Pasture-based Milk Production Systems Research Articles

Productivity, Profitability and Nitrogen Utilisation Efficiency of Two Pasture-Based Milk Production Systems Differing in the Milking Frequency and Feeding Level

The aim of this study was to model the productivity, profitability and the nitrogen (N) utilisation efficiency (NUE) of two spring-calving pasture-based milk production systems differing in milking frequency and intensification levels in New Zealand. For this purpose, physical performance data from a low-intensity production system where cows were milked once per day (OAD-LI) and from a high-intensity production system where cows were milked twice per day (TAD-HI) were employed. OAD-LI cows were milked once-daily with a stocking rate (SR) of 2.1 cows/ha and fed diets with low supplementation (304 kg pasture silage/cow) with applications of 134 kg N fertiliser/ha and TAD-HI cows were milked twice-daily with a SR of 2.8 cows/ha and fed diets of higher supplementation (429 kg pasture silage and 1695 kg concentrate/cow) with applications of 87 kg N fertiliser/ha. The Moorepark Dairy System Model was used to evaluate production, economic performance and N balance on an annual basis. Despite the higher feed costs of TAD-HI as more supplementation was utilised, profitability per hectare was 16% higher because more cows were milked with a higher milk yield per cow (milking frequency) when compared to OAD-LI. At the cow level, the NUE was higher in TAD-HI (30% vs. 27%) reflecting the better balanced diet for energy and crude protein and higher milk yields as a result of milking frequency. At the farm scale the NUE was higher (38% vs. 26%) in the TAD-HI due to the losses associated with the imported feed being excluded and higher N captured in milk. These results suggest that milking frequency, the use of feed supplementation and application of N fertiliser as management tools on grazing dairy systems affect productivity, profitability and N balance. Further studies are required to find optimal stocking rates in combination with the use of supplementary feed and N fertiliser application that maximize milk production and profitability for OAD and TAD milking production systems but minimize N losses.

Nitrogen use efficiency, reticulo-ruminal pH and behaviour of lactating cows grazing either in a full-time system or in a part-time system with indoor feeding of fresh herbage and concentrate

AbstractIncreasing herd size and fragmented or non-uniform land challenge pasture-based milk production systems. Therefore, part-time grazing combined with indoor feeding of fresh herbage (‘cut-and-carry’) allows to maintain the advantages of fresh-herbage-based diets. However, consequences for environmental nitrogen losses, animal behaviour and ruminal metabolism are decisive for its implementation. Animal performance, nitrogen use efficiency (NUE), reticulo-ruminal pH and feeding/locomotive behaviour were compared in six cows subjected either to full-time grazing (system F) or to part-time grazing system with indoor feeding of fresh herbage and concentrate (system P). Biweekly samplings were performed on six lactating cows per treatment in spring, summer and autumn. Nitrogen (N) balance was calculated from estimated N intake and N output via faeces, urine and milk. Reticulo-ruminal pH was recorded using a wireless data recorder. Eating and locomotive behaviour were measured with noseband sensors and pedometers. In spring, but not later, herbage intake per unit of body weight was higher for system F compared to system P cows. Eating time was shorter by 15% in P compared to F cows. Across systems, NUE declined from spring to summer/autumn from about 300 to 200 g milk protein N/kg N intake. Reticulo-ruminal pH was more variable under part-time grazing conditions, with a pronounced decrease subsequent to offering the herbage indoors. The part-time grazing system resulted in similar or higher NUE than full-time grazing. Additional studies with larger numbers of experimental units are required to be able to develop comprehensive recommendations for the improvement of the two grazing systems.

Carbon Footprint of Milk from Pasture-Based Dairy Farms in Azores, Portugal

The dairy sector is a major contributor to greenhouse gas emissions. Pasture-based dairy production is sometimes credited as environmentally friendlier but is less studied than more intensive production systems. Here we characterize and calculate the carbon footprint (CF), using life cycle assessment, of the “Vacas Felizes” pasture-based milk production system, in the Azores archipelago. Impacts were determined for multiple functional units: mass, energy and nutritional content, farm, area and animal. We performed multivariate analysis to assess the contribution of production parameters to the CF. Finally, we performed a literature review to compare these results with other production systems. Results show that emissions from enteric fermentation, concentrated feed production and (organic and mineral) fertilizer application are the three main sources of impact. Milk yield is a key production feature for the determination of emissions. The average CF is 0.83 kg CO2/kg raw milk. At each milk yield level, the farms are approximately homogeneous. Compared with other studies, “Vacas Felizes” milk has a lower CF than 80 (out of 84) published CFs and on average it is approximately 32% lower.

Factors associated with profitability in pasture-based systems of milk production

The global dairy industry needs to reappraise the systems of milk production that are operated at farm level with specific focus on enhancing technical efficiency and competitiveness of the sector. The objective of this study was to quantify the factors associated with costs of production, profitability, and pasture use, and the effects of pasture use on financial performance of dairy farms using an internationally recognized representative database over an 8-yr period (2008 to 2015) on pasture-based systems. To examine the associated effects of several farm system and management variables on specific performance measures, a series of multiple regression models were developed. Factors evaluated included pasture use [kg of dry matter/ha and stocking rate (livestock units/ha)], grazing season length, breeding season length, milk recording, herd size, dairy farm size (ha), farmer age, discussion group membership, proportion of purchased feed, protein %, fat %, kg of milk fat and protein per cow, kg of milk fat and protein per hectare, and capital investment in machinery, livestock, and buildings. Multiple regression analysis demonstrated costs of production per hectare differed by year, geographical location, soil type, level of pasture use, proportion of purchased feed, protein %, kg of fat and protein per cow, dairy farm size, breeding season length, and capital investment in machinery, livestock, and buildings per cow. The results of the analysis revealed that farm net profit per hectare was associated with pasture use per hectare, year, location, soil type, grazing season length, proportion of purchased feed, protein %, kg of fat and protein per cow, dairy farm size, and capital investment in machinery and buildings per cow. Pasture use per hectare was associated with year, location, soil type, stocking rate, dairy farm size, fat %, protein %, kg of fat and protein per cow, farmer age, capital investment in machinery and buildings per cow, breeding season length, and discussion group membership. On average, over the 8-yr period, each additional tonne of pasture dry matter used increased gross profit by €278 and net profit by €173 on dairy farms. Conversely, a 10% increase in the proportion of purchased feed in the diet resulted in a reduction in net profit per hectare by €97 and net profit by €207 per tonne of fat and protein. Results from this study, albeit in a quota limited environment, have demonstrated that the profitability of pasture-based dairy systems is significantly associated with the proportion of pasture used at the farm level, being cognizant of the levels of purchased feed.

Effect of stocking rate and animal genotype on dry matter intake, milk production, body weight, and body condition score in spring-calving, grass-fed dairy cows.

The objective of the experiment was to quantify the effect of stocking rate (SR) and animal genotype on milk production, dry matter intake (DMI), energy balance, and production efficiency across 2 consecutive grazing seasons (2014 and 2015). A total of 753 records from 177 dairy cows were available for analysis: 68 Holstein-Friesian and 71 Jersey × Holstein-Friesian (JxHF) cows each year of the experiment under a pasture-based seasonal production system. Animals within each breed group were randomly allocated to 1 of 3 whole-farm SR treatments defined in terms of body weight per hectare (kg of body weight/ha): low (1,200 kg of body weight/ha), medium (1,400 kg of body weight/ha), and high (1,600 kg of body weight/ha), and animals remained in the same SR treatments for the duration of the experiment. Individual animal DMI was estimated 3 times per year at grass using the n-alkane technique: March (spring), June (summer), and September (autumn), corresponding to 45, 111, and 209 d in milk, respectively. The effects of SR, animal genotype, season, and their interactions were analyzed using mixed models. Milk production, body weight, and production efficiency per cow decreased significantly as SR increased due to reduced herbage availability per cow and increased grazing severity. As a percentage of body weight, JxHF cows had higher feed conversion efficiency, higher DMI and milk solids (i.e., kg of fat + kg of protein) production, and also required less energy intake to produce 1 kg of milk solids. The increased production efficiency of JxHF cows at a similar body weight per hectare in the current analysis suggests that factors other than individual cow body weight contribute to the improved efficiency within intensive grazing systems. The results highlight the superior productive efficiency of high genetic potential crossbred dairy cows within intensive pasture-based milk production systems at higher SR where feed availability is restricted.

Evaluation of the RumiWatchSystem for measuring grazing behaviour of cows

Feeding behaviour is an important parameter of animal performance, health and welfare, as well as reflecting levels and quality of feed available. Previously, sensors were only used for measuring animal feeding behaviour in indoor housing systems. However, sensors such as the RumiWatchSystem can also monitor such behaviour continuously in pasture-based environments. Therefore, the aim of this study was to validate the RumiWatchSystem to record cow activity and feeding behaviour in a pasture-based system. The RumiWatchSystem was evaluated against visual observation across two different experiments. The time duration per hour at grazing, rumination, walking, standing and lying recorded by the RumiWatchSystem was compared to the visual observation data in Experiment 1. Concordance Correlation Coefficient (CCC) values of CCC=0.96 for grazing, CCC=0.99 for rumination, CCC=1.00 for standing and lying and CCC=0.92 for walking were obtained. The number of grazing and rumination bouts within one hour were also analysed resulting in Cohen‘s Kappa (κ)=0.62 and κ=0.86 for grazing and rumination bouts, respectively. Experiment 2 focused on the validation of grazing bites and rumination chews. The accordance between visual observation and automated measurement by the RumiWatchSystem was high with CCC=0.78 and CCC=0.94 for grazing bites and rumination chews, respectively. These results indicate that the RumiWatchSystem is a reliable sensor technology for observing cow activity and feeding behaviour in a pasture based milk production system, and may be used for research purposes in a grazing environment.

Life cycle environmental impacts of high and low intensification pasture-based milk production systems: A case study of the Waikato region, New Zealand

Intensification of pasture-based dairy farming systems is usually associated with increased use of off-farm inputs to increase milk production per hectare, and additional environmental impacts may be associated with upstream production and delivery of these off-farm inputs. The objective of this study was to compare two levels of dairy farming intensification (i.e. high versus low) in pasture-based milk production systems with respect to (i) farm production and (ii) life cycle environmental indicators. The study considered 53 dairy farms in the Waikato region, New Zealand. Three farm attributes (stocking rate, amount of brought-in feeds and amount of nitrogen fertilizer used) were used to define a level of farming intensification (Intensification Index) for each dairy farm. The upper and lower quartiles of the dairy farms ranked on their Intensification Indices were chosen to represent high and low intensification groups, respectively. Twelve midpoint environmental indicators of the dairy systems were assessed using attributional Life Cycle Assessment with one kg of fat- and protein-corrected milk as a functional unit and a cradle-to-farm gate perspective as a system boundary. Compared with the low intensification group, the high intensification group had (on a per hectare basis) higher (P < 0.001) stocking rate, total brought-in feeds and nitrogen fertilizer use, which led to greater milk yield per cow (P < 0.01) and per hectare (P < 0.001). The different levels of farming intensification did not affect total feed conversion efficiency (P > 0.05). However, for the high intensification group, the results for 10 out of 12 environmental indicators per kg of fat- and protein-corrected milk were higher (P < 0.05) than those in the low intensification group. The main drivers for the increases in most environmental indicator results were the production of brought-in feeds, manufacturing of agrichemicals and transportation of off-farm inputs for use on dairy farms. In contrast, increased pasture intake was negatively correlated (P < 0.05) with all environmental impacts, indicating that efficient pasture management is also critical to mitigate environmental impacts. In conclusion, while an increase in intensification of pasture-based dairy farming systems led to increased milk production per cow and per hectare, it also resulted in increased environmental impacts for most indicators. Apart from increased resource use efficiency, increased pasture utilization efficiency is a promising measure to improve environmental sustainability of pasture-based dairy farming systems.

Effect of stocking rate on milk and pasture productivity and supplementary feed use for spring calving pasture fed dairy systems

The productivity of grazing systems is primarily limited by the scale and efficiency of systems applied to the grazable land platform adjacent to the milking parlor. The objective of this study was to compare forage production, utilization and quality, milk production, and requirement for supplementary feeds for 2 different grazing platform stocking rate (GPSR) treatments over 4 yr. Animals were randomly allocated to 1 of 2 GPSR treatments: high-closed (HC; 3.1 cows/ha) and high-open (HO; 4.5 cows/ha), which were designed to represent alternative GPSR in a post-European Union milk quota, spring calving, pasture-based milk production system. Animal production data were analyzed using Proc MIXED of SAS with GPSR, year, and parity included as fixed effects in the final model. Within a seasonal spring calving grazing system, at high GPSR and offering moderate amounts of additional supplements based on pasture supply deficits, both systems produced more milk and fat plus protein per hectare in comparison with Irish commercial dairy farms. Although requiring additional supplementation, increased GPSR resulted in increased milk production per hectare but also in an increased requirement for concentrate and forage supplementation during lactation. No significant influence of GPSR was found on body weight and body condition score or reproductive performance during the 4-yr study period. In addition, GPSR also had no effect on pasture production, utilization, or quality during the study period. The strategic use of additional supplements with restricted pasture availability at higher GPSR maintained milk production per cow and significantly increased milk production per hectare.

The multi-year cumulative effects of alternative stocking rate and grazing management practices on pasture productivity and utilization efficiency

The production and utilization of increased quantities of high quality pasture is of paramount importance in pasture-based milk production systems. The objective of this study was to evaluate the cumulative effects of alternative integrated grazing strategies, incorporating alternative stocking rate (SR) and grazing severities, on pasture productivity and grazing efficiency over multiple years within farm systems using perennial ryegrass dominant pastures. Three whole-farm SR treatments were compared over 4 complete grazing seasons (2009 to 2012 inclusive): low (2.51 cows/ha; LSR), medium (2.92 cows/ha; MSR), and high (3.28 cows/ha; HSR). Each system had its own farmlet containing 18 paddocks and remained on the same treatment for the duration of the study. Stocking rate had a significant effect on all grazing variables with the exception of soil fertility status and sward density. Increased SR resulted in increased total annual net pasture accumulation, improved sward nutritive value, and increased grazed pasture utilization. Total annual net pasture accumulation was greatest in HSR [15,410kg of dry matter (DM)/ha], intermediate for MSR (14,992kg of DM/ha), and least for LSR (14,479kg of DM/ha) during the 4-yr study period. A linear effect of SR on net pasture accumulation was detected with an increase in net pasture accumulation of 1,164.4 (SE=432.7) kg of DM/ha for each 1 cow/ha increase in SR. Pregrazing pasture mass and height and postgrazing residual pasture mass and height were greatest for LSR, intermediate for the MSR, and lowest for the HSR. In comparison with the LSR, the imposition of a consistently increased grazing severity coupled with increased whole farm SR in MSR and HSR treatments arrested the decline in sward nutritive value, typically observed during mid-season. Incorporating the individual beneficial effects of SR on pasture accumulation, nutritive value, and utilization efficiency, total proportional energy (unité fourragère lait) utilization per hectare increased significantly with increasing SR (+0.026 and +0.081 for MSR and HSR, respectively). These results quantify the significant effect of grazing management practices on the feed production capability of modern perennial ryegrass pastures for intensive grazing dairy production systems. Furthermore, these results highlight the importance of consistently imposing grazing treatments over multiple years, and within integrated whole farm systems, to accurately assess the longer term effects of alternate grazing management practices on pasture productivity.

The energy expenditure of 2 Holstein cow strains in an organic grazing system

Until recently, measurements of energy expenditure (EE; herein defined as heat production) in respiration chambers did not account for the extra energy requirements of grazing dairy cows on pasture. As energy is first limiting in most pasture-based milk production systems, its efficient use is important. Therefore, the aim of the present study was to compare EE, which can be affected by differences in body weight (BW), body composition, grazing behavior, physical activity, and milk production level, in 2 Holstein cow strains. Twelve Swiss Holstein-Friesian (HCH; 616kg of BW) and 12 New Zealand Holstein-Friesian (HNZ; 570kg of BW) cows in the third stage of lactation were paired according to their stage of lactation and kept in a rotational, full-time grazing system without concentrate supplementation. After adaption, the daily milk yield, grass intake using the alkane double-indicator technique, nutrient digestibility, physical activity, and grazing behavior recorded by an automatic jaw movement recorder were investigated over 7d. Using the 13C bicarbonate dilution technique in combination with an automatic blood sampling system, EE based on measured carbon dioxide production was determined in 1 cow pair per day between 0800 to 1400h. The HCH were heavier and had a lower body condition score compared with HNZ, but the difference in BW was smaller compared with former studies. Milk production, grass intake, and nutrient digestibility did not differ between the 2 cow strains, but HCH grazed for a longer time during the 6-h measurement period and performed more grazing mastication compared with the HNZ. No difference was found between the 2 cow strains with regard to EE (291±15.6kJ) per kilogram of metabolic BW, mainly due to a high between-animal variation in EE. As efficiency and energy use are important in sustainable, pasture-based, organic milk production systems, the determining factors for EE, such as methodology, genetics, physical activity, grazing behavior, and pasture quality, should be investigated and quantified in more detail in future studies.

Synchronisation of ovulation for management of reproduction in dairy cows

Important developments have occurred in the last two decades, since the advent of the Ovsynch protocol, on the understanding and use of synchronisation programmes for management of reproduction in dairy herds. This better understanding of oestrus cycle control associated with suboptimal reproductive performance in dairy herds has led dairy producers to quickly adopt timed artificial insemination (AI) protocols. Recent surveys have documented that fixed-time AI has become an important component of management of reproduction in high-producing herds. Furthermore, timed AI protocols have also demonstrated benefits in pasture-based milk production systems because of the ability to increase insemination rate. In general, successful use of the Ovsynch protocol requires some fundamental physiological principles to be respected, including: induction of ovulation to synchronise follicle growth in the first 2 days of the programme such that a young antral follicle is recruited; maintenance of high concentrations of progesterone during the development of the ovulatory follicle, but also effectively lyse the corpus luteum to result in very low concentration of progesterone at AI; and having a healthy pre-ovulatory follicle of moderate diameter that is highly oestrogenic and responsive to gonadotropins to synchronously ovulate 12 to 18 h after insemination. Current methods oestrous and ovulation synchronisation are still not optimal and future improvements will likely require new technologies for hormone formulation and delivery such that additional interventions are minimised to maintain acceptance by producers.

Challenges in ration formulation in pasture-based milk production systems

The profitability of temperate dairy systems in southern Australia and New Zealand is underpinned by the efficient use of the home-grown feed-base and purchased supplements. While there are well established principles on which to devise nutritionally balanced rations for ruminants, it is evident that in grazing systems the variable supply of nutrients from forage often results in suboptimal performance compared with predicted responses. This review discusses the key factors likely to directly affect responses to supplementation in grazing systems, namely pasture supply, nutritive characteristics and intake, feeding systems, substitution, associative effects and nutrient synchrony. Intake is the primary driver for metabolisable energy (ME) and nutrient supply and uncertainties on the actual ME and nutrients supplied by pasture will remain due to variability within and between pasture types and within paddocks. It is suggested that improvements in ration formulation will require a better understanding of the kinetics of intake and digestion within the rumen and of nutrients supplied to the tissues from grazed pasture with and without supplements. Research which clearly elucidates which mechanism or mechanisms determine substitution rates with concentrate and forage supplements would be valuable. Such knowledge could also be used to develop feeding strategies that result in a reduction in the degree of substitution and, in turn, lead to higher DM and ME intake compared with existing feeding practices. This in turn would lead to a better understanding of associative effects and nutrient synchrony with benefits in better design of rations and feed-use efficiency.

Suitability of different dairy cow types for an Alpine organic and low-input milk production system

The implementation of a seasonal, site adapted, pasture-based milk production system as applied in New Zealand and North Western Europe might be an alternative for the near future in Alpine regions. As in such a low-input system the fertility and secondary traits such as robustness against metabolic challenges of dairy cows is of crucial importance, it is questionable whether conventional dairy cow types, which were selected primarily for milk production under high-input conditions, are most suitable. Therefore the aim of this study was to compare two different genotypes concerning their suitability for an organic, low-input milk production system under Alpine conditions. Between 2008 and 2011 records from 91 lactations of Brown Swiss (BS) and a special strain of Holstein Friesian (HFL) were collected at an experimental organic dairy farm. The BS cows represented the average Austrian BS population and were primarily selected for high milk yield. HFL cows were for decades selected under low-input conditions and were primarily bred for lifetime performance and fitness. The dataset was analysed using multi-factorial statistical models. BS was superior for most milk production parameters, but not for milk yield per unit of metabolic body weight. HFL had a lower body weight throughout the lactation, but mobilised significantly less body reserves, indicating a shorter and less negative energy balance. Both breeds had similar total feed and energy intake, but HFL had a significantly higher feed intake per unit of metabolic body weight. Furthermore, HFL conceived earlier and had a significantly shorter calving interval and an overall superior reproductive performance. Due to this, HFL has some advantages over conventional BS if managed in a pasture based, low-input milk production system, especially when block calving is involved.

A biological and economic comparison of 2 pasture-based production systems on a wetland drumlin soil in the northern region of Ireland

The objective of this study was to compare the biological and economic efficiencies of 2 likely future pasture-based systems of milk production differing in overall stocking rate and concentrate supplementation level on a wetland drumlin soil in the Border Midlands Western region of Ireland. Physical performance data were obtained from a 3-yr systems comparison at Ballyhaise College, Co. Cavan, comparing 2 production systems: a high grass (HG) system (578kg of concentrate/cow at 2.45 livestock units per hectare) and a high intensity (HI) system (1,365kg of concentrate/cow at 2.92 livestock units/ha). Animal production data were analyzed using a mixed model, with feed system, year, and parity included as fixed effects in the final model. Feed system had a significant effect on all yield variables with higher yields in the HI system. Production system had no significant influence on reproductive performance. The Moorepark Dairy Systems Model, a stochastic budgetary simulation model, was used to simulate a model farm integrating biological data from each feed system to identify the economic effect of each system at 3 future milk prices of 22, 27, and 33 euro cents per liter (€c/L). Two economic scenarios were investigated within the model: scenario 1 (S1) assumed fixed cow numbers (n=55 cows) and scenario 2 (S2) assumed fixed land area (n=40ha). At a milk price of 27 or 33 €c/L, profit per cow, per kilogram of milk solids, and per hectare were similar for HG and HI in S1 and higher for HI in S2. At a milk price of 22 €c/L, all systems were unprofitable, with increased losses realized in the HI system (both S1 and S2) compared with the HG system. Pasture-based systems of milk production in the northern region of Ireland are capable of highly efficient and profitable milk production. Moreover, the efficacy of increased supplementation to remove the constraints of pasture seasonality will depend on the cost of supplementation and the price paid for additional milk produced.

Meta-analysis of the impact of stocking rate on the productivity of pasture-based milk production systems

The objective of this study is to quantify the milk production response per cow and per hectare (ha) for an incremental stocking rate (SR) change, based on a meta-analysis of published research papers. Suitable experiments for inclusion in the database required a comparison of at least two SRs under the same experimental conditions in addition to details on experimental length and milk production results per cow and per ha. Each additional increased SR treatment was also described in terms of the relative milk production change per cow and per ha compared to the lower base SR (b_SR). A database containing 109 experiments of various lengths with 131 comparisons of SR was sub-divided into Type I experiments (common experimental lengths) and Type II experiments (variable experimental lengths). Actual and proportional changes in milk production according to SR change were analysed using linear mixed model procedures with study included as a random effect in the model. Low residual standard errors indicated a good precision of the predictive equations with the exception of proportional change in milk production per cow. For all milk yield variables analysed, the results illustrate that while production per cow is reduced, a strong positive relationship exists between SR and milk production per ha. An SR increase of one cow/ha resulted in a decrease in daily milk yield per cow of 7.4% and 8.7% for Type I and Type II data, respectively, whereas milk yield per ha increased by 20.1% and 19.6%, respectively. Within the Type II data set, a one cow/ha increase in SR also resulted in a 15.1% reduction in lactation length (equivalent to 42 days). The low predictability of proportional change in milk production per cow according to the classical SR definition of cows per ha over a defined period suggests that SR may be more appropriately defined in terms of the change in available feed offered per animal within each treatment.

Dry matter intake and feed efficiency profiles of 3 genotypes of Holstein-Friesian within pasture-based systems of milk production

The primary objective of the study was to quantify the effect of genetic improvement using the Irish total merit index (Economic Breeding Index) on dry matter intake and feed efficiency across lactation and to quantify the variation in performance among alternative definitions of feed efficiency. Three genotypes of Holstein-Friesian dairy cattle were established from within the Moorepark dairy research herd: 1) low Economic Breeding Index North American Holstein-Friesian representative of the Irish national average dairy cow, 2) high genetic merit North American Holstein-Friesian, and 3) high genetic merit New Zealand Holstein-Friesian. Animals from within each genotype were randomly allocated to 1 of 2 possible intensive pasture-based feed systems: 1) the Moorepark pasture system (2.64 cows/ha and 500kg of concentrate supplement per cow per lactation) and 2) a high output per hectare pasture system (2.85 cows/ha and 1,200kg of concentrate supplement per cow per lactation). A total of 128 and 140 spring-calving dairy cows were used during the years 2007 and 2008, respectively. Each group had an individual farmlet of 17 paddocks, and all groups were managed similarly throughout the study. The effects of genotype, feed system, and the interaction between genotype and feed system on dry matter intake, milk production, body weight, body condition score, and different definitions of feed efficiency were studied using mixed models with factorial arrangements of genotypes and feed systems accounting for the repeated cow records across years. No significant genotype-by-feed-system interactions were observed for any of the variables measured. Results showed that aggressive selection using the Irish Economic Breeding Index had no effect on dry matter intake across lactation when managed on intensive pasture-based systems of milk production, although the ranking of genotypes for feed efficiency differed depending on the definition of feed efficiency used. Performance of animals grouped on alternative definitions of feed efficiency showed that conventional definitions such as feed conversion efficiency or residual feed intake may be inappropriate measures of efficiency for lactating dairy cows. An alternative definition, residual solids production, is proposed. This definition of feed efficiency identifies animals that produce greater volumes of milk solids at similar levels of feed intake without excessive body tissue mobilization and with improved fertility performance. The results also suggest that although there are differences in feed efficiency between strains of Holstein-Friesian, there is also variation within genotypes so that improvements in feed efficiency can be realized if the appropriate definition of feed efficiency is incorporated into breeding programs.

Increasing milk solids production across lactation through genetic selection and intensive pasture-based feed system

The objective of the study was to quantify the effect of genetic improvement using the Irish total merit index, the Economic Breeding Index (EBI), on overall performance and lactation profiles for milk, milk solids, body weight (BW), and body condition score (BCS) within 2 pasture-based systems of milk production likely to be used in the future, following abolition of the European Union's milk quota system. Three genotypes of Holstein-Friesian dairy cattle were established from within the Moorepark dairy research herd: LowNA, indicative of animals with North American origin and average or lower genetic merit at the time of the study; HighNA, North American Holstein-Friesians of high genetic merit; and HighNZ, New Zealand Holstein-Friesians of high genetic merit. Animals from within each genotype were randomly allocated to 1 of 2 possible pasture-based feeding systems (FS): 1) The Moorepark pasture (MP) system (2.64 cows/ha and 344kg of concentrate supplement per cow per lactation) and 2) a high output per hectare (HC) system (2.85 cows/ha and 1,056kg of concentrate supplement per cow per lactation). Pasture was allocated to achieve similar postgrazing residual sward heights for both treatments. A total of 126, 128, and 140 spring-calving dairy cows were used during the years 2006, 2007, and 2008, respectively. Each group had an individual farmlet of 17 paddocks and all groups were managed similarly throughout the study. The effects of genotype, FS, and the interaction between genotype and FS on milk production, BW, and BCS across lactation were studied using mixed models with factorial arrangements of genotype and FS accounting for the repeated cow records across years. No significant genotype by FS interaction was observed for any of the variables measured. Results show that milk solids production of the national average dairy cow can be increased across lactation through increased EBI. High EBI genotypes (HighNA and HighNZ) produced more milk solids per cow and per hectare than the LowNA genotype (2.7 and 4.1%, respectively). The results also suggest that when concentrate supplementation is used to facilitate increased stocking rates, increased herbage utilization and decreased substitution of concentrate for herbage can be achieved. When implemented, the HC FS could increase the overall productivity of pasture-fed dairy farming systems where land availability is the primary limiting factor of production.

A meta-analysis of the impact of stocking rate on the productivity of pasture-based milk production systems

The objective of this study is to quantify the milk production response per cow and per hectare (ha) for an incremental stocking rate (SR) change, based on a meta-analysis of published research papers. Suitable experiments for inclusion in the database required a comparison of at least two SRs under the same experimental conditions in addition to details on experimental length and milk production results per cow and per ha. Each additional increased SR treatment was also described in terms of the relative milk production change per cow and per ha compared to the lower base SR (b_SR). A database containing 109 experiments of various lengths with 131 comparisons of SR was sub-divided into Type I experiments (common experimental lengths) and Type II experiments (variable experimental lengths). Actual and proportional changes in milk production according to SR change were analysed using linear mixed model procedures with study included as a random effect in the model. Low residual standard errors indicated a good precision of the predictive equations with the exception of proportional change in milk production per cow. For all milk yield variables analysed, the results illustrate that while production per cow is reduced, a strong positive relationship exists between SR and milk production per ha. An SR increase of one cow/ha resulted in a decrease in daily milk yield per cow of 7.4% and 8.7% for Type I and Type II data, respectively, whereas milk yield per ha increased by 20.1% and 19.6%, respectively. Within the Type II data set, a one cow/ha increase in SR also resulted in a 15.1% reduction in lactation length (equivalent to 42 days). The low predictability of proportional change in milk production per cow according to the classical SR definition of cows per ha over a defined period suggests that SR may be more appropriately defined in terms of the change in available feed offered per animal within each treatment.

Embryo survival in dairy cows managed under pastoral conditions

Efficient pasture-based milk production systems require a compact calving pattern aligned to the onset of the grazing season, a 365-day calving interval and low culling rates for infertility. Achievement of these targets requires high herd reproductive performance. While high genetic merit Holstein cows produce more milk in grass-based systems their fertility is compromised. Management of the modern high genetic merit Holstein dairy cow presents a major challenge in pasture-based systems of production. It appears that the extent of early embryo loss is greater (up to 20% points greater) in the modern high-producing dairy cow and that a much higher proportion of the embryos die before day 7 following insemination in contrast to heifers and lower yielding cows. About 7–8% of pregnancies are lost between days 30 and 90 of gestation with no evidence that loss rate is related to cow genetic merit, parity or level of production. Systemic concentrations of progesterone during both the cycle preceding and following insemination affect embryo survival rate with evidence that too low or indeed too high a concentration of progesterone been negatively associated with embryo survival rate. Peripheral concentrations of both progesterone and oestradiol are lowered by increased plane of feed intake due to increased metabolic clearance rate of the steroids, which is related to liver blood flow. It appears that high producing dairy cows have an increased risk of embryo death as a result of lowered peripheral concentrations of progesterone as a consequence of increased hepatic metabolism of progesterone. Uterine expression of mRNA for progesterone receptor, oestradiol receptor and retinol binding protein mRNA appears to be sensitive to changes in peripheral concentrations of progesterone during the first week after AI. It would appear that energy balance and dry matter intake during the 4 weeks, immediately after calving are critically important in determining conception rate when cows are inseminated at 70–100 days post-calving. Concentrate supplementation of cows at pasture during the breeding period has minimal affects on conception rates though sudden reduction in dietary intake should be avoided. For pasture-based systems of milk production more balanced breeding strategies, with greater emphasis on fertility and feed intake must be developed.

A systems approach to quantify greenhouse gas fluxes from pastoral dairy production as affected by management regime

A model was developed to determine what effect management practices would have on the production of the greenhouse gases (GHG) within pastorally based dairy production systems typical of those practiced in Ireland. The model simulates two levels of GHG emissions, firstly the on-farm GHG emissions of methane, nitrous oxide and carbon dioxide for example from the pastorally spreading of slurry and secondly, off-farm GHG emissions associated with both inputs brought onto the farm to maintain productivity (for example emissions arising from manufacture of concentrate feeds and fertiliser) as well as from indirect GHG emissions associated with nitrate leaching and ammonia. The aim of this work was to allow the development of effective GHG mitigation strategies at the farm level capable of reducing GHG emissions per litre of milk. Greenhouse gas emissions were modelled for nine farming systems differing in the level of concentrate supplementation (376, 810 and 1540 kg per cow per lactation) and genotype for milk production as assessed by their pedigree index (<100, 100–200 and 200–300 kg) of milk production. A three-year study to evaluate the influence of cow genetic potential for milk production and concentrate supplementation level on profitability of pasture-based systems of milk production was used to drive the Moorepark Dairy Systems Model (MDSM). Output from this model then described farm size, feed budgets, animal numbers and farm profitability when annual milk quota was set to 468,000 kg of milk year. Relating GHG emissions to annual milk sales revealed that for these pastorally based systems increasing concentrate usage reduced both on-farm and off-farm emissions, but that increasing the genotype of the dairy cow (i.e., the genetic capacity of the animal to produce milk) will increase both on-farm and off-farm GHG emissions. Lowest GHG emissions per kilogram of milk were achieved for an intermediate genotype type cow fed within a high concentrate system whilst the highest emissions were associated with high genotype cows fed within a low concentrate system. Maximum profitability was obtained when either a high concentrate feeding regime was combined with high genotype cows or where low concentrate systems were fed to low genotype cows. Relating farm profitability to GHG emissions allowed the identification of scenarios where changing from one management systems to another would achieve a simultaneous reduction in GHG emissions whilst improving farm profitability. By implementing this approach of assessing management induced change on both GHG emissions arising from the farm together with farm profitability, individual whole farm GHG mitigation strategies could be developed with a high degree of acceptability to the producer.