandstone bed constrains the Adamanian-Revueltian land vertebrate faunachron boundary. Our new data permit a significant time overlap between the lower Chinle sequence and the dinosauromorph-rich Ischigualasto Formation of northwestern Argentina. Near-contemporaneity of the trans-American deposits and their faunal similarities imply that early dinosaur evolution occurred rapidly across the Americas.

High-resolution U–Pb ages from the Upper Triassic Chinle Formation (New Mexico, USA) support a diachronous rise of dinosaurs

Though the Late Triassic preserves major paleoenvironmental fluctuations and is key for understanding the evolution of Mesozoic and modern terrestrial ecosystems, comparisons of Late Triassic non-marine sedimentary and fossil records are difficult because global correlations lack precise radioisotopic ages, and have instead been based upon unconstrained biostratigraphic ranges of palynomorph and vertebrate fossils. The Chinle Formation in southwestern North America preserves a major Late Triassic record of paleoenvironmental and biotic change, including significant early dinosaur fossils. Previous high-resolution radioisotopic age constraints for the formation are limited to a single U–Pb zircon age from the upper third of the formation. The extraction of a geologically meaningful age is challenging from these redeposited units and preference is given to considering the youngest age of a deposit as a maximum age and closest approximation of the depositional age. Because calculating a weighted mean age (or median age) from a group of ages from such deposits is often not adequate, the precision of our two new CA-TIMS single crystal zircon U–Pb ages from the Chinle Formation of New Mexico is limited to ca 0.3% (or ± 0.7 Ma) of the youngest crystal age. Our 206Pb/ 238U age of ~ 218 Ma from the Blue Mesa Member in Six Mile Canyon, western New Mexico, demonstrates that strata, palynomorphs, and vertebrate fossils previously considered to be late Carnian in age are actually middle Norian in age. Our new age of ~ 212 Ma from the Hayden Quarry within the Petrified Forest Member at Ghost Ranch, northern New Mexico, provides the first maximum age for important vertebrate assemblages from this area that record the rise of dinosaurs, and demonstrates that basal dinosauromorphs (‘dinosaur precursors’) co-existed with dinosaurs for at least 18 Ma. These new radioisotopic data allow a new correlation of the Chinle Formation to the Late Triassic timescale, suggesting that most if not all of the lower Chinle is Norian in age. This new correlation has global implications as it allows us to make more precise comparisons with early dinosaur assemblages from the Ischigualasto Formation of Argentina, indicating that Chinle dinosaur assemblages are significantly younger than those from South America. The revised age of the Chinle Formation also demonstrates that dinosaurs were much rarer in North America at a time when they were abundant in South America, supporting hypotheses of paleolatitudinal variation during the rise of dinosaurs.

Hindlimb osteology and distribution of basal dinosauromorphs from the Late Triassic of North America

ABSTRACT The recent discovery of early dinosauromorphs from North America demonstrates that they were contemporaries with dinosaurs and other basal archosaurs during a substantial portion of the Late Triassic Period. Hindlimb material (femora, tibiae, a fibula, astragalocalcanea, and phalanges) of Dromomeron romeri, a non-dinosauriform dinosauromorph from the Petrified Forest Member of the Chinle Formation from north-central New Mexico, is described. A new species of Dromomeron from the lower portion of the Chinle Formation (eastern Arizona) and Dockum Group (northern Texas) is also described, based on several disarticulated femora and tibiae. D. romeri, Lagerpeton, and the new taxon form the sister group to all other dinosauromorphs and demonstrate that this clade, Lagerpetidae, persisted well into the Norian. Lagerpetidae is supported by several synapomorphies: femoral head hook-shaped in medial and lateral views; ventral emargination on the anterolateral side of the femoral head; an enlarged posteromedial tuber of the proximal end of the femur; femoral crista tibiofibularis larger than the medial condyle; anteromedial corner of the distal end of the femur forms 90° or acute (>90°) angle; and a posterior ascending process of the astragalus. An ontogenetic series of the femur of Dromomeron indicates that some character states previously used in phylogenetic analyses of early dinosaurs may be ontogenetically variable.

A revision ofPoposaurus gracilis (Archosauria: Suchia) based on two new specimens from the Late Triassic of the southwestern U.S.A.

Two associated partial skeletons, one from the McCarty ranch (MOTTU site 0690) near Kalgary, Texas, Tecovas Formation, and the other from the Petrified Forest Member of the Chinle Formation, outside the Petrified Forest, Arizona, represent the most complete specimens ofPoposaurus gracilis found to date, and increase our understanding of the osteology of this rare pseudosuchian archosaur. The cervical centrum of TTUP-10419 possesses an accessory dorsal parapophyseal rib facet, as in “Lythrosuchus”. The ilia of these new fossils are the first ofPoposaurus to preserve the complete preacetabular blade, which is long and “blade-like” rather than “paddle-shaped” as previously thought. These specimens also confirm that isolated elements previously referred toPoposaurus (fused sacral vertebrae, the pubes and the ischia), but not present in the holotype indeed belong to that taxon. The morphology of the cervical centrum and the preacetabular blade of the ilium are identical to the much larger type of“Lythrosuchus” langstoni, suggesting that “Lythrosuchus” is a junior synonym ofPoposaurus. However, we retain the speciesPoposaurus langstoni nov. comb. on the basis of larger size, absence of a distinct ridge on the lateral surface of the ilium, and the lack of a pit on the proximal end of the ischium. A phylogenetic analysis suggests a monophyletic Poposauroidea and a paraphyletic “Rauisuchia”.

Multi-proxy paleosol evidence for middle and late Triassic climate trends in eastern Utah

Prior investigations suggest that alluvial lowlands of Pangea in the southwestern U.S.A. during the middle to late Triassic experienced an arid to semiarid climate with significant seasonality. Our investigation finds evidence for a global pluvial episode from paleosols in the middle to late Triassic section of eastern Utah. Multi-proxy paleosol evidence is used to quantify rainfall amounts, ambient temperatures and atmospheric CO 2 concentrations and infer associated plant formations related to ecosystem persistence. Rainfall estimates are derived from geochemical molecular weathering ratios; temperature and atmospheric CO 2 levels from stable oxygen and carbon isotopes of pedogenic carbonate, respectively; and ecosystem reconstruction by a combination of climate indexes and paleosol characteristics. Eight pedotypes were identified and include poorly drained aquic Entisols (Fisher), well-drained non-aquic Entisols (Salt Valley), calcic Aridisols (Castle Valley), calcic Inceptisols (Moab), cambic (Ute) and calcic (La Sal) Vertisols, dystric Inceptisols (Kokopeli), and argillic Alfisols (Slickrock). The middle Triassic Moenkopi Formation (Anisian) was dominated by Castle Valley paleosols with mean annual rainfall between 300 and 400 mm and mean annual temperature between 13 and 23 °C (mesic to hyperthermic). Based on these climate indexes and root traces, the associated plant formation was most likely desert shrub or dry woodland. The lower Chinle Formation (Carnian) contains a succession of Ute and La Sal paleosols that changed up-section to Kokopeli, Fisher and Slickrock paleosols. In accordance with geochemical and isotopic signatures from these pedotypes, rainfall amounts initially increased to between 700 and 900 mm, and then to between 1300 and 1400 mm. Temperatures estimated from the La Sal paleosol are approximately 18 °C (thermic) at this time. The lower Chinle marks a transition from dry woodland to open forest that appears to correlate with the formation of the Petrified Forest Member in Arizona. The upper Chinle Formation (Carnian–Norian) reveals a return to semiarid to subhumid conditions and the formation of Castle Valley, Moab and Salt Valley paleosols, all of which appear to have supported desert shrub or dry woodlands. Rainfall amounts decreased to between 400 and 600 mm with temperatures eventually increasing to 29 °C (hyperthermic). Using the paleosol isotopic barometer, atmospheric CO 2 estimates generally correlate with other proxy for the Triassic. The well-developed Alfisols and noncalcareous Inceptisols identified during the Carnian of the lower Chinle correlate with a previously identified global pluvial episode based on sedimentological and marine isotopic evidence, possibly in response to rifting of Pangea. It should not be assumed that the middle to late Triassic in continental alluvial lowlands supported a uniform, semiarid to arid climate, with strong seasonality.

Vertebrate Burrows from Triassic and Jurassic Continental Deposits of North America and Antarctica: Their Paleoenvironmental and Paleoecological Significance

Comparisons of recently identified Triassic and Jurassic continental trace fossils in North America and Antarctica to modern mammal and reptilian burrows facilitate the identification and interpretation of the ancient burrows as vertebrate in origin, indicating advanced behaviors. Hollow, bowl-shaped depressions in the Petrified Forest Member of the Upper Triassic Chinle Formation in Petrified Forest National Park, Arizona, are interpreted as nest-holes constructed possibly by phytosaurs, aeotosaurs, turtles, or rauisuchians. Large-diameter, multiple-branching, and interconnected burrows in the Owl Rock Member of the Chinle Formation in southeastern Utah are tentatively interpreted as vertebrate burrows indicating communal behavior. Complex, large-diameter burrows in the Salt Wash Member of the Upper Jurassic Morrison Formation near the Henry Mountains in southern Utah are interpreted as burrow systems of fossorial mammals. Large-diameter, gently dipping, simple, subhorizontal burrows in the Salt Wash Member are interpreted as possible dwelling burrows of sphenodontids. Other vertebrate trace fossils, such as the large-diameter burrows from the Lower Triassic Fremouw Formation in the Queen Maud Mountains of Antarctica, are reinterpreted as vertebrate burrows and were likely constructed by small mammal-like reptiles. These burrows were thought to have been enigmatic in origin and different from very large-diameter burrows interpreted as therapsid dwelling burrows. Descriptions and interpretations of all these trace fossils are important because most vertebrate ichnology research to date has focused on trackways or locomotion experiments with modern reptiles and birds. These Triassic and Jurassic ichnofossils represent fossorial and nesting behavior of several different groups of vertebrates. The Fremouw Formation burrows indicate fossorial behavior in several sizes of vertebrates, including small and large therapsids. The burrows were likely used for shelter, giving birth, raising young, and hibernation. During the early Mesozoic, the Fremouw landscapes were thought to have had cold winters due to their high-paleolatitude position. The bowl-shaped depressions from the Petrified Forest Member of the Chinle Formation likely represent the earliest known structures excavated by reptilians for the sole purpose of reproduction. Basic nest-hole architecture in extant reptiles with early Mesozoic ancestry has changed minimally in nearly 220 million years. Large-diameter, multiple branching, and interconnected burrows in the Owl Rock Member of the Chinle Formation were likely constructed by fossorial vertebrates that had communal family groups. Simple, gently dipping, subhorizontal burrows in the Morrison Formation were used by crocodiles or sphenodontids as dwelling structures constructed in firm, subaerially exposed substrata close to open bodies of water. Complex, large-diameter burrows in the Salt Wash Member suggest subsocial behavior of fossorial mammals, where the burrow was used for raising young, storage and disposal of food and wastes, and coping with episodic water inundation. Research demonstrates that fossorial behavior of reptiles, therapsids, and mammals was established by the beginning of the Mesozoic and prior to the break-up of Pangea. The basic architecture of vertebrate nest construction has changed little in nearly 280 million years. Fossorial burrowing behavior likely evolved several times in different vertebrate groups during this time. These basic burrow architectures are also used by invertebrate groups. This overlap in burrow architectures between vertebrates and invertebrates suggests strongly that paleoenvironmental and paleoclimatic organism-substrate relationships dictate the architecture used by the organism. These burrow morphologies indicate particular physicochemical conditions in terrestrial and freshwater settings that are unique to the continental realm.

A new crocodylomorph archosaur from the Upper Triassic of North Carolina

ABSTRACT A new taxon of sphenosuchian crocodylomorph, Dromicosuchus grallator, is described on the basis of a well-preserved, largely articulated partial skeleton from Late Triassic strata in the Durham sub-basin of the Deep River basin (Newark Supergroup) of Durham County, North Carolina. The holotype was preserved directly beneath the skeleton of a rauisuchian archosaur; this association, along with apparent bite marks to the head and neck of the crocodylomorph, suggests that the two animals died and were buried together during the act of predation. Dromicosuchus grallator is most closely related to Hesperosuchus agilis from the Petrified Forest Member of the Chinle Formation (late Carnian or early Norian) of Arizona and New Mexico and Saltoposuchus connectens from the Middle Stubensandstein (Löwenstein Formation; middle Norian) of Württemberg, Germany. The monophyly of Sphenosuchia is only weakly supported at present.

Isotopic age of the Black Forest Bed, Petrified Forest Member, Chinle Formation, Arizona: An example of dating a continental sandstone

Zircons from the Black Forest Bed, Petrified Forest Member, Chinle Formation, in Petrified Forest National Park, yield ages that range from Late Triassic to Late Archean. Grains were analyzed by multigrain TIMS (thermal-ionization mass spectrometry), single-crystal TIMS, and SHRIMP (sensitive, high-resolution ion-microprobe). Multiple-grain analysis yielded a discordia trajectory with a lower intercept of 207 6 2 Ma, which because of the nature of multiple-grain sampling of a detrital bed, is not considered conclusive. Analysis of 29 detrital-zircon grains by TIMS yielded UPb ages of 2706 6 6M a to 2066 6 Ma. Eleven of these ages lie between 211 and 216 6 6.8 Ma. Our statistical analysis of these grains indicates that the mean of the ages, 213 6 1.7 Ma, reflects more analytical error than geologic variability in sources of the grains. Grains with ages of ca. 1400 Ma were derived from the widespread plutons of that age exposed throughout the southwestern Cordillera and central United States. Twelve grains analyzed by SHRIMP provide 206 Pb*/ 238 U ages from 214 6 2M a to 200 6 4 Ma. We use these data to infer that cores of inherited material were present in many zircons and that single-crystal analysis provides an accurate estimation of the age of the bed. We further propose that, even if some degree of reworking has occurred, the very strong concentration of ages at ca. 213 Ma provides a maximum age for the Black Forest Bed of 213 6 1.7 Ma. The actual age of the bed may be closer to 209 Ma. Dating continental successions is very difficult when distinct ash beds are not clearly identified, as is the case in the Chinle Formation. Detrital zircons in the Black Forest Bed, however, are dominated by an acicular morphology with preserved delicate terminations. The shape of these crystals and their inferred environment of deposition in slow-water settings suggest that the crystals were not far removed from their site of deposition in space and likely not far in time. Plinian ash clouds derived from explosive eruptions along the early Mesozoic Cordilleran margin provided the crystals to the Chinle basin, where local conditions insured their preservation. In the case of the Black Forest Bed, the products of one major eruption may dominate the volcanic contribution to the unit. Volcanic detritus in the Chinle Formation was derived from multiple, distinct sources. Coarse pebble- to cobble-size material may have originated in eastern California and/or western Arizona, where Triassic plutons are exposed. Fine-grained detritus, in contrast, was carried in ash clouds that derived from caldera eruptions in east-central California or western Nevada.

Mapping sedimentary and volcanic units within and surrounding Petrified Forest National Park, Arizona, using Landsat-5 and SPOT panchromatic data

Landsat Thematic Mapper (TM) and SPOT panchromatic data were analysed to determine the lateral variation of the sedimentary and volcanic formations within and surrounding the Petrified Forest National Park in northeastern Arizona. Detailed geological mapping of portions of Petrified Forest National Park allowed for a correlation between TM images and the geological formations that aided in identifying these formations in unmapped areas outside and inside the park. A variety of image processing techniques, including band ratios, RGB composite images, selective principal component analysis and Brovey transformations were used in distinguishing these geological formations. Newly determined regions of the Lower and Upper parts of the Petrified Forest Member of the Chinle Formation were identified south of Pilot Rock and east of the Blue Mesa region. A final spectral reflectance-based geological map created by a maximum likelihood supervised classification algorithm provides a first-order approximation to the ...

Mapping sedimentary and volcanic units within and surrounding Petrified Forest National Park, Arizona, using Landsat-5 and SPOT panchromatic data

Landsat Thematic Mapper (TM) and SPOT panchromatic data were analysed to determine the lateral variation of the sedimentary and volcanic formations within and surrounding the Petrified Forest National Park in northeastern Arizona. Detailed geological mapping of portions of Petrified Forest National Park allowed for a correlation between TM images and the geological formations that aided in identifying these formations in unmapped areas outside and inside the park. A variety of image processing techniques, including band ratios, RGB composite images, selective principal component analysis and Brovey transformations were used in distinguishing these geological formations. Newly determined regions of the Lower and Upper parts of the Petrified Forest Member of the Chinle Formation were identified south of Pilot Rock and east of the Blue Mesa region. A final spectral reflectance-based geological map created by a maximum likelihood supervised classification algorithm provides a first-order approximation to the location of numerous sedimentary and volcanic units within and surrounding Petrified Forest National Park.