Permian-Triassic Interval Research Articles

Identification of the subaerial unconformity, genesis, and distribution in Lower Triassic Baikouquan Formation, Mahu Sag, Junggar Basin, Western China

The genesis of unconformities involves multiple influences on basin formation and evolution, which affects the favorable reservoir and hydrocarbon distribution. Within the Early Triassic Baikouquan Formation in the Mahu Sag of the Junggar Basin, we identified subaerial unconformities (SU), namely SU1 and SU2. SU1 is characterized by regional, irregular valleys, whereas SU2 displays extensive lateral extent with comparatively lower relief. These distinctions stem from detailed analyses involving cores, well data, seismic profiles, and spatial distributions, enriching our understanding of SUs in the Junggar Basin. Key findings from our study include: (1) SU occurrences within the Baikouquan Formation span the falling-stage systems tract (FSST) and expanding system tract (EST), corresponding to relative lake level changes. SU1 emerges during descending lake levels, whereas SU2 develops during rising levels. (2) The extensive uplift of the Junggar Basin's perimeter, driven by the late Hercynian movement, heightened topographic gradient shapes a steeper lake shoreline profile, fostering SU development. Arid paleoclimatic conditions during the Permian-Triassic interval accelerate SU formation in the FSST. (3) SU typically comprises regional compound scour surfaces (RCS) and river intermittent exposure surfaces (IES). Continuous winnowing at RCS led to relatively high primary porosities, IES can develop better secondary porosity with superposed freshwater leaching. The investigation into the subaerial unconformity in the study area holds broader implications for comprehending depositional processes, reservoir distribution, and petroleum prospects, both locally and within a global lake basin context.

Deep-water osmium-isotope record of the Permian–Triassic interval from Niushan, China reveals potential delayed volcanic signal post the mass extinction

The end Permian mass extinction event is one of the most severe biotic crisis of the Phanerozoic. The trigger of this event has been widely linked to massive volcanic activity associated with the Siberian Traps Large Igneous Province through the application of multiple geochronological (e.g., UPb and ArAr dating) and geochemical proxies (e.g., C, Hg, Zn, Os, etc.). Of these proxies, a few osmium (187Os/188Os) records are available for near-shore/shallow water depositional settings (e.g., Meishan, China; Opal Creek, Canada), which suggest multiple episodes of volcanism before, during and after the mass extinction event. Here, we present a new initial 187Os/188Os (Osi) stratigraphy across the deep-water Niushan Permian–Triassic boundary interval section of South China that reveals a single unradiogenic Os-isotope shift (from ~0.6 to 0.3) after the end Permian mass extinction event interval, which is interpreted to be related to volcanic activity. Above the unradiogenic excursion, a radiogenic shift to ~1.1 is detected across the Permian–Triassic boundary at Niushan that is correlative to the radiogenic Osi shift observed in the Meishan section. This increase in Os-isotope values is taken to reflect enhanced continental weathering. Both the unradiogenic and radiogenic excursions in the deep-water Niushan section are smaller in magnitude compared with the Os isotope profiles from more shallow-water sections of Meishan and Opal Creek. Our data suggest investigations of multiple sections across a variety of depositional environments may yield a more comprehensive understanding of the scale of the global perturbations related to volcanism and continental weathering intensity of the Permian–Triassic boundary interval.

Sedimentary and pedogenic record of seasonal humidity during the Permian-Triassic transition on the SE margin of Central European Basin (Holy Cross Mountains, Poland)

The transition from the late Permian to Early Triassic was associated with one of the biggest biotic crises in the entire Phanerozoic which affected life in both marine and continental settings. Unlike in the marine realm, the environmental dynamics in continental palaeoenvironments across the Permian-Triassic interval are still poorly understood. In order to fill this knowledge gap, we studied the Permo-Triassic continental succession from the SE part of the intracontinental Central European Basin (CEB). High-resolution sedimentological logging of over 1700 m of cores, supplemented by micromorphology of palaeosols, palynofacies, and quantitative spore-pollen analyses, allowed reconstructing major steps in palaeoenvironmental evolution, and assessing the magnitude of extrinsic factors responsible for such development. The studied interval is dominated by redbeds deposited in arid, semi-arid, and sub-humid subenvironments in which palaeosol profiles are important elements of facies architecture. Vertical and lateral facies arrangement indicate three major development stages associated with progradation of a large distributary fluvial system towards the centre of the basin. These stages correspond to the main lithostratigraphic units in the studied area. In the first stage, the saline mudflats were replaced by muddy floodplain facies and then by medial and proximal parts of the ephemeral fluvial system of the Podzamcze Formation. The overlying Siodła Formation represents the stage of environmental stabilisation associated with widespread pedogenesis resulting in the formation of thick cumulative palaeosol profiles. The last stage was marked by the influx of coarse-grained fluvial sediments of the Jaworzna Formation and change in soil character towards less evolved types in overbank facies of the Szczukowice Formation. The change in the sedimentary record and soil types reflects the successive humidification in the SE part of the CEB, supported by increasing contribution of the hygrophytic over mostly xerophytic palynomorphs in the upper Permian palynological assemblage. This humidification trend was likely an environmental response to global warming in the late Permian.

Floral Assemblages and Patterns of Insect Herbivory during the Permian to Triassic of Northeastern Italy.

To discern the effect of the end-Permian (P-Tr) ecological crisis on land, interactions between plants and their insect herbivores were examined for four time intervals containing ten major floras from the Dolomites of northeastern Italy during a Permian–Triassic interval. These floras are: (i) the Kungurian Tregiovo Flora; (ii) the Wuchiapingian Bletterbach Flora; (iii) three Anisian floras; and (iv) five Ladinian floras. Derived plant–insect interactional data is based on 4242 plant specimens (1995 Permian, 2247 Triassic) allocated to 86 fossil taxa (32 Permian, 56 Triassic), representing lycophytes, sphenophytes, pteridophytes, pteridosperms, ginkgophytes, cycadophytes and coniferophytes from 37 million-year interval (23 m.yr. Permian, 14 m.yr. Triassic). Major Kungurian herbivorized plants were unaffiliated taxa and pteridosperms; later during the Wuchiapingian cycadophytes were predominantly consumed. For the Anisian, pteridosperms and cycadophytes were preferentially consumed, and subordinately pteridophytes, lycophytes and conifers. Ladinian herbivores overwhelming targeted pteridosperms and subordinately cycadophytes and conifers. Throughout the interval the percentage of insect-damaged leaves in bulk floras, as a proportion of total leaves examined, varied from 3.6% for the Kungurian (N = 464 leaves), 1.95% for the Wuchiapingian (N = 1531), 11.65% for the pooled Anisian (N = 1324), to 10.72% for the pooled Ladinian (N = 923), documenting an overall herbivory rise. The percentage of generalized consumption, equivalent to external foliage feeding, consistently exceeded the level of specialized consumption from internal feeding. Generalized damage ranged from 73.6% (Kungurian) of all feeding damage, to 79% (Wuchiapingian), 65.5% (pooled Anisian) and 73.2% (pooled Ladinian). Generalized-to-specialized ratios show minimal change through the interval, although herbivore component community structure (herbivore species feeding on a single plant-host species) increasingly was partitioned from Wuchiapingian to Ladinian. The Paleozoic plant with the richest herbivore component community, the coniferophyte Pseudovoltzia liebeana, harbored four damage types (DTs), whereas its Triassic parallel, the pteridosperm Scytophyllum bergeri housed 11 DTs, almost four times that of P. liebeana. Although generalized DTs of P. liebeana were similar to S. bergeri, there was expansion of Triassic specialized feeding types, including leaf mining. Permian–Triassic generalized herbivory remained relatively constant, but specialized herbivores more finely partitioned plant-host tissues via new feeding modes, especially in the Anisian. Insect-damaged leaf percentages for Dolomites Kungurian and Wuchiapingian floras were similar to those of lower Permian, north-central Texas, but only one-third that of southeastern Brazil. Global herbivore patterns for Early Triassic plant–insect interactions remain unknown.

An extensive anoxic event in the Triassic of the South China Block: A pyrite framboid study from Dajiang and its implications for the cause(s) of oxygen depletion

Water column oxygen deficiency has been considered as a potent driver of the extinction of marine benthos, and is a main feature of marine environments in the aftermath of the end-Permian mass extinction. The record of Permian-Triassic anoxia is more complex than previously thought, and is seen to vary between different palaeogeographic settings, but a full understanding is hindered by a paucity of evidence. During the Permian-Triassic interval the South China Block was located equatorially with Palaeotethys to the north and western Panthalassa to the south. This specific configuration provides a unique opportunity to compare the extent and duration of oxygen deficiency in Palaeotethys and Panthalassa under broadly similar climatic conditions. Sedimentary facies and pyrite framboid size-frequency distributions suggest that the oxygen-poor conditions became widespread across the shallow-marine carbonate platform of the South China Block immediately above the Permian-Triassic boundary and mass extinction level. Oxygen deficiency was most intense at the southern margin of the block where it met Panthalassa. Proposed drivers of the expansion of oxygen minimum zones into platform settings include enhanced terrigenous input and/or ocean stratification, or alternatively the upwelling of nutrient-rich deep ocean water. The former mechanisms are theoretically more likely to have operated in the relatively restricted Palaeotethys which was surrounded by ancient lands. In contrast, Panthalassa would likely have experienced stronger oceanic circulation and therefore be more susceptible to the effects of upwelling. Although variations in the record of the South China Block anoxic event might reflect local factors, the greater intensity of oxygen deficiency and a concomitant larger negative shift in carbonate carbon isotopes on its Panthalassan margin point to a key role for upwelling. This mechanism was likely a major driver of the Permian-Triassic global oceanic anoxic event, which itself was at least partly responsible for the ongoing inhospitable conditions and delayed recovery following the end-Permian extinction.

Lilliput effect in freshwater ostracods during the Permian–Triassic extinction

Abstract The Lilliput effect following the Permian–Triassic mass extinction and its aftermath has been documented in a variety of marine animal groups, but it is less known in terrestrial and freshwater invertebrates. Here we present new investigations of the size variations of terrestrial ostracods of the genus Darwinula based on fossil records from a Permian–Triassic section on the northern limb of the Dalongkou Anticline section in Northwest China. Quantitative analyses reveal that ostracod test sizes decreased sharply through the terrestrial Permian–Triassic mass extinction interval. The Lilliput effect in terrestrial ostracods is characterized by the extinction of large taxa and the rise of small-sized and elongate new forms, coupled with the dramatic loss of conchostracans, charophytes, and the blooming of lycopod spores. The size decrease in terrestrial ostracods, following the biotic crisis through the Permian–Triassic interval, was probably triggered by several interacting events, including global warming, anoxia, and enhanced sediment input following acid rain and wildfire.

Palynostratigraphy at the Permian-Triassic boundary of the Amb section, Salt Range, Pakistan

Palynostratigraphy across the Permian–Triassic of the Salt Range and Surghar Range area remains poorly known because of the predominance of unsuitable facies and preservation (Nammal, Chhidru, Chitta-Landu and Narmia). Dolomite and limestone in the basal Mianwali Formation prevented the establishment of a palynostratigraphic scheme for the basal Early Triassic (Hermann et al. 2012). Based on the study of a new Permian–Triassic section in the Amb valley where siltstone layers are intercalated in the basal Mianwali Formation (Kathwai Member), we describe a new record of palynological assemblages from the uppermost Permian Chhidru Formation and the lowermost part of the Early Triassic Mianwali Formation. The latest Permian Chhidru 2 sporomorph association occurs in the uppermost part of the Chhidru Formation, the so-called ‘white sandstone unit’. It is marked by the dominance of bisaccate pollen grains (mainly Protohaploxypinus spp.) and cavate trilete spores (mainly Kraeuselisporites spp.). Here, the previously described lowermost Triassic palynozone PTr 1 is subdivided into two subzones. PTr 1a assemblages are described from the Griesbachian Kathwai Member. They are characterized by continued high bisaccate pollen grain abundances and the presence of the lycopod genera Lundbladispora spp. and Densoisporites spp. In these assemblages, typical ‘Permian taxa’ and typical ‘Triassic taxa’ are mixed. The composition of the middle Dienerian PTr 1b assemblages corresponds to the previously described PTr 1 assemblage from Nammal and Chitta-Landu. Lundbladispora spp. and Densoisporites spp. account for ca. 80% of the total assemblage in PTr 1b. Although the Griesbachian assemblages record a decrease in diversity, the middle Dienerian assemblages represent the poorest assemblages of the studied Permian–Triassic interval. Reduviasporonites occurs only sporadically throughout the Amb succession.

Floral biodiversity and geology of the Talcher Basin, Orissa, India during the Permian–Triassic interval

The review paper provides an updated account of the previous and recently published records concerning the palaeobiology and the geology of the Talcher Basin of Orissa State, India. We conclude that fossil floral species in this basin originated in the earliest Permian Talchir Formation and evolved and diversified through the Karharbari Fm., Barakar Fm., Barren Measures Fm. and the uppermost Kamthi Fm. (Late Permian–Triassic). The megaflora and the palynology of the different formations of the basin are also discussed briefly. The geological setting of the basin along with the status of different formations (especially the Kamthi Formation) has been redefined. The post‐Barakar Fm. rocks, earlier retained in the Raniganj/Kamthi, Panchet and Mahadeva formations in this basin, have been critically assessed and redefined as the Lower and Upper Kamthi formations of Late Permian and Triassic ages, respectively. Accordingly, the geological map of the basin has been modified. Permian deposits (particularly the Barakar and the lower Kamthi formations) not only have the best preserved flora but also possess the highest diversity, whereas the upper Kamthi Triassic sediments have a meagre number of taxa. The plant diversity of the basin has been discussed in detail to interpret the development of the flora, evolutionary trends and palaeoenvironments of the basin. The patchy Gangamopteris vegetation of the Talchir glacial phase has ultimately evolved and diversified through time (Karharbari Fm. to Lower Kamthi Fm.) and gave rise to the thick dense swampy forests consisting of large Glossopteris trees and other shade‐loving under‐storied pteridophytes. Several groups of plants including spores and pollen have disappeared in a ladder pattern during the Permian–Triassic interval (Lower Kamthi–Upper Kamthi Fm.) and, similarly, in steps, many new fore‐runners appeared in the Upper Kamthi Formation.Records of marine acritarchs and ichnofossils in this basin at various Permian–Triassic levels demonstrate that there were marine influences. These features suggest a paralic (coastal marine to deltaic) mode of origin of the coal beds and associated sediments in the basin. The present study also advocates the continued survival of plants, rather than a mass extinction near the vicinity of the Permian–Triassic (P–T) boundary in this basin. Copyright © 2012 John Wiley & Sons, Ltd.

Global diversity and phylogeny of the Asteroidea (Echinodermata).

Members of the Asteroidea (phylum Echinodermata), popularly known as starfish or sea stars, are ecologically important and diverse members of marine ecosystems in all of the world's oceans. We present a comprehensive overview of diversity and phylogeny as they have figured into the evolution of the Asteroidea from Paleozoic to the living fauna. Living post-Paleozoic asteroids, the Neoasteroidea, are morphologically separate from those in the Paleozoic. Early Paleozoic asteroid faunas were diverse and displayed morphology that foreshadowed later living taxa. Preservation presents significant difficulties, but fossil occurrence and current accounts suggests a diverse Paleozoic fauna, which underwent extinction around the Permian-Triassic interval was followed by re-diversification of at least one surviving lineage. Ongoing phylogenetic classification debates include the status of the Paxillosida and the Concentricycloidea. Fossil and molecular evidence has been and continues to be part of the ongoing evolution of asteroid phylogenetic research. The modern lineages of asteroids include the Valvatacea, the Forcipulatacea, the Spinlosida, and the Velatida. We present an overview of diversity in these taxa, as well as brief notes on broader significance, ecology, and functional morphology of each. Although much asteroid taxonomy is stable, many new taxa remain to be discovered with many new species currently awaiting description. The Goniasteridae is currently one of the most diverse families within the Asteroidea. New data from molecular phylogenetics and the advent of global biodiversity databases, such as the World Asteroidea Database (http://www.marinespecies.org/Asteroidea/) present important new springboards for understanding the global biodiversity and evolution of asteroids.

Stratigraphy of Upper Permian and Lower Triassic Strata of the Žiri Area (Slovenia)

The paper deals with the stratigraphy of Late Permian and Early Triassic strata of the Lukač section in the Žiri area of western Slovenia. This is the only section presently known in the External Dinarides where the Permian- Triassic boundary is defined following international criteria based on the first appearance of the conodont Hindeodus parvus. The following lithostratigraphic units have been formalized: the Bellerophon Limestone and Evaporite-dolomite Members of the Bellerophon Formation and the Lukač Formation with the three members, the Transitional Beds, Streaky Limestone and Carbonate-clastic Member. The paper presents the results of micropaleontological study based on foraminifers and conodonts as well as petrographic and sedimentologic research results. The investigation of conodont assemblages enabled the conodont biozonation of the Permian-Triassic interval of the studied Lukač section.

Conodont biostratigraphy and lithostratigraphy across the Permian-Triassic boundary at the Lukač section in Western Slovenia

Detailed conodont biostratigraphy and lithostratigraphy of the Late Permian and Early Triassic were studied at the LukaC section in western Slovenia. The analyzed section is composed of the Bellerophon Formation (evaporite-dolomite member) and the newly introduced Lukae Formation (transitional beds, streaky limestone member and carbonate-clastic member). The Permian-Triassic boundary interval is represented by transitional beds of carbonate facies deposited in shallow restricted marine conditions. The presence of H. parvus in sample L1 in the transitional beds marks the systemic boundary between Permian and Triassic. The studied interval is characterized by a diverse microfauna that contain conodonts, foraminifers, ostracods and gastropods. Six conodont zones have been recognized, in ascending order, the latest Changhsingian (uppermost Permian) praeparvus Zone, and the Griesbachian (lowermost Triassic) parvus, lobata, staeschei- isarcica , postparvus and anceps zones. This faunal succession represents the first known and the most complete conodont biozonation across the Permian-Triassic interval from the entire Dinaric region. The recognized conodont biozones can be correlated with the biozonation of the Southern Alps and of the GSSP Meishan D section.

Evolutionary dynamics of the Permian–Triassic foraminifer size: Evidence for Lilliput effect in the end-Permian mass extinction and its aftermath

There is growing evidence of the Lilliput effect in the end-Permian mass extinction and its aftermath. However, very little has been published on assessing the Lilliput effect on protozoans in the Early Triassic based on quantitative analysis of a great quantity of samples from outcrop sections. This study investigates foraminiferal test size variations through the Changhsingian to the Olenekian based on fossil records from three outcrop sections in South China. The sharp reduction in foraminifer sizes coincides with the first or main episode of the end-Permian mass extinction. Foraminifers remained small in sizes in the early Induan. Their body sizes increased slowly in the late Induan to Olenekian, but never recovered to the pre-extinction test size levels. The Lilliput effect on foraminifera in the aftermath of the end-Permian crisis is indicated by the extinction of large taxa, decrease in size for surviving taxa and rise of small-sized new forms. Physiological reactions of various foraminifer groups reveal that several defaunation events such as anoxia, ocean acidification, and global warming that prevailed through the Permian–Triassic interval may be responsible for both the end-Permian crisis and the Lilliput effect.

No abrupt change in redox condition caused the end-Permian marine ecosystem collapse in the East Greenland Basin

Multiple observations have revealed that environmental disturbances may have been linked to the end-Permian mass extinction and delayed biotic recovery. Biogeochemical constraints on the temporal and spatial changes of oceanic redox chemistry during the Permian–Triassic interval are essential to evaluate global significance of previous hypotheses and to improve our understanding of extinction and recovery processes. To investigate redox ocean chemistry change associated with the end-Permian extinction and subsequent delayed biotic recovery, we examine framboidal pyrites as well as sulfur isotopic compositions of pyrites from the East Greenland Basin. The size distributions of framboidal pyrites in sediments from a continuous section across the Permian–Triassic boundary reveal that sulfidic conditions in water columns were established about 0.7 m above the extinction event in the East Greenland Basin. Our detailed examination of framboidal pyrites challenges a leading hypothesis that euxinia in the photic zone caused the end-Permian ecosystem collapse. We identify several positive and negative S-isotopic shifts before and after the extinction event and demonstrate that a positive S-isotopic shift is not indicative of an abrupt change of redox chemistry in water columns, in contrast to previous claims. The integration of isotope and framboidal pyrite data provides a nearly continuous record of ocean chemistry evolution and new insights into the end-Permian extinction and delayed biotic recovery in the East Greenland Basin.

Timing of the terrestrial Permian-Triassic boundary biotic crisis: Implications from U-Pb dating of authigenic zircons

The Late Permian to Early Triassic transition represents one of the most important Phanerozoic mass extinction episodes. The cause of this event is still in debate between catastrophic and gradual mechanisms. This study uses the U-Pb method on zircons from the uppermost Permian/lowermost Triassic clay deposits at Chahe (Guizhou Province, SW China) to examine time constraints for this event. The results of both this and previous studies show that the ages of Bed 68a and 68c (the upper clay bed of the terrestrial Permian-Triassic boundary (PTB)) respectively are 252.6±2.8 and 247.5±2.8 Ma. This age (within the margin of error) almost accords with the upper clay bed (Bed 28) age of Meishan and the eruption age of Tunguss Basalt, and is so far the most accurate age obtained from terrestrial PTB. The claystone of Bed 68 was formed in the earliest Triassic. The biotic crisis occurred at nearly the same time in terrestrial and marine environments during Permian-Triassic interval; however the extinction patterns and processes are different. The extinction pattern of the terrestrial plants shows a major decline at the PTB after long-term evolution, followed by a retarded extinction of the relicts in the earliest Triassic.

Conodont biostratigraphy across the Permian–Triassic boundary at the Dawen section, Great Bank of Guizhou, Guizhou Province, South China: Implications for the Late Permian extinction and correlation with Meishan

Several continuous Permian–Triassic boundary (PTB) sections are well exposed in the interior of the Great Bank of Guizhou (GBG) on the east limb of the Bianyang syncline, Luodian County, Guizhou Province, South China. Fourteen conodont taxa are identified, including Clarkina kazi, Clarkina lehrmanni n. sp., Clarkina taylorae, Clarkina zhejiangensis, Hindeodus eurypyge, Hindeodus inflatus, Hindeodus sxlatidentatus, Hindeodus parvus erectus, Hindeodus parvus parvus, Hindeodus postparvus, Hindeodus praeparvus, Hindeodus typicalis, Isarcicella staeschei, and Merrillina ultima based on a detailed study of the Permian-Triassic interval at the Dawen section. The first occurrence (FO) of H. parvus parvus in the lower Daye Formation, at about 7.45 m above the contact surface between the Upper Permian skeletal packstone and a calcimicrobial framestone unit, is correlated with the Permian–Triassic boundary; the occurrence of H. eurypyge, H. praeparvus and M. ultima immediately below and H. postparvus above the interval supports this interpretation. A morphometric analysis of 31 Hindeodus specimens helped distinguish H. parvus erectus from H. parvus parvus and H. postparvus. Correlation with the Meishan section (PTB GSSP) using both conodont biostratigraphy and carbon isotopes, indicates that the major extinction at the two localities is simultaneous and coincides with the top of the skeletal packstone at Dawen. The contact between the skeletal packstone and the calcimicrobialite is very irregular and has previously been interpreted as a dissolution surface and correlated with a surface in the lower part of bed 27 at Meishan. Our results confirm this interpretation and reveal that the dissolution event postdated the extinction.

A comparison of the Iberian and Ebro Basins during the Permian and Triassic, eastern Spain: A quantitative subsidence modelling approach

The Permian–Triassic sediments of the Iberian Plate are a well studied case of classical Buntsandstein–Muschelkalk–Keuper facies, with good sedimentological interpretations and precise datings based on pollen and spore assemblages, ammonoids and foraminifera. Synrift–postrift cycles are recorded in these facies, but there are only a few studies of quantitative subsidence analysis (backstripping method) and only a previous one using forward modelling for the quantification of synrift–postrift phases of this period. Here we present the results obtained by the quantitative analysis of fourteen field sections and oil-well electric log records in the Iberian and Ebro Basins, Spain. Backstripping analysis showed five synrift phases of 1 to 3 million years duration followed by postrift periods for the Permian–Triassic interval. The duration, however, shows lateral variations and some of them are absent in the Ebro Basin. The forward modelling analysis, assuming local isostatic compensation, has been applied to each observation point using one-layer and two-layer lithospheric configurations. The second one shows a better fit between observation and model prediction in a systematic way, so a two layer configuration is assumed for the sedimentary basin filling analysis. Lithospheric stretching factors β and δ obtained in the forward modelling analysis are never higher than 1.2, but sometimes β < 1 and simultaneously δ > 1 in the same section. If surficial extension is compensated by deep compression either at the roots of the rift basins or in far-away zones is not yet clear, but this anomaly can be explained using a simple shear extensional model for the Iberian and Ebro basins.

Taxon characteristics that promote survivorship through the Permian–Triassic interval: transition from the Paleozoic to the Mesozoic brachiopod fauna

Examination of organismal characteristics which promote survivorship through both background and mass extinctions may reveal general ecological principles potentially critical to modern conservation efforts. This study explores survivorship of brachiopods, a highly diverse and abundant Paleozoic clade, through the mid-Permian to mid-Triassic interval, which includes the greatest mass extinction in the history of metazoan life. This interval of time separates two of the major Phanerozoic evolutionary faunas. In this regard, survivorship across any one extinction during the interval would not have been relevant if the survivor went extinct shortly after the extinction event; surviving background extinction is as important as surviving a mass extinction. Similarly, taxa that survived but failed to rediversify also were not major elements of the Mesozoic evolutionary fauna. Thus, the analysis aims to analyze survivorship not just across a single extinction but across the entire mid-Permian to mid-Triassic; only survivors through the entire interval can be the ancestors of the Mesozoic clades.Fewer brachiopod genera survived the interval than did brachiopod clades, suggesting that pseudoextinction or insufficient sampling could be a problem in analyzing these extinctions; thus, survivorship analysis should be conducted at the clade level. Nine characteristics were examined for generic representatives of 20 North American brachiopod clades, five of which survived both Permian extinctions and the subsequent earliest Triassic transitional interval. Characteristics include both those that operate on global scales and those that operate on the higher-resolution scales of individuals and populations.Survivors were significantly smaller and occurred less frequently than victims. Mean diversity of communities in which survivors were present was significantly greater. The finding that rare taxa belonging to high-diversity communities were more likely to survive runs counter to traditional predictions. However, these results are consistent with recent studies suggesting that higher diversity within a trophic level may create a buffer, as surviving taxa quickly occupy the vacant niche space of the victims. As size, abundance, and community diversity are all statistically related, the small size of survivors may be an artifact of reduced biovolume per taxon in a diverse community.No significant relationship exists between global-scale processes and survivorship of brachiopods through the mid-Permian to mid-Triassic. The results suggest that ecological processes can strongly influence global extinction patterns.

A LATE PERMIAN TO EARLY TRIASSIC BIVALVE FAUNA FROM THE DONGPAN SECTION, SOUTHERN GUANGXI, SOUTH CHINA

A bivalve fauna from the Permian—Triassic interval of the Dongpan section, Guangxi, South China is described herein. This bivalve fauna includes 16 species belonging to 10 genera. New taxa are Euchondria fusuiensis, Palaeoneilo qinzhouensis, and Claraia liuqiaoensis. Bed 12 of the interval (Talung Formation) contains the typical late Late Permian ammonoids: Huananoceras sp., Laibinoceras cf. L. compressum Yang, and Qiangjiangoceras sp. Bed 13 (Luolou Formation) contains typical Early Triassic bivalves Claraia dieneri Nakazawa, C. cf. C. wangi (Patte), and C. griesbachi (Bittner), which coexist with the earliest Triassic ammonoid Ophiceras sp. The bivalve and ammonoid distributions at the Dongpan section indicate that bed 12 should be assigned to the Late Permian, and bed 13 should be assigned to the Early Triassic. From the early Changhsingian to the Induan, the byssal notches of Claraia species tend to become progressively narrower, and change from being ventrally extended to being horizontally extended. The species level diversity of Claraia also increases through this interval.

Permian marine paleoecology and its implications for large-scale decoupling of brachiopod and bivalve abundance and diversity during the Lopingian (Late Permian)

The transition during the Permian–Triassic interval from brachiopod-rich benthic marine assemblages to communities with diverse molluscs has been well constrained by measures of global taxonomic richness but its ecological context has not been studied using quantitative relative abundance data. New relative abundance data from Early, Middle, and Late Permian silicified fossil assemblages from offshore shelf carbonate environments indicate that molluscs increased significantly in relative abundance, from 0.8% in the Middle Permian (Guadalupian) to 65.4% in the Late Permian (Lopingian). These numerical changes are accompanied by increases in the abundance of infaunal bivalves and motile gastropods in the Late Permian. Although this ecological change was coincident with the end-Guadalupian extinction, a new global diversity compilation indicates that the overall genus-level severity of that biotic crisis was substantially lower than previous estimates. The pronounced ecological shift was instead concurrent with the onset of deep-marine anoxic or euxinic conditions around the Guadalupian–Lopingian boundary, suggesting that increased environmental variability in proximity to the deep water mass may have given eurytopic molluscs a competitive advantage over more stenotopic brachiopods. However, the lack of global changes in relative genus richness implies dramatic decoupling of global taxonomic and local ecological processes. The pronounced increase in molluscan abundance and alpha diversity in the silicified assemblages contrasts with their unchanged global taxonomic richness and may also have resulted from increased environmental stress in offshore shelf environments. These results demonstrate that the ecological transition from the brachiopods to molluscs was more complex than inferred from global measures of taxonomic richness and imply that initial stages of Permo–Triassic environmental deterioration may have been responsible for the decoupling of local ecological from global taxonomic processes during the dramatic Late Permian rise of the Modern Fauna.

Early Triassic recovery of echinoderms

The Permian–Triassic mass extinction interval was an important time in the evolutionary history of the echinoderms. Details of the extinction and, in particular the immediate post-extinction recovery in the Early Triassic, are seldom addressed because of a perception that the Permian–Triassic echinoderm fossil record is too poor. However, only the Holothuroidea and Asteroidea lack any Early Triassic fossil representatives. Even in these groups, details of the extinction and recovery can be inferred from recent cladistic analyses. The Holothuroidea are unique amongst the echinoderms in showing no family level extinction through the Permian–Triassic interval, possibly due to their deposit-feeding lifestyle. In contrast, the Echinoidea, Crinoidea and probably the Asteroidea underwent severe evolutionary bottlenecks during that time. In the echinoids, significant post-Permian radiation occurred from the Late Triassic (Carnian), although it may have begun in the Early Triassic. In the Crinoidea, fossil diversity increases dramatically from the Late Ladinian, although cladistic analyses suggest that initial diversification took place in the Earliest Triassic (Induan). Many undescribed crinoid remains from Lower Triassic strata worldwide also imply that the post-Permian radiation in this group may have been more rapid than currently thought. Locally in the Spathian, crinoid ossicles may approach rock-building densities. The presence of at least seven Early Triassic fossil ophiuroid species may indicate rapid post-Permian radiation in the Ophiuroidea, although the higher level affinities of these taxa are presently unresolved and the Late Permian record is poorly known. Ophiuroid remains are the most diverse echinoderm fossils during the Early Triassic, comprising both complete body fossils and disarticulated ossicles. Holothuroids possibly radiated in the Early Triassic, but current evidence from cladistic analysis favours a largely Anisian age for the post-Permian radiation in this group. All known Early Triassic echinoderms were small-sized animals that inhabited very shallow, oxygenated, low palaeolatitude environments within wave base. To cite this article: R. Twitchett, T. Oji, C. R. Palevol 4 (2005) .