Palatable Grass Species Research Articles

Implications of Ecological Drivers on Roan Antelope Populations in Mokala National Park, South Africa

Climate change has massive global impacts and affects a wide range of species. Threatened species such as the roan antelope (Hippotragus equinus) are particularly vulnerable to these changes because of their ecological requirements. Attempts to address concerns about the roan’s vulnerability have not been well documented in South African protected areas. This study identifies the landscape use and distribution of the roan as well as habitat and forage suitability changes to help inform management decisions for the conservation of roan. We used fine- and broad-scale data from Mokala National Park, South Africa that includes roan occurrence data, vegetation condition indices, vegetation (structure and plant species composition), elevation and temperature differences, and precipitation strata to construct a suitability framework using the Maximum Entropy (Maxent) and Random Forest statistical package. In Mokala National Park, roan occurred in the Schmidtia pappophoroides–Vachellia erioloba sparse woodland, Senegalia mellifera–Vachellia erioloba closed woodland, Senegalia mellifera–Vachellia tortilis open shrubland, Vachellia erioloba–V. tortilis closed woodland and Rhigozum obovatum–Senegalia mellifera open shrubland. The veld (vegetation) condition index (VCI) improved from 2019 (VCI < 50%) to 2021 (VCI > 60%), with the proportion of palatable grass species (Schmidtia pappophoroides and Eragrostis lehmanniana) also increasing. This study identified four key climatic conditions affecting roan distribution, namely annual mean daily temperature range, temperature seasonality, minimum temperatures of the coldest month, and precipitation of the wettest month. These results suggest that the conservation of roan antelope should consider these key variables that affect their survival in preferred habitats and foraging areas in anticipation of changing ecological conditions.

Ecological impacts of Rosa rubiginosa on rangelands: Case of Mphaki, in Quthing district, Lesotho

The capacity of many rangelands in Lesotho is compromised due to plant invasion, resulting from unsuitable agricultural practices. Despite a high number and abundance of invasive plants in Lesotho, there are very few studies determining the invasion ecology to inform management interventions. Rosa rubiginosa L. is one of the most common invasive species invading the rangelands of Lesotho and has notable negative ecological impacts. The aim of this study was to assess rangeland condition following the invasion by Rosa rubiginosa and determine the ecological impacts thereof. This was achieved by assessing the herbaceous species cover and abundance, as well as edaphic factors following R. rubiginosa invasion. The results show that R. rubiginosa negatively affects rangelands by reducing the abundance of herbaceous species, including palatable grass species. In addition, it is shown that the plant alters soil conditions by increasing pH by 27% and affecting nutrient availability. This enhances growth of invasive species since they have competitive advantage in accessing available nutrients over native plants. It is therefore recommended that R. rubiginosa should be managed in the rangelands to improve rangeland productivity.

Enhancing ecosystem services through collaborative grass removal and fire exclusion in the Eastern Ghats

AbstractHuman activities in mesic savanna ecosystems have resulted in plant communities that are heavily dominated by fire‐tolerant grass species, are less diverse, and offer fewer ecosystem services such as palatable plant biomass. Experimental studies manipulating fire and grass presence have mostly been conducted in ecosystems invaded by exotic grass species. However, these experiments are also relevant to ecosystems that have become dominated by native grass species due to changes in human activities. Our study compared three different management treatments in the Eastern Ghats of India, where mesic savanna ecosystems are highly dominated by the unpalatable native Cymbopogon grasses, specifically: (1) control (both Cymbopogon grasses and fire are present), (2) fire exclusion, and (3) manual removal of Cymbopogon grasses and fire exclusion. We found that both grass removal and fire exclusion were necessary to significantly increase palatable herbaceous plant biomass and species diversity, and that species diversity responses were only significant at larger spatial grains of investigation. High site‐to‐site variation in the grass removal and fire exclusion treatment prevented us from detecting significant differences in species composition across treatments, even though particular palatable grass species benefited from the treatment. Our study is in line with research from other mesic savanna systems showing that fire management alone is not sufficient to transition ecosystems to a desired or historical state. We demonstrate how normally costly management practices, such as manual removal of dominant grasses, can be reasonably achieved through collaboration between scientists, community governed village institutions, and government and nongovernment agencies in socioecological systems.

Naïve plant communities and individuals may initially suffer in the face of reintroduced megafauna: An experimental exploration of rewilding from an African savanna rangeland.

Excluding large native mammals is an inverse test of rewilding. A 25-year exclosure experiment in an African savanna rangeland offers insight into the potentials and pitfalls of the rewilding endeavor as they relate to the native plant community. A broad theme that has emerged from this research is that entire plant communities, as well as individual plants, adjust to the absence of herbivores in ways that can ill-prepare them for the return of these herbivores. Three lines of evidence suggest that these "naïve" individuals, populations, and communities are likely to initially suffer from herbivore rewilding. First, plots protected from wild herbivores for the past 25 years have developed rich diversity of woody plants that are absent from unfenced plots, and presumably would disappear upon rewilding. Second, individuals of the dominant tree in this system, Acacia drepanolobium, greatly reduce their defences in the absence of browsers, and the sudden arrival of these herbivores (in this case, through a temporary fence break), resulted in far greater elephant damage than for their conspecifics in adjacent plots that had been continually exposed to herbivory. Third, the removal of herbivores favoured the most palatable grass species, and a large number of rarer species, which presumably would be at risk from herbivore re-introduction. In summary, the native communities that we observe in defaunated landscapes may be very different from their pre-defaunation states, and we are likely to see some large changes to these plant communities upon rewilding with large herbivores, including potential reductions in plant diversity. Lastly, our experimental manipulation of cattle represents an additional test of the role of livestock in rewilding. Cattle are in many ways ecologically dissimilar to wildlife (in particular their greater densities), but in other ways they may serve as ecological surrogates for wildlife, which could buffer ecosystems from some of the ecological costs of rewilding. More fundamentally, African savannah ecosystems represent a challenge to traditional Western definitions of "wilderness" as ecosystems free of human impacts. We support the suggestion that as we "rewild" our biodiversity landscapes, we redefine "wildness" in the 21st Century to be inclusive of (low impact, and sometimes traditional) human practices that are compatible with the sustainability of native (and re-introduced) biodiversity.

Distribution of Blue Bull (Boselaphus tragocamelus) and its Conservation Threats in Bardia National Park, Nepal


 
 
 Blue bull is Asia’s largest antelope, a species of least concern in IUCN Red data list of 2020. In Nepal, it is vulnerable and is often considered as a problem animal for its crop raiding habit. Although, its population is restricted in India and Nepal, there are insufficient studies conducted on the distribution and threats of the species at local level. This study aimed to assess the distribution of blue bull and its conservation threats in Bardia National Park and its buffer zone. Field survey was carried out to identify the potential area with the information provided by park staff and buffer zone people and by the transect method in the selected habitat to determine the distribution of blue bull population. Additionally, six focus group discussions (1 in each of thefive sites and 1 with park staff) and a half-day workshop (involving 25 participants representing each site and park office) were organized to assess the existing threats to the species. Data were analyzed descriptively using MS Excel, while the distribution map was prepared using Arc GIS. Also, 8 major identified threats were ranked using relative threat ranking procedure and classified into four severity classes. We found that the population of blue bull was dispersed from core area of Bardia National Park towards the buffer zone area. Open grazing, invasive species, predation by tiger and flash flood were the major threats to the blue bull as perceived by the local people. Habitat management activities including control of grazing, removal of invasive plant species, plantation of palatable grass species, increase in other prey species of tiger and control of flood in blue bull’s habitats are recommended to protect the species and thus sustain their threatened population.
 
 

Nutrient concentrations in two savannah grass species and their implications for grazers

The diversity of grazers on savannahs exerts pressure on the grass to provide nutrition. In this study the concentrations of major and trace animal nutrients in above ground grass tissue as influenced by grazing were examined. Two palatable grass species grazed by both livestock and wild grazers were sampled: Brachiaria nigropedata and Eragrostis lehmanniana. Samples of grass tissue were collected from widely spread sampling sites in both livestock and wildlife grazing sites, at the end of the growing season. Although B. nigropedata had higher nutrient concentrations, there was general covariance in nutrient levels in the two grasses. The covariance indicated that the respective sites generally had location context-dependent low or high nutrient concentrations in all grasses. The adequacy of the nutrient concentrations for grazer nutrition was assessed using literature-sourced minimum and maximum tolerable limit requirements of cattle and the high population gazelle-like species as illustration. The results showed that, in these nitrogen-limited savannahs, the grass generally met grazer requirements in terms of crude protein, potassium, molybdenum, nickel and zinc but that it had deficiencies in phosphorous, sodium, and boron, as well as grazing intensity-related localised toxic levels of potassium, phosphorous, magnesium, iron and manganese. The deficiencies and toxicities were inferred to result in abnormalities in grazer physical condition and reproduction. Given the importance of livestock in semiarid savannahs, these effects on livestock grazers were inferred as threatening human food security. The study shows that high grazing intensity can raise the concentrations of some animal nutrients in above ground grass tissue to levels above grazer tolerable limits. Grazing rotation to reduce grazing intensity is recommended as a proactive grazing management strategy to limit the effects.

Coexistence between wildlife and livestock is contingent on cattle density and season but not differences in body size.

Many studies on the coexistence of wildlife with livestock have focused primarily on similar-sized species. Furthermore, many of these studies have used dietary overlap as a measure of potential competition between interacting species and thus lack the important link between dietary overlap and any negative effects on a particular species–a prerequisite for competition. Consequently, the mechanisms that drive interspecific interactions between wildlife and cattle are frequently overlooked. To address this, we used an experimental setup where we leveraged different cattle stocking rates across two seasons to identify the drivers of interspecific interactions (i.e. competition and facilitation) between smaller-bodied oribi antelope and cattle. Using direct foraging observations, we assessed dietary overlap and grass regrowth, and also calculated oribi nutritional intake rates. Ultimately, we found that cattle compete with, and facilitate, smaller-bodied oribi antelope through bottom-up control. Specifically, cattle facilitated oribi during the wet season, irrespective of cattle stocking density, because cattle foraging produced high-quality grass regrowth. In contrast, during the dry season, cattle and oribi did not co-exist in the same areas (i.e. no direct dietary overlap). Despite this, we found that cattle foraging at high densities during the previous wet season reduced the dry season availability of oribi’s preferred grass species. To compensate, oribi expanded their dry season diet breadth and included less palatable grass species, ultimately reducing their nutritional intake rates. Thus, cattle competed with oribi through a delayed, across-season habitat modification. We show that differences in body size alone may not be able to offset competitive interactions between cattle and wildlife. Finally, understanding the mechanisms that drive facilitation and competition are key to promoting co-existence between cattle and wildlife.

Evaluating Restoration Methods A Cross A Range of Plant Communities Dominated by Invasive Annual Grasses to Native Perennial Grasses

The objectives of the study were to determine whether there are differences in plant community response to restoration methods & to create awareness among the pastoralists/ agro-pastoralists on how to restore degraded rangeland. Six treatments were established: (1) ripping, mulching and seeding (2) ripping, manuling and seeding (3) ripping, mulching, manuling and seeding (4) control. The treatments were arranged in a randomized complete block design with factorial and replicated three times in each plant community. Each treatment plot was 3m by 6m, and depending on the treatment assigned to the plot, measurements for the vegetations were taken from those plots. Seeds of perennials, large tufted and palatable grass species were selected for reseeding purpose i.e., Andropogon canaliculatus, Tetrapogon cenchriformis, Panicum coloratum and Chrysopogon plumolosus. The study was carried out over the last three rainy seasons from 2015/16 and 2017/ 2018. Data were collected from germination to dry matter yield within three years of the experimental years. Accordingly, the result showed as variation clearly observed among the restoration treatments to different plant community. There is a significant difference at (P<0.05) rangeland parameters collected in all Restoration methods in each seasons. A total of 18 species of grasses were identified in the study district. Of the grass species 11(61.1%) species were perennials and 7 (38.9%) species were annuals. In Perennial plant community with Restoration Methods of Ri+Ma+Mu+Se was significant difference at (P<0.05) in the three consecutive seasons in Biomass production. The highest Biomass production was observed in the third season of implementation time which was 2.37 ton/ha while the lowest biomass production was observed in the first season of implementation period in control one which was 1 ton/ha. In Mixed plant community with Restoration Methods of Ri+Ma+Mu+Se was significant difference at (P<0.05) in the three consecutive seasons in Biomass production. The highest Biomass production was observed in the third season of implementation time which was 1.75 ton/ha while the lowest biomass production was observed in the first season of implementation period in control one which was 0.83 ton/ha. Therefore, we conclude that from this finding, perennial plant community with Restoration methods of Ri+Ma+Mu+Se is more effective method to rehabilitate denuded rangeland areas in semi-arid areas and also Annual plant community with Restoration methods of Ri+Ma+Mu+Se has shown dramatic result to improved range resources within short period of time. However, continuous onsite training & practical demonstration was very crucial to enhance the adoption rate of those Restoration methods. Keywords: Restoration Methods, plant community, Germination, Basal cover & Biomass Yield DOI : 10.7176/JBAH/9-19-01 Publication date :October 31 st 2019

Combined effects of grazing management and climate on semi‐arid steppes: Hysteresis dynamics prevent recovery of degraded rangelands

Abstract Livestock grazing has degraded many arid and semi‐arid rangelands around the world, and the drier climate predicted by climate change scenarios may amplify these effects and even lead to catastrophic vegetation shifts. We assess the long‐term effects (1900–2100) of grazing and rainfall on various aspects of vegetation structure including the grass‐shrub balance, the maintenance of spatial vegetation patterns, and the decline or recovery of palatable grasses (e.g. Poa ligularis) on a cover and/or density basis. We used the eco‐hydrological and individual‐based simulation model DINVEG for this purpose, which describes the spatiotemporal dynamics of Patagonian grass‐shrub steppes based on six decades of field research (1955–2018). Rainfall and grazing affected the simulated vegetation structure in different ways. Total plant cover was mostly influenced by rainfall, but the cover of palatable grasses was mostly influenced by stocking rate. Dry conditions and low stocking rates (122 mm/year and <0.2 sheep/ha) favoured grasses over shrubs, whereas shrub encroachment occurred only in the high rainfall scenario combined with high stocking rates (181 mm/year and >0.2 sheep/ha). High stocking rates and/or drier conditions caused only gradual shifts in spatial vegetation patterns, but maintained the observed positive association for grasses around shrubs. In contrast, shrub encroachment was associated with repulsion between grasses and shrubs and the formation of shrub clusters into a matrix of scattered less palatable grasses. Plant compositional changes occurred through grass species replacement (e.g. P. ligularis is replaced by Pappostipa humilis) and the associated hysteresis effect of palatable grass species: model simulations suggest that 2–3 decades of heavy and year‐long continuous grazing can drive palatable grasses to close to extinction, whereas natural recovery of degraded steppes may take 100 years or longer. Synthesis and applications. Desertification and climate change challenge grazing management in semi‐arid rangelands, especially in already degraded ecosystems. Management that alternates between years of grazing and resting was effective to maintain the cover of palatable grasses, but this allowed for only very slow recovery of degraded steppes. While drier climate and grazing may not change the overall spatial patterns of vegetation, our results are rather pessimistic regarding the short‐term recovery of palatable grasses. This will require increasing complexity in ecosystem restoration efforts, combined with interventions such as sowing, watering, reseeding or major changes in land use.

Restoration potential of invaded abandoned agricultural fields: what does the seed bank tell us?

Soil seed banks can play an important role in the restoration of degraded ecosystems, especially where indigenous species are well represented in, and invasive species are largely absent from, the seed bank. Here, we studied the potential contribution of the soil seed bank to the restoration of invaded, abandoned agricultural fields in the Eastern Cape, South Africa. We recorded the aboveground cover and belowground abundance of all vascular plant species from 120 quadrats that differ in cover of the extralimital woody invader, Pteronia incana. Our results show that higher cover of P. incana is associated with lower species richness, aboveground cover, and belowground seed abundance. Furthermore, community similarity between the above‐ and belowground component was low, with the seed bank and standing vegetation having only 15 species in common and 49 species being recorded only from the seed bank. We suggest that this large number of seed bank‐only species is a relic of previous vegetation, prior to large‐scale invasion by P. incana. The most important finding from our study is the absence of P. incana from the soil seed bank. This finding, combined with the large number of mostly native species from the seed bank, holds promise from a restoration perspective. However, given the susceptibility of the invaded systems to erosion, coupled with the low grazing value of the seed bank species, we suggest that P. incana removal should be accompanied by both erosion control measures and reseeding with palatable grass species, to secure the livelihoods of local communities.

Behavioral and Ecological Implications of Bunched, Rotational Cattle Grazing in East African Savanna Ecosystem

Rangelands and the wildlife and livestock they support are critical to human livelihoods, but rangeland ecosystems increasingly suffer from overgrazing and degradation. Planned grazing, a strategy that commonly involves time-controlled rotations of high-density (bunched) groups of cattle across a pasture, is marketed as a method to enhance rangeland health and lessen livestock impacts. However, the behavioral mechanisms underlying any potential rangeland improvements resulting from rotational, high-density planned grazing have rarely been examined. To investigate these mechanisms, we compared planned grazing with conventional continuous grazing management in a savanna ecosystem in Kenya. We surveyed cattle grazing behavior, measured changes in vegetation characteristics through surveys conducted before and after cattle grazing, and measured native ungulate abundance following grazing using camera traps. Stocking rates were held constant across treatments, resulting in a commensurate decline in total foliar hits per pin (a proxy for vegetative biomass) across treatments. Planned grazing management altered cattle behavior and reduced grazing selectivity by restricting movements, causing cattle to walk more slowly while grazing and to take more bites per step. Vegetation survey results supported this finding: cattle in the planned grazing treatment ate significantly more Pennisetum grasses (typically avoided because of their unpalatability), creating the opportunity for regrowth of more palatable species after seasonal rains. We also documented significantly higher zebra presence in planned grazing plots after cattle grazing, likely due to increased relative abundance of more palatable grass species. This investigation of grazing behavior, and specifically decreased grazing selectivity as a mechanism underpinning the benefits of planned grazing, shows that when conducted at appropriate stocking densities, planned grazing has the potential to help mitigate rangeland degradation and improve rangeland sustainability for both livestock and wildlife in pastoral African savanna ecosystems.

Short duration overnight cattle kraaling in natural rangelands: Implications for grass composition, quality, above ground biomass, species diversity and basal cover

Short duration overnight cattle kraaling in natural rangelands is an innovative management practice aimed at improving grazing availability. We tested the ability of this practice to alter grass composition in favour of palatable species, increase aboveground grass biomass, species diversity and basal cover using a chronosequence consisting of sites kraaled 6, 12, 24 and 36 months prior to the study and compared them to the surrounding vegetation. We also determined grass quality in terms of fibre content, digestibility and nutrient content [nitrogen (N), phosphorus (P), calcium (Ca), potassium (K) and magnesium (Mg)] 12, 16 and 20 months after kraal use and in surrounding vegetation. Previously kraaled sites had higher proportions of palatable grass species, diversity, N, P, K and Mg than surrounding vegetation. Aboveground grass biomass and basal cover were lower in previously kraaled sites than surrounding vegetation, while grass fibre, digestibility and Ca content did not change with kraaling. These results demonstrate that establishing short duration overnight cattle kraals in natural rangelands improve grazing quality in terms of abundance of palatable grass species and grass nutrient content. However, improved grass quality attracted grazers such as warthogs (Phacochoerus aethiopicus), which reduced grass biomass and basal cover.

What are the long-term effects of high-density, short-duration stocking on the soils and vegetation of mesic grassland in South Africa?

High-density, short-duration stocking (HDG) is gaining popularity amongst farmers in the South African mesic grasslands, but little is known about its potential impact on natural resources. Using a fence-line contrast approach, this study compared the long-term effects of HDG with those of other rotational grazing systems at lower densities (LDG) on soil properties and vegetation composition at two sites. Soils at Kokstad and Cedarville were 31% and 19% more compacted under HDG than LDG, respectively, but did not differ in total nitrogen, total carbon, available phosphorus or pH. At Kokstad, HDG had promoted unpalatable grass species at the expense of palatable grass species. Forb species composition differed substantially between HDG and LDG, with HDG having 13 and LDG having 16 unique species. A radical prostrate forb growth habit was favoured over a cauline erect habit under HDG. At Cedarville, few grass or forb species showed differences between HDG and LDG, possibly because of an overriding influence of the absence of burning in both grazing systems. These results suggest that HDG potentially has negative impacts on soil health and vegetation composition of South African mesic grassland.

Comparison of soil physico-chemical properties under different land-use and cover types in northeastern China's Horqin Sandy Land

The Horqin Sandy Land of northeastern China was originally a grassland with plenty of water and lush vegetation dominated by palatable grass species along with sparsely scattered woody species. However, it has experienced severe desertification in recent decades due to its fragile ecology together with inappropriate human activities. Currently, the landscape of the Horqin Sandy Land is dominated by irrigated croplands and sand dunes with different degrees of vegetation cover, as the region has become the most important part of the semiarid agro-pastoral ecotone of northern China. In this study, we compared soil physical and chemical properties under different land-use and cover types (irrigated cropland, rainfed cropland, sandy grassland, fixed dunes, and mobile dunes) . We found that soil particle size distribution; organic C, total N, and total mineral element, microelement, and available microelement and nutrient contents; pH; CEC; and bulk density differed significantly among the land-use and cover types. In general, soil quality was highest in the cropland, intermediate in the sandy grassland, and lowest in the dunes. The most important soil quality attribute, soil organic carbon (SOC) storage, decreased in the following order: irrigated cropland (5, 699 g/m 2 ) ＞ sandy grassland (3, 390 g/m 2 ) ＞ rainfed cropland (2, 411 g/m 2 ) ＞ fixed dunes (821 g/m 2 ) ＞ mobile dunes (463 g/m 2 ) . SOC was significantly positively correlated with a large proportion of the other soil physico-chemical parameters. Our results suggest that the key issue in restoration of the degraded soils will be to increase SOC storage, which would also create a high potential for sequestering soil C in desertified areas of the Horqin Sandy Land.

Effect of short-duration overnight cattle kraaling on grass production in a southern African savanna

Ranch owners in eastern and southern Africa have embraced a new rangeland management practice of keeping cattle overnight in temporary livestock corrals (hereafter referred to as kraals) within rangelands for short durations (for example, one week) to improve grass production. To better understand the influence of this practice on the grass community, we determined grass species diversity, richness, biomass and quality (crude protein, fibre, rumen fermentation and metabolisable energy) from a chronosequence consisting of kraals abandoned 1, 2, 3 and 4 years prior to the study and compared them to the surrounding vegetation. Grass species diversity, richness, biomass and quality were higher in abandoned kraals than the surrounding vegetation. Grass biomass increased five-fold in the first year of kraal abandonment and thereafter declined three-fold after four years. Metabolisable energy, a measure of grass nutritive value, increased from 8.92 to 11.90 MJ kg−1 with kraaling. The number of grass species also increased with kraaling. Palatable grass species, such as Urochloa mosambicensis and Panicum maximum, were more abundant in abandoned kraal sites than the surrounding vegetation. We conclude that short-duration overnight cattle kraaling improves grass quality and biomass.

Rangeland management effects on soil properties in the savanna biome, South Africa: A case study along grazing gradients in communal and commercial farms

Although the savanna biome of South Africa is a major resource for rangeland management, little is known about how differences in rangeland management systems affect soil properties in such biomes. Near to Kuruman, commercial farms have practiced rotational grazing for decades. In communal areas of former homeland Bophuthatswana, similar strategies were used prior to 1994. Nowadays, a continuous grazing system is common. We hypothesized that these changes in management affected soil properties. To test this, we sampled soils at communal and commercial land along a gradient with increasing distance to water points. The results revealed that communal systems with continuous grazing showed enlarged spatial gradients. The soils were depleted in most nutrients close to the water relative to those of commercial systems. In contrast, as the distance to the water increased, the nutrient stocks of these communal systems were higher. Changes in soil nutrient stocks were related to a zone of increased bush encroachment (up to 25%). Specific analyses (phosphorus fractions, particulate organic carbon, δ13C) confirmed that the soils of the communal grazing systems benefited from the shift of grass-dominated to bush-dominated system with woody Acacia vegetation, while the rangeland degraded in the sense that it lost palatable grass species.

Checklist of Palatable Grass Species from Peninsular India

Grasslands are common landscapes of India and dominate peninsular Indian landmass. Though secondary, these grasslands are important from cultural as well as economical point of view. Domestic cattle population of this region largely depends on these grasslands as food source. The grasslands, exposed to anthropocentric activities like burning and grazing, are facing several changes like replacement of palatable species by unpalatable ones. As an attempt to understand the fodder potential of grasslands of this region, a comprehensive checklist of palatable grass species, based on literature survey and field experience, was compiled. Local pastoral communities were interviewed for information on palatable grass species and their utility potentials. Various herbaria were consulted for confirmation of habitats, presence and absence of awns and grass phenology. The grasses with awns are consumed before the maturation and after the dispersal of awns. A total of 143 grass species were documented as palatable, of which 64 are awned and 79 are unawned. The palatable grasses were classified in 9 habitats and the palatability grade based on their use value is assessed. This documentation will be helpful for understanding and better management of grasslands of peninsular India.

Checklist of Palatable Grass Species from Peninsular India

Checklist of Palatable Grass Species from Peninsular India

Influence of combined grazing by wild boar and small ruminant on soil and plant nutrient contents in a coppice oak forest

AbstractLimited information is available about the impact of grazing on soil and plant nutrient contents in Mediterranean deciduous oak forests grazed by both livestock and wild life. Thus, the main objective of this study was to investigate the effect of wild boar and small ruminants grazing in a coppice oak forest in N Greece. Eight even‐aged stands of similar canopy cover and site quality were selected: four stands with long‐time grazing mainly by wild boar and four stands with long‐time grazing by sheep and goats. A plot of 150 m2in each of the stands was fenced in order to be protected from grazing. A similarly sized plot open to grazing was assigned next to the fenced plot. Species composition was recorded in fenced and grazed plots. Soil samples and individual plants of the dominant palatable grass speciesDactylis glomerata andBrachypodium sylvaticumwere collected in every fenced and grazed plot. These samples were analysed for contents of nutrients. The results indicated that grazing increased organic matter, C, and total N, P and Ca, while it had no effect on the plant nutrients Mg, Na, K and all micronutrients, and pH as well. Wild boar grazing had a significantly negative impact on the concentrations of Na, Fe, Zn, and Mn in the soil. Beside a decrease in K and Mn, grazing had minimal effects on the contents of plant nutrients. Conversely, wild boar grazing increased the contents of several nutrients such as N, Na, Cu and Fe in plant tissue. It seems that species composition was more affected by the topography and management than by the soil features as revealed by RDA analysis. The results of this study indicate that it is important to maintain moderate small ruminant grazing intensity for the sustainable multi‐disciplinary management of coppice oak forests. It is also important to control wild boar population in order to facilitate ecosystem process, soil preservation and forage quality.

Refuge effect of an unpalatable forb on community structure and grass morphology in a temperate grassland

The role of unpalatable plant species as biological antiherbivore refuges for palatable species is well-documented at community level particularly in harsh environments. In productive sub-humid temperate grassland subjected to domestic grazing, we examined the protective effect of Eryngium horridum on plant community structure and floristic composition, and evaluated whether these changes impacted on a number of morphological traits of grasses, related to grazing resistance. We also investigated, for a palatable grass species (Stipa neesiana) the existence of morphological differences between protected and unprotected plants and if this eventual variation was either plastic or genetic. The study consisted of a field survey where we compared paired patches, with and without E. horridum, and a greenhouse experiment where we evaluated individuals of S. neesiana coming from both patch types over a 11 months period. Patches dominated by E. horridum had lower richness and cover of forbs than patches without the forb, and similar richness but greater cover of cool-season tussock palatable grasses, which suggests a protective role on the latter. Grasses in these patches also had longer blades and sheaths and lower specific leaf area. The morphological differences of S. neesiana individuals collected from both patch types disappeared after 11 months growth in a common environment which revealed significant phenotypic plasticity in this species. These results suggest the existence of plant-to-plant facilitation in a productive ecosystem not only at community level, through changes in species richness and the promotion of palatable grasses, but also at population level, through plastic changes in aboveground morphological traits. Both facilitation and plasticity, would contribute to the persistence of threatened palatable grasses in the heavy grazed productive ecosystems.