In this paper, the vibration characteristics (free vibration and forced vibration) of fiber-reinforced thin-walled conical–cylindrical composite shells (FTCCS) are analyzed by combining theory and experiment. Based on the classical shell theory and Kirchhoff–Love assumption, the overall structure of the FTCCS is theoretically modeled. The artificial spring technology is used to simulate the arbitrary boundary conditions at the joint, which is divided into a main spring and a secondary spring. At the same time, the displacement functions of the two sub-structure shells of the FTCCS are established by the orthogonal polynomial method, and the energy function of the FTCCS is constructed according to the continuity condition of the bolt joint. Then, the vibration characteristics of the FTCCS are solved using the Rayleigh–Ritz method and the modal superposition method. Finally, the TC300/epoxy resin-based fiber-reinforced thin-walled conical–cylindrical composite shells are selected as the research object, and the rationality of the mathematical model is verified by pulse, frequency sweeping and resonance excitation tests. The experimental results show that the errors between the natural frequency and the resonance displacement calculated by the theory and the experimental results are 1.75–4.76% and 3.79–9.28%, which confirms the rationality of the proposed model. By changing the length of the shell, the lamination angle and the semi-cone angle of the conical shell, the vibration characteristics parameters under different conditions are calculated to evaluate the influence of three physical parameters on the vibration characteristics of the FTCCS.