Patchy Landscapes Research Articles

Quantifying the dynamic of cereals and broadleaf plants in semi-arid grasslands using a high-spatial-resolution satellite imaging

The vegetation phenology in pasture grasslands temporarily changes during the growing season due to the different physiological properties of broadleaf and cereal species. Determining their dynamics is required for maximal pasture utilization and sustainability. Widely used low-spatial resolution satellites are suitable for plains but are less effective in patchy landscapes. Therefore, the study's objectives are: (i) to determine the dynamic of cereals and broadleaf vegetation in arid grasslands using high-spatial resolution satellite imaging data, and (ii) to determine the influence of different grazing intensities on grassland vegetation phenology. The study site is in Migda LTER farm, the northern Negev, Israel. Fifty-three grassland plots, sized 10x10m, were located. Seventeen plots were grazed: six heavily (90 min. per plot), five moderately (60 min.), and six lightly (30 min.). A sheep and goats' herd of 120–130 animals grazed on these plots once a year between April to May, from 2019 to 2023. As a reference, fenced, ungrazed plots were located, mainly covered by cereals (Conserved). Imaging data from PlanetScope® was acquired (Spatial resolution: 3 m pixel−1, Temporal: Once in three days, Spectral: RGB and NIR). Using a novel timeline analysis, based on normalizing the different treatments' NDVI by the Conserved values, yielded a three-phased result: a parabolic increase in fresh vegetation reflectance (FVR) from the treatment in the first and third phases, and a parabolic increase of FVR of the Conserved in the second phase. In most grazing treatments, phase duration and maximal NDVI were positively correlated to the rainfall amounts. Rehabilitation trends were calculated for the Lightly grazed, which was correlated to increased broadleaf biomass, while a degradation trend was found for the Heavily grazed. The findings indicate that a parabolic change takes place in vegetation composition in arid grasslands. The novel methodology we used can be adapted and utilized in other ecosystems.

Natural grazing by horses and cattle promotes bird diversity in a restored European alluvial grassland.

A challenge in grassland conservation is to maintain both the openness and the heterogeneity of the habitat to support the diversity of their animal communities, including birds-a taxon that is known to be sensitive to disturbance. An increasingly used management tool in European grassland conservation, especially in rewilding projects, is grazing by large herbivores such as horses and cattle. These grazers are believed to create and maintain patchy landscapes that promote diversity and richness of other species, but their influence on birds is often debated by conservationists, who raise concerns about the impact of disturbance by the grazers. Our aim was to examine the relationship between the abundance and species richness of birds across four foraging guilds and the area utilization patterns of Highland cattle and Konik horses in an alluvial grassland in France. We also aimed to examine the influence of land cover and season on the spatial distribution, including abundance and species richness, of different bird guilds present in the grazed area. We used GPS-collars on all grazers and recorded their positions on an hourly basis over a study period of 1.5 years, assessing patterns of area usage. We counted birds weekly along three transects to describe their distribution within the grazed area and carried out land-cover surveys to describe the habitat. To assess how species richness and abundance of birds of different guilds were related to grazer density, season, and habitat characteristics, we used GAMM models in a spatially explicit framework. We also compared bird numbers at our main study site with a nearby non-grazed control area. The number of birds in the grazed area was about twice the number in the non-grazed control area. Within the grazed area, the abundance of open-area foraging birds increased with increasing grazer density. The number of woodland-foraging birds was also positively correlated with grazer density but less so than open-area foraging birds. The number of individuals in the aerial and wetland bird guilds was not correlated with the density of grazers. Most bird species and individuals were observed on open landscapes scattered with woody patches and waterbodies, and on areas with moderate grazer density. Low-intensity grazing represents a potentially important management tool in creating heterogeneity in alluvial grasslands, thereby promoting suitable habitat for a diverse assemblage of bird species.

Patchy energy landscapes promote stability of small groups of active particles

Group formation and coordination are fundamental characteristics of living matter, essential for performing tasks and ensuring survival. Interactions between individuals play a key role in group formation, and the impact of resource distributions is a vibrant area of research. As of now, an understanding of how patchy resource distributions determine group dynamics is not yet fully understood. Studying active particles in controlled optical landscapes as energy sources, we demonstrate a non-monotonic dependency of group size on landscape patchiness, with the smallest groups forming when the patches match the active particles’ size. A similar relationship is observed in terms of group stability, evidenced by a reduced rate of individual exchange in patchy environments compared to homogeneous conditions. Reduced group sizes can be beneficial to optimise resources in heterogeneous environments and to control information flow within populations. Our results provide insights into the role of patchy landscapes and uneven energy distributions in active matter and hold implications for refining swarm intelligence algorithms, enhancing crowd management techniques, and tailoring colloidal self-assembly.

The influence of resource patch condition and size on social interactions, reproductive behaviors, and oviposition in a fungivorous beetle

Variation among resource patches can structure spatial patterns of social behavior. Individual preferences and requirements for heterogeneous resources can lead to differences in where behaviors take place and which kinds of individuals interact. In this study we examined whether properties of habitat patches predict presence of and interactions among wild forked fungus beetles (Bolitotherus cornutus). B. cornutus use shelf-like fruiting bodies of wood decaying fungi (brackets) as social arenas, courtship and mating sites, oviposition locations, and food sources. Brackets on a single log vary in size and condition and represent a dynamic resource as individual brackets decay over the years that they persist on a subpopulation. We found that size and condition significantly predicted how a bracket was used by B. cornutus. Both males and females were found more often on larger, living brackets. Larger brackets hosted proportionally more social interactions and mating behaviors relative to the overall visitation by individuals. The relationship between these resource properties and frequencies of behaviors was stronger for reproductive, between-sex social interactions. Live brackets also attracted more oviposition events by females, but bracket size had no effect on this behavior. Our results indicate that the dynamics of growth and aging of a critical resource can impact how social and reproductive behaviors are distributed in time and space, which in turn could affect the social structure of subpopulations.Significance statementSpecies that depend on critical host resources are often faced with a patchy landscape of resource quality. Those patches represent the infrastructure within which social behaviors take place, and can have formative effects on where, when, and how social interactions occur. We demonstrated that the size and condition of a fungal resource predicted male and female forked fungus beetle presence and behavior. Larger, living brackets hosted more beetles overall, while larger brackets in any condition hosted more social interactions and mating behaviors. Female beetles were more likely to lay eggs on living brackets, regardless of size. This study demonstrates how ecological dynamics of aging resource patches can indirectly structure social landscapes by influencing where and how individuals interact.

A reaction-diffusion model for population dynamics in patchy landscapes

We address the linear eigenvalue problem associated with reaction-diffusion models for patchy linear domains with any finite number of unique patches. Interface conditions between adjacent patches may be chosen independently to accurately reflect organism behavior. We show that there exists a real principal eigenvalue with a corresponding non-negative eigenfunction that is non-zero interior to the domain. We demonstrate that our model is a generalization of several existing two-patch models. We consider simple three- and four-patch landscapes representing fragmented reserves to illustrate how our model can be used to analyze population persistence, spatial distribution, and migration dynamics. Fragmented landscapes with increased heterogeneity capture migration dynamics not found in landscapes with only two patch types, such as favorable habitat patches that experience net immigration. Conditions for the existence and non-existence of zero-flux conditions interior to three-patch systems are shown, which provide insight to the location of sources and direction(s) of migration.

Predatory Dogs as Drivers of Social Behavior Changes in the Central Himalayan Langur (Semnopithecus schistaceus) in Agro-Forest Landscapes.

Globally, habitat fragmentation has increased the proximity between wildlife, humans, and emerging predators such as free-ranging dogs. In these fragmented landscapes, encounters between primates and dogs are escalating, with primates often falling victim to dog attacks while navigating patchy landscapes and fragmented forests. We aim to investigate how these primates deal with the simultaneous threats posed by humans and predators, specifically focusing on the adaptive strategies of Central Himalayan langur (CHL) in the landscape of fear. To address this, we conducted a behavioral study on the CHL in an agro-forest landscape, studying them for a total of 3912 h over two consecutive years. Our results indicate that, compared to their most common resting behavior, CHLs allocate more time to feeding and locomotion, and less time to socializing in the presence of humans and predatory dogs. Additionally, they exhibit increased feeding and locomotion and reduced social behavior in agro-forest or open habitats. These behavioral patterns reflect adaptive responses to the landscape of fear, where the presence of predators significantly influences their behavior and resource utilization. This study suggests measures to promote coexistence between humans and wildlife through the integration of effective management strategies that incorporate both ecological and social dimensions of human-wildlife interactions.

Regional and local factors interact to shape colonization and extinction dynamics of invasive Hydrilla verticillata in a patchy landscape.

Understanding the response of species to global change requires disentangling the drivers of their distributions across landscapes. Colonization and extinction processes, shaped by the interplay of landscape-level and local patch-level factors, are key determinants of these distributions. However, disentangling the influence of these factors, when larger-scale processes manifest at local scales, remains a challenge. We addressed this challenge by investigating the colonization and extinction dynamics of the aquatic plant, Hydrilla verticillata, in a complex riverine rock pool system. This system, with hundreds of rock pools experiencing varying flooding frequencies, provided a natural laboratory to examine how a single landscape-level disturbance can differentially impact colonization and extinction depending on local patch characteristics to shape species distributions. Using 5 years of data across over 500 sites and more than 5000 surveys, we employed dynamic occupancy models to model colonization, extinction, and changes in Hydrilla patch occupancy while accounting for imperfect detection. Our results revealed that larger, infrequently flooded pools closer to the river were more likely to be colonized. In contrast, local extinction of Hydrilla was more likely in smaller pools closer to the river that flooded frequently. These findings underscore the importance of considering context-dependence in species distribution models. The same landscape-level disturbance (flooding) had opposing effects on colonization and extinction, with the direction and magnitude of these effects varying with local patch characteristics. Our study highlights the need for integrating local and landscape-level factors, and considering how larger-scale processes play out at the patch level, to understand the complex dynamics that shape species distributions.

Trait variation in patchy landscapes: Morphology of spotted salamanders (Ambystoma maculatum) varies more within ponds than between ponds.

The influence of intraspecific trait variation on species interactions makes trait-based approaches critical to understanding eco-evolutionary processes. Because species occupy habitats that are patchily distributed in space, species interactions are influenced not just by the degree of intraspecific trait variation but also the relative proportion of trait variation that occurs within- versus between-patches. Advancement in trait-based ecology hinges on understanding how trait variation is distributed within and between habitat patches across the landscape. We sampled larval spotted salamanders (Ambystoma maculatum) across six spatially discrete ponds to quantify within- and between-pond variation in mass, length, and various metrics associated with their relationship (scaling, body condition, shape). Across all traits, within-pond variation contributed more to total observed morphological variation than between-pond variation. Between-pond variation was not negligible, however, and explained 20-41% of total observed variation in measured traits. Between-pond variation was more pronounced in salamander tail morphology compared to head or body morphology, suggesting that pond-level factors more strongly influence tails than other body parts. We also observed differences in mass-length relationships across ponds, both in terms of scaling slopes and intercepts, though differences in the intercepts were much stronger. Preliminary evidence hinted that newly constructed ponds were a driver of the observed differences in mass-length relationships and morphometrics. General pond-level difference in salamander trait covariation suggest that allometric scaling of morphological traits is context dependent in patchy landscapes. Effects of pond age offer the hypothesis that habitat restoration through pond construction is a driver of variation in trait scaling, which managers may leverage to bolster trait diversity.

Community of Small Mammals in the Patchy Landscape of Northern Kulunda (Western Siberia)

Community of Small Mammals in the Patchy Landscape of Northern Kulunda (Western Siberia)

“Patchiness” in mechanical stiffness across a tumor as an early-stage marker for malignancy

Mechanical phenotyping of tumors, either at an individual cell level or tumor cell population level is gaining traction as a diagnostic tool. However, the extent of diagnostic and prognostic information that can be gained through these measurements is still unclear. In this work, we focus on the heterogeneity in mechanical properties of cells obtained from a single source such as a tissue or tumor as a potential novel biomarker. We believe that this heterogeneity is a conventionally overlooked source of information in mechanical phenotyping data. We use mechanics-based in-silico models of cell-cell interactions and cell population dynamics within 3D environments to probe how heterogeneity in cell mechanics drives tissue and tumor dynamics. Our simulations show that the initial heterogeneity in the mechanical properties of individual cells and the arrangement of these heterogenous sub-populations within the environment can dictate overall cell population dynamics and cause a shift towards the growth of malignant cell phenotypes within healthy tissue environments. The overall heterogeneity in the cellular mechanotype and their spatial distributions is quantified by a “patchiness” index, which is the ratio of the global to local heterogeneity in cell populations. We observe that there exists a threshold value of the patchiness index beyond which an overall healthy population of cells will show a steady shift towards a more malignant phenotype. Based on these results, we propose that the “patchiness” of a tumor or tissue sample, can be an early indicator for malignant transformation and cancer occurrence in benign tumors or healthy tissues. Additionally, we suggest that tissue patchiness, measured either by biochemical or biophysical markers, can become an important metric in predicting tissue health and disease likelihood just as landscape patchiness is an important metric in ecology.

Vegetation mosaic shaping bryophyte diversity in a threatened ecosystem: A case study of Brazilian Cangas (ferruginous rupestrian fields)

The Canga vegetation in Brazil is a unique ecosystem found on ironstone outcrops, known as ferruginous rupestrian fields, and is considered one of the most diverse in the world. This mosaic landscape, comprised of trees, shrubs, and herbaceous vegetation, is home to a diverse range of plants, including angiosperm, bryophytes, and lichens. To understand the effect of Cangas' patchy landscape, here called mesohabitats, on bryophyte communities, we asked the following questions: Do the diversity parameters such as richness, abundance and composition of bryophytes differ along the different Cangas’ mesohabitats? Do bryophyte assemblages group by functional traits and filter per mesohabitat? We surveyed eight sites in the Quadrilátero Ferrífero (Iron Quadrangle), southeast Brazil. We collected and analysed data on bryophyte diversity and functional traits in three different mesohabitats: exposed areas (EA), shrub associations (SA), and tree associ ations (TA). The diversity of both groups was tested using one-way ANOVA; and functional traits were addressed with a Factorial Analysis of Mixed Data (FAMD). There are significant differences in the diversity of bryophytes among the mesohabitats, with TA having the highest diversity and abundance of liverworts and mosses, followed by SA and EA. The study also found bryophyte assemblies with similar functional traits in similar mesohabitats across the Brazilian Quadrilátero Ferrífero. The research revealed that the different mesohabitats provided important and distinct niches for bryophytes in Cangas, and this threatened ecosystem's high diversity must be considered when developing conservation strategies. This entails land managers adopting effective approaches aimed at improving both the quality and connectivity of habitats, fostering biodiversity, and bolstering the resilience of the ecosystem.

Symbiont plasticity as a driver of plant success.

We discuss which plant species are likely to become winners, that is achieve the highest global abundance, in changing landscapes, and whether plant-associated microbes play a determining role. Reduction and fragmentation of natural habitats in historic landscapes have led to the emergence of patchy, hybrid landscapes, and novel landscapes where anthropogenic ecosystems prevail. In patchy landscapes, species with broad niches are favoured. Plasticity in the degree of association with symbiotic microbes may contribute to broader plant niches and optimization of symbiosis costs and benefits, by downregulating symbiosis when it is unnecessary and upregulating it when it is beneficial. Plasticity can also be expressed as the switch from one type of mutualism to another, for example from nutritive to defensive mutualism with increasing soil fertility and the associated increase in parasite load. Upon dispersal, wide mutualistic partner receptivity is another facet of symbiont plasticity that becomes beneficial, because plants are not limited by the availability of specialist partners when arriving at new locations. Thus, under conditions of global change, symbiont plasticity allows plants to optimize the activity of mutualistic relationships, potentially allowing them to become winners by maximizing geographic occupancy and local abundance.

Consistent individual differences in cattle grazing patterns

Cattle grazing patterns can have a significant impact on the economic and environmental sustainability of rangelands and cattle production. Grazing patterns appear to be driven by consistent individual differences in cattle, deemed ‘grazing personalities’. Optimal distribution of cattle on rangelands can be manipulated by alteration of the environment, like adding water, supplement, or fencing to targeted sites, but few studies address how these management tools are impacting grazing patterns of individual cattle. Our study sought to fill this gap in knowledge by investigating consistency of grazing patterns among fifty cattle, fitted with GPS collars, across two years given the addition of a water site at higher elevation prior to year two. We used Bayesian multivariate mixed models to evaluate consistency, evidenced by among-individual correlations across years with confidence intervals that do not cross 0. We found that cattle were consistent in all rangeland use metrics across years after the addition of the new water site: elevation (Estimate = 0.38, CI = [0.07,0.64]), slope (Estimate = 0.59, CI = [0.21,0.86]), distance to water (Estimate = 0.38 CI = [0.03,0.67]), distance to supplement (Estimate = 0.42, CI = [0.04,0.72]), distance to loafing sites (Estimate = 0.68, CI = [0.24,0.95]), except for distance traveled (Estimate = 0.16,CI = [−0.18, 0.48]). Cows that ventured higher on rangeland in year one (Estimate = 0.84 CI = [0.59,0.98]) and year two (Estimate = 0.93 CI = [0.79,0.99]) and further from watering sites in year two (Estimate = 0.72 CI = [0.41,0.92]) also had more within-individual variability in their grazing patterns than those that stayed at lower elevation and closer to water. These results indicate differences in plasticity or variability can be linked to grazing personalities. There was strong evidence that temperature affected grazing distribution such that on hotter days, cows clumped closer to water and loafing sites, were found at lower elevation, and traveled less far, not reaching supplement sites that were placed at the higher elevations. Based on results of this study, ranchers should consider the effectiveness of management tools on individual cattle and conduct cost-benefit analyses of tools implemented for specific behavior types of cattle. Cattle with behavior types demonstrating more within-cow variability and more ‘hill-climbing’ will impact how cattle graze especially patchy landscapes where resources are distributed across elevation, thus ranchers could potentially match cattle types to rangeland characteristics.

Effects of Season, Habitat, and Host Characteristics on Ectoparasites of Wild Rodents in a Mosaic Rural Landscape.

Despite the large number of studies on rodent ectoparasites-most of them vectors of epidemiologically important pathogens-infestation patterns remain poorly understood in various ecological contexts, such as the highly patchy agricultural landscapes. We aimed to relate the infestation of rodents to temporal, habitat, and host variables. We assessed the difference in parasite prevalence and mean abundance depending on host sex, age, and body weight, season, and land use intensity. Furthermore, we analysed the effect of host species abundance and the differential responses of parasites in main and minor host species. The field survey was conducted in a rural landscape in southern Transylvania (Romania) between June and September 2010-2011. We live-trapped small mammals, collected the ticks and fleas, and recorded the presence of lice and mites. Overall, we found the same infestation patterns largely reported in the literature: higher prevalence and mean abundance in heavier adult males, significant seasonality and differences among host species, and evidence of the dilution effect. The uniqueness of our study system was the negative effect of the land use intensity on the prevalence and mean abundance of parasites, explained by the highly patchy mosaic landscape.

The patchy landscape of supervision for child protection professionals in Albania

ABSTRACT To provide a snapshot of the state of child protection supervision in Albania, this article presents findings from a small-scale study undertaken to explore the views and experiences of professionals in multidisciplinary child protection teams. Our aims were to provide a description of current supervisory practice and explore perceptions and experiences of supervision. Data were collected via five key informant interviews and surveys completed by 17 frontline workers. We found a patchy landscape of understanding and provision of supervision in Albania.

Uneven distributions of unique species promoting N niche complementarity explain the stability of degraded alpine meadow

Alpine meadow degradation, usually involving decreased soil nitrogen (N) and patchy landscapes, is a challenge for natural restoration. However, the mechanism underlying plant species coexistence under degradation is unclear. In this study, we evaluated plant N niche complementarity in degraded alpine meadows on the Qinghai–Tibet Plateau using a 15N-labeling (15NO3−, 15NH4+, and 15N-glycine) experiment. At the community level, the concentration of 15NO3− in the degraded alpine meadow was 1.5 times higher than that in the undegraded alpine meadow; both alpine meadows had a significant preference for NO3− (60.72 % and 66.84 % for the degraded and undegraded alpine meadows, respectively), and the degree of glycine preference was significantly higher in the degraded alpine meadow (30.77 %) relative to the undegraded alpine meadow (21.85 %). At the species level, dominant species in both alpine meadows consistently preferred NO3−; the generalist species that can be found in both meadows and unique species of the two alpine meadows generally showed NO3− preferences, while the other plant species that were unevenly distributed in the degraded alpine meadow tended to show increased utilization of glycine, which could reduce N competition. We observed that differentiation among N sources and the uneven distribution of unique species may explain the stability of degraded alpine meadows. Our results suggested that uneven distributions of plants could have strong impacts on community stability and highlighted the importance of considering fine-scale analysis in studies of niche theory. This study has important implications for the restoration of degraded alpine meadows.

Patchy indirect effects of predation: predators contribute to landscape heterogeneity and ecosystem function via localized pathways

Predators are widely recognized for their irreplaceable roles in influencing the abundance and traits of lower trophic levels. Predators also have irreplaceable roles in shaping community interactions and ecological processes via highly localized pathways (i.e. effects with well‐defined and measurable spatio–temporal boundaries), irrespective of their influence on prey density or behavior. We synthesized empirical and theoretical research describing how predators – particularly medium‐ and large‐sized carnivores – have indirect ecological effects confined to discrete landscape patches, processes we have termed ‘patchy indirect effects (PIEs) of predation'. Predators generate PIEs via three main localized pathways: generating and distributing prey carcasses, creating ecological hotspots by concentrating nutrients derived from prey, and killing ecosystem engineers that create patches. In each pathway, the indirect effects are limited to discrete areas with measurable spatial and temporal boundaries (i.e. patches). Our synthesis reveals the diverse and complex ways that predators indirectly affect other species via patches, ranging from mediating scavenger interactions to influencing parasite/disease transmission risk, and from altering ecosystem biogeochemistry to facilitating local biodiversity. We provide basic guidelines on how these effects can be quantified at the patch and landscape scales, and discuss how predator‐mediated patches ultimately contribute to landscape heterogeneity and ecosystem functioning. Whereas density‐ and trait‐mediated indirect effects of predation generally occur through population‐scale changes, PIEs of predation occur through individual‐ and patch‐level pathways. Our synthesis provides a more holistic view of the functional role of predation in ecosystems by addressing how predators create patchy landscapes via localized pathways, in addition to influencing the abundance and behavior of lower trophic levels.

Late Middle Pleistocene sequences in the lower Ob’ and Irtysh (West Siberia) and new multi-proxy records of terrestrial environmental change

The Middle Pleistocene environmental history of the north of West Siberia is very incomplete. Only a few poorly preserved paleoarchives are known so far in this huge part of Eurasia encompassing up to one million square kilometers. Here we present new data (micromammals, insects, plant macrofossils and luminescence dating) from the lower reaches of the Ob’ and Irtysh Rivers. We reconstruct the circumstances of the formation of the fossil-bearing sites Bolshaya Ob 440 km – Khashgort (BOB440) and Gornopravdinsk 2 (GP2) and compare the results with the current state of knowledge of the environmental change in this area. The combined evidence suggests that the fossil-bearing unit known as the Khashgort diagonal sands at BOB440 yields 1) a late Middle Pleistocene complex of fauna and flora of a cold terrestrial environment with tundra-like vegetation, 2) a Middle Pleistocene flora of interglacial environment with temperate vegetation, and 3) reworked fossils form the Late Cretaceous – Oligocene. Geochronometric data suggest that the Khashgort sand unit at BOB440 might have been formed during the Middle/Late Pleistocene transition. However, because of a high variation in results obtained on repeated samples from the same strata, some uncertainty of the luminescence dating remains, which requires further verification. The fossil-bearing fluvial sequence at GP2 represents the lower part of the Chembakchino Formation and encompasses the lower part of the Tobolian Horizon (correlated with Holsteinian). The micromammal complex comprises Dicrostonyx simplicior (intermediate morphological stage S2 sensu Smirnov et al., 1997), Lasiopodomys gregalis, Microtus nivaloides lidiae and other taxa, which are characteristic of the early evolutionary stage of late Middle Pleistocene faunas in the north of West Siberia. The paleoecological analysis of plant and animal remains at GP2 suggests an environmental change from cold, tundra-like landscapes at the Shaitanian/Tobolian transition (MIS 12 – MIS 11) to non-analogous open patchy landscapes with highly productive moss and herbaceous vegetation during the early Tobolian time.

Undesigned World: what educating through walking means in the Anthropocene

For this essay, we draw from a collaborative experience between two educators and thirty students in a course called Designed World, taught at a public university, to explore how attention and intention develop within the practice of walking across “patchy landscapes” (Tsing 2015, 20) or the anthropocentric landscape, and how they can assist thinking about the design of “possible worlds” (Haraway 1991). This essay adopts an interdisciplinary approach that integrates ecofeminist theory with ideas of Romantic empiricism and relates them to aesthetic education and theories about walking. This journey to new places of learning – beyond the indoor structure of the classroom or studio – begins with becoming attentive to the complexity around us. Attention is a lack of intention (Masschelein 2010, 48). In contrast to most design activities that begin with an intent to produce a desired outcome, and a rich methodology, such as Design Thinking, there is no method here and no sequence to follow. Instead, the goal is to allow the present to be sublime to us so that we open ourselves up in a vulnerable way to being changed by that through which we move. This form of teaching helps us practise attention, presence, and responsibility. We do not expect anything in return. The simple act of walking can help us navigate an indeterminate time and place and create a social art and commons.

Food webs coupled in space: Consumer foraging movement affects both stocks and fluxes.

The exchange of material and individuals between neighboring food webs is ubiquitous and affects ecosystem functioning. Here, we explore animal foraging movement between adjacent, heterogeneous habitats and its effect on a suite of interconnected ecosystem functions. Combining dynamic food web models with nutrient-recycling models, we study foraging across habitats that differ in fertility and plant diversity. We found that net foraging movement flowed from high to low fertility or high to low diversity and boosted stocks and flows across the whole loop of ecosystem functions, including biomass, detritus, and nutrients, in the recipient habitat. Contrary to common assumptions, however, the largest flows were often between the highest and intermediate fertility habitats rather than highest and lowest. The effect of consumer influx on ecosystem functions was similar to the effect of increasing fertility. Unlike fertility, however, consumer influx caused a shift toward highly predator-dominated biomass distributions, especially in habitats that were unable to support predators in the absence of consumer foraging. This shift resulted from both direct and indirect effects propagated through the interconnected ecosystem functions. Only by considering both stocks and fluxes across the whole loop of ecosystem functions do we uncover the mechanisms driving our results. In conclusion, the outcome of animal foraging movements will differ from that of dispersal and diffusion. Together we show how considering active types of animal movement and the interconnectedness of ecosystem functions can aid our understanding of the patchy landscapes of the Anthropocene.