Objective Synonym Research Articles

Synonymic notes in the Southeast Asian genus Lutera Westwood, 1875 (Coleoptera: Scarabaeidae: Rutelinae)

To resolve the taxonomic confusion surrounding two Asian Rutelinae species, Lutera nigromaculata Ohaus, 1900 and L. nigrita Ohaus, 1930, all primary types of relevant taxa have been revisited. A redescription of L. nigromaculata is given, and diagnosis and color illustrations are provided for both L. nigromaculata and L. nigrita. Parastasia ohausi Chûjô, 1966 (new synonym), an unnecessary replacement name for and junior objective synonym of L. nigromaculata, was found in a neglected publication by the late Michio Chûjô. A lectotype is designated for L. nigromaculata guttata (Frey, 1970) (new synonym) which is recognized as another junior synonym of L. nigromaculata. All known distribution records of the two species are mapped, and L. nigromaculata is newly recorded from India and Vietnam. The exact location of “Lakhon” in Indo-China, which is currently Nakhon Phanom of Thailand, is also discussed.

A second species of the genus Manodactylus Moser, 1919 (Coleoptera, Scarabaeidae, Melolonthinae, Macrodactylini) from the highlands of Colombia

Moser in 1919 described the genus Manodactylus for the new species Manodactylus gaujoni Moser, 1919, which was described based on specimens of Abbé Gaujon from Loja, Ecuador; a syntype of this species was designated in 2017 as the lectotype. Earlier, Macrodactylus gaujoni was described by Ohaus in 1909 based on a specimen collected also in highlands of Loja; however, this type specimen is presumably lost. We designate the lectotype as a neotype for Macrodactylus gaujoni Ohaus, 1909 based on the congruence between the type localities and the diagnostic characters mentioned in the descriptions. Thus, Manodactylus gaujoni Moser, 1919 is a junior objective synonym of Macrodactylus gaujoni Ohaus, 1909, which is presented here in the new combination under the genus Manodactylus. Also, a second species of the genus Manodactylus is described from the Paramo ecosystem in the Andes of southern Colombia. Manodactylus paramicolasp. nov. differs from M. gaujoni by the shape of clypeus and punctation of pronotum and elytra. Diagnosis for the genus and for the species, the description of M. paramicolasp. nov., and a distribution map are presented.

The discovery and naming of the Sumatran Rhinoceros (Dicerorhinus sumatrensis) after 1793, with details of the Rhinoceros Sumatricus of Bertuch (1805) and Wilhelm (1808)

There was a considerable gap of 21 years between the illustrated description of a specimen of the Sumatran Rhinoceros (Dicerorhinus sumatrensis) by William Bell in 1793 and the proposal to accept it as a named new species by Gotthelf Fischer in 1814. In the meantime some 17 authors reported the possibility that the animal should be regarded as a new species. Hitherto overlooked, the German writer Friedrich Justin Bertuch used a new name in the caption to a reproduction of Bell’s plate, as Rhinoceros bicornis Sumatricus. This was published in 1805 in a second edition of his “Bilderbuch für Kinder”, a serial magazine aimed at young people. This work has an extremely complex bibliography through many re-issues and translations, which is only partly resolved. The same name in the combination Rhinoceros Sumatricus was included by Gottlieb Tobias Wilhelm in a companion to the “Bilderbuch” available in 1808. The complicated circumstances of Bertuch’s proposal in 1805 and his unusual usage of a trinomen lead to the suggestion that his book might not be consistently binominal, making the name unavailable. The naming by Wilhelm in 1808 is straightforward and correct, hence Rhinoceros sumatricus Wilhelm, 1808 is an available name and a senior objective synonym of Rhinoceros sumatrensis G. Fischer, 1814. The International Code of Zoological Nomenclature allows the suggestion that Wilhelm’s name was a nomen oblitum (forgotten name). The various plates of the rhinoceros found in the “Bilderbuch” by Bertuch and the “Unterhaltungen aus der Naturgeschichte” are figured and explained.

Taxonomic review of the genus Lemyra Walker (Erebidae: Arctiinae) from India

In the present manuscript, we review the genus Lemyra Walker, 1856 from India by examining the specimens collected freshly from different parts of India from 2016 to 2019 and from the National Zoological Collection of the Zoological Survey of India (NZCZSI). The revised status of Lemyra in India is presented by describing seven new species: Lemyra (Thyrgorina) rubripecta sp. nov., L. (T.) densimacula sp. nov., L. (T.) fortiorspina sp. nov., L. (T.) apatani sp. nov., L. (T.) adunca sp. nov., L. (T.) obsoleta sp. nov. and L. (T.) pinus sp. nov., synonymizing equal number of species: L. (T.) punctilinea (Moore, 1879), syn. nov. under L. (T.) flavalis (Moore, 1865), L. (T.) magnaproteus N. Singh & Kirti, 2016, syn. nov. under L. (T.) proteus (De Joannis, 1928), Thanatarctia costalis Kaleka, 2012: 51, syn. nov., Thanatarctia collarlis Kaleka, 2012: 53, syn. nov., L. (T.) collarlis N. Singh, Kirti & Kaleka, 2016: 15, syn. nov. (objective synonym of T. collarlis Kaleka, 2012), L. (T.) pseudocollarlis N. Singh, Kirti & Kaleka, 2016: 16, syn. nov. under L. (T.) nigrescens (Rothschild, 1910) and L. (T.) angularis (Strand, 1919) under L. (T.) spilosomata (Walker, [1865]). The status of L. (T.) aurantiaca (Fang, 1985), a new record to India, is upgraded to species level. Additionally, two species i.e., L. (T.) kaikarisi Saldaitis et al., 2019 and L. (T.) persephone Saldaitis et al., 2020 are reported for the first time from India. An updated checklist of 117 species (including the nominotypical subspecies) and 10 subspecies of Lemyra is provided with 48 species (including the new records and new species described herein) from India. Of the 48 species from India, the systematics of 42 species is provided based on their voucher specimens. We could not study the rest of the six species with their voucher specimens. The detailed diagnoses with closely similar congeners, genitalia illustrations, distributions, individual variations and sexual dimorphism, if any, and bionomics is provided.

Two invalid genera in the family Bythinellidae Locard, 1893 (Caenogastropoda: Truncatelloidea)

Within the genus Bythinella Moquin-Tandon, 1856, with about 250 nominal species, the separation of several genera would be helpful in understanding the phylogenetic relationships between these snails, but neither morphology nor molecular loci studied so far can be used to distinguish any natural taxon over the range of species within this genus. Apart from Bythinella there are two other genera listed in the Bythinellidae by WoRMS: Terrestribythinella Sitnikova, Starobogatov et Anistratenko, 1992, and Strandzhia Georgiev et Glöer, 2013. Considering the cytochrome c oxidase subunit I (COI) sequences, as well as poor morphological data, Terrestribythinella should be considered a junior subjective synonym of Bythinella. In the case of Strandzhia, both COI and 18S sequences studied by us, as well as morphology re-examined (misinterpretation of the penis morphology in the original description) clearly show the assignment of Strandzhia to the genus Grossuana Radoman, 1983, thus Strandzhia becomes a junior objective synonym of Grossuana.

Revision of Lower Devonian clam shrimp (Branchiopoda, Diplostraca) from the Rhenish Massif (Eifel, SW-Germany), and the early colonization of non-marine palaeoenvironments

Three collections of clam shrimp from the Lower Devonian (upper lower Emsian) Klerf Formation of Willwerath and Waxweiler in the western Eifel Mountains are herein studied. Four discernible morphotypes are present in the Willwerath assemblage. These morphotypes correspond to the previously described species from this locality: Pseudestheria diensti (Gross), Pseudestheria subcircularis Raymond, and Palaeolimnadiopsis? eifelensis Raymond. The fourth morphotype also closely corresponds to Pseudestheria diensti, but is slightly different in outline. Asmussia willweratica (Novozhilov) is an objective synonym of Pseudestheria subcircularis. In the Willwerath clam shrimp sample, it is difficult to disentangle taphonomic/preservational versus ontogenetic/sexual variation. If taphonomic/preservational causes could be confirmed, all the Willwerath specimens may represent just one taxon and Pseudestheria subcircularis and Palaeolimnadiopsis? eifelensis would be potential subjective synonyms of Pseudestheria diensti. However, we refrain from a formal taxonomic act, which requires a larger number of specimens to be analysed.In addition, we investigated two samples from Waxweiler, which originate from distinct centimetre-thick layers and therefore represent associations. One of these associations comprises two discernible clam shrimp morphotypes. Because of limited preservation of morphological characters, one is here left in open nomenclature as Spinicaudata incertae sedis, whereas the other is recognised as a new species of the family Palaeolimnadiopseidae, Palaeolimnadiopsis frankeorum sp. nov. The second association from Waxweiler comprises at least four, possibly six different clam shrimp taxa. In addition to Pseudestheria diensti, Palaeolimnadiopsis sp., and poorly preserved Asmussia- and Ulugkemia-like specimens, two taxa can be described as new, namely the vertexiid Cornia wasvilrensis sp. nov. and the leaiine Eicheleaia wenndorfi gen. nov. et sp. nov. Altogether, the Klerf Formation at Waxweiler yields a total of at least five (possibly eight) clam shrimp taxa. The palaeoenvironment of these early clam shrimp from the Klerf Formation is interpreted as a non-marine, relatively proximal deltaic setting with no or very minor marine influence, such as a deltaic freshwater pond or lake. The comparatively high diversity of these lower Devonian clam shrimp suggests a considerably earlier origin and cryptic evolution of the group.

New combinations and synonymies in the weevil genus Lyterius Schnherr (Coleoptera, Curculionidae), with a conspectus of historical works on Daldorffs Sumatran beetles.

In preparation for a future taxonomic revision, we explore provenance, collectors, original owners and current repositories of the type material for three available genus-group names, Lyterius Schnherr, 1844, Barisoma Motschulsky, 1863 and Plaxes Pascoe, 1885, which have been applied to a group of small, relatively flat weevils associated with Pandanaceae. Lectotypes are designated for Rhynchaenus musculus Fabricius, 1802 and Plaxes impar Pascoe, 1885. The lectotype of Rh. musculus is designated as neotype for Curculio abdominalis Weber, 1801, making the species names objective synonyms (reestablished synonymy), with C. abdominalis having date priority. The names Barisoma Motschulsky and Plaxes are placed in synonymy with Lyterius (new synonymies). Five valid species names are recognised in Lyterius, L. abdominalis (Weber), L. dispar (Faust, 1896) new combination, L. impar (Pascoe) new combination, L. instabilis Boheman, 1844 and L. pandanicola (Motschulsky, 1863) new combination. We also provide an overview of the early descriptive works on Sumatran beetles collected by Daldorff and their original owners.

Checklist of Yunnan Papilionidae (Lepidoptera: Papilionoidea) with nomenclatural notes and descriptions of new subspecies.

A checklist of the Papilionidae of Yunnan is presented, with nomenclatural and taxonomic changes made. In the nomenclatural section, the junior homonym Papilio bootes nigricans Rothschild, 1895 is replaced by Papilio bootes nigricauda Lamas & Cotton nom. nov., Chilasa (Cadugoides) epycides muhabbet Koak, 2005 is synonymised with Papilio epycides camilla Rousseau-Decelle, 1947 syn. nov., Graphium cloanthus nyghmat Koak & Kemal, 2000 is placed as a junior objective synonym syn. nov. of Graphium cloanthus clymenus (Leech, 1893), and Papilio astorion Westwood, 1842 is shown to have priority over Papilio varuna White, 1842, thus the valid species name is Atrophaneura astorion (Westwood, 1842) comb. nov. In the main checklist, five new subspecies are described: Parnassius cephalus haba Hu & Cotton ssp. nov., Lamproptera curius hsinningae Hu, Zhang & Cotton ssp. nov., Lamproptera curius yangtzeanus Hu & Cotton ssp. nov., Graphium macareus vadimi Cotton & Hu ssp. nov., and Papilio krishna benyongi Hu & Cotton ssp. nov. The First Reviser Principle under the ICZN Code is invoked to solve four taxonomic problems, and 18 names are synonymised with explanations, notably Papilio machaon venchuanus Moonen, 1984 syn. nov., which is synonymised with Papilio machaon schantungensis Eller, 1936. Byasa genestieri (Oberthr, 1918) stat. nov. is separated from Byasa latreillei (Donovan, 1826), and Papilio everesti Riley, 1927 stat. nov. and P. verityi Fruhstorfer, 1907 stat. nov. are separated from Papilio machaon Linnaeus, 1758 as species. Taxa that require further confirmation of their presence in Yunnan and those that do not occur in Yunnan are enumerated.

Taxonomic Notes and Nomenclatural Corrections on Four Sphaeromatid Isopod Generic Names (Crustacea: Isopoda: Sphaeromatidae)

Details regarding the synonymy of Nesaea Leach, 1814 and Dynamene Leach, 1814 are given and a type species is selected for Dynamene. The genus Heteruropus Verhoeff, 1942 is shown to be the senior objective synonym of Harrieta Kensley, 1987 and an expanded synonymy list for the type species, Heteruropus faxoni (Richardson, 1905) is provided.

Solving nomenclatural problems of genus-group names of the cuckoo-wasps (Hymenoptera, Chrysididae): objectively invalid and unavailable names, new type-species designations, new names, a new genus and new synonymies.

Information is provided for 143 objectively invalid or unavailable genus-group names used in Chrysididae; 34 are objectively invalid, of which 18 are junior homonyms (five of these are also currently junior subjective synonyms), 18 are junior objective synonyms (two of these are also junior homonyms) of which seven are unjustified emendations, and two are unnecessary replacement names. The remaining 109 names are unavailable; 96 are incorrect subsequent spellings, most of them were originally lapsus calami or misprints, one is hereby deemed to be an incorrect original spelling, eight are nomina nuda, three were originally published as junior synonyms and never adopted before 1961 as valid names of any taxa, and one is a vernacular name previously considered as valid. Gender, type species, taxonomic history and status are given for each name except for incorrect subsequent spellings and unjustified emendations. Some cases of homonymies with taxa currently classified as non-animal are discussed. For two such junior homonyms, denoting genus-group taxa considered as valid in the present work, new replacement names are proposed: Linsenmaierella Rosa & Pavesi nom. nov. for Chrysidella Linsenmaier, nec Pascher; and Rhipidochrysis Rosa & Pavesi nom. nov. for Pleurochrysis Bohart, nec Pringsheim. New generic synonymies are established: consequent on new type-species designations, Leptoglossa Klug becomes a junior objective synonym of Parnopes Latreille syn. nov., and Pyrochloris Klug becomes a junior objective synonym of Euchroeus Latreille syn. nov.; Pseudodichrysis Trautmann is a junior objective synonym of Dichrysis Lichtenstein syn. nov. and the latter is in turn a junior subjective synonym of Chrysis Linnaeus. Platycelia Dahlbom is selected by the authors, acting as First Revisers, as the correct original spelling, and the simultaneously published Platycoelia is therefore deemed to be an incorrect subsequent spelling. The validation of Philoctetes Abeille de Perrin, not available when originally proposed, is discussed. Morphochrysis Rosa & Pavesi gen. nov. (type species: Chrysis pulchella Spinola) is here described; it includes members of the Chrysis pulchella species group previously included in Gonodontochrysis Semenov-Tian-Shanskij & Nikol'skaya, an unavailable name. The Chrysis zaravshanica species group is merged with the pulchella species group. A new specific synonymy within this group is proposed: Chrysis zaravshanica Tarbinsky syn. nov. of C. personata Semenov-Tian-Shanskij.

A taxonomic revision of Boiga multomaculata (Boie, 1827) and B. ochracea (Theobald, 1868), with the description of a new subspecies (Squamata, Serpentes, Colubridae).

The analyses of molecular genetic data (mtDNA markers 16S, ND4, CYTB, and the nuclear marker c-mos) provided evidence that the Asian cat snake taxa Boiga multomaculata and B. ochracea actually represent a single species. They form mixed clades of low intraclade genetic differentiation. This evidence for conspecificy is supported by the lack of differentiation in all examined pholidotic and morphometric characters. Therefore, we formally place Dipsas ochracea Theobald, 1868 in the synonymy of Dipsas multomaculata Boie, 1827. We provide a summary of the tangled taxonomic history of the taxa involved in this study. Also, we resurrect Dipsadomorphus stoliczkae Wall, 1909 from synonymy of B. ochracea, for specimens exhibiting 21 midbody dorsal scale rows. Boiga stoliczkae is found in the Himalayas north and west of the Brahmaputra valley. Finally, based on the detection of historical genetic lineages within the newly defined species Boiga multomaculata we recognize three subspecies: Boiga multomaculata multomaculata (Boie, 1827), Boiga multomaculata ochracea (Theobald, 1868), and Boiga multomaculata septentrionalis n. ssp. which is distributed in northern Myanmar and Assam and Nagaland, India. We designate BMNH 1946.1.2.60 (1) as neotype of Dipsas ochracea Theobald, (2) as lectotype of D. ochraceus Günther, and (3) as lectotype of Boiga ochracea walli Smith, thereby making these names objective synonyms. Finally, we designate BMNH 94.12.31.55 as lectotype of Dipsadomorphus stoliczkae Wall.

The double life of Bulimus ovoideus Bruguière, 1789: taxonomic implications for Streptaxidae and Camaenidae (Mollusca, Stylommatophora)

ABSTRACT The name Bulimus ovoideus Bruguière, 1789 is in current use for two different species, one from East Africa and one from the Philippines. The name has also been declared as the type species of both Edentulina Pfeiffer, 1856 (Streptaxidae) and Cochlostyla Férussac, 1821 (Camaenidae). We demonstrate that the use of this name for a Philippine camaenid is based on an early misidentification. To clarify the situation, we invoke Article 70.3 of the Code of Zoological Nomenclature, concerning misidentified type species, to select the taxonomic species actually involved, Bulinus luzonicus Sowerby I, 1833, as the type species of Cochlostyla. Cochlostyla thus becomes a senior objective synonym of Canistrum Mörch, 1852. Under Article 40.2 of the Code, Helicostylinae Ihering, 1909 takes precedence of Cochlostylinae Möllendorff, 1890. We further show that the Philippine camaenids Orthostylus Beck, 1837 and Dryocochlias Möllendorff, 1898 are objective synonyms with the former name having priority. Finally, we designate a neotype for Bulimus ovoideus to stabilise its taxonomy. These actions permit the current nomenclature to be maintained with a minimum of disruption. The currently accepted name of the taxon in question is Edentulina ovoidea (Bruguière, 1789).

Resolving inconsistencies between Plötz’s descriptions and presumed type specimens of some Hesperiidae (Lepidoptera)

Comparing specimens curated in MfN as primary types of Hesperiidae names proposed by Carl Plötz with the original descriptions and unpublished drawings reveals a number of inconsistencies that we address. Lectotypes are designated for Telegonus labriaris Butler, 1877, Eudamus jalapus Plötz, 1881, and Apaustus interpunctata Plötz, 1884. Neotypes are designated for Netrocoryne seneca Plötz, 1882 and Hesperia irma Plötz, 1882. Hesperia ulphila Plötz, 1883 is treated as a nomen dubium. As a result of these designations, the following are junior objective synonyms: Netrocoryne seneca Plötz, 1882 of Telegonus labriaris Butler, 1877 and Hesperia irma Plötz, 1882 of Pamphila irma Möschler, 1879, the latter two names being homonyms. In all these instances, the original descriptions are satisfied, and, except for A. interpunctata, the current application of these names is unaffected. Taxonomically, Callimormus diaeses Schaus, 1902, stat. rest. is a valid species, not a synonym of A. interpunctata, which is a junior subjective synonym of Eutocus vetulus (Mabille, 1883).

The Hybotidae of the Our Planet Reviewed in Corsica 2019-2021 survey, with the description of three new species of Platypalpus and Tachydromia (Diptera, Empidoidea)

In the framework of the Our Planet Reviewed program, a 3-years survey of Corsica was conducted between 2019 and 2021. A total of 3,287 Empidoidea (Diptera) were retrieved from about 17% or 152 out of 876 Diptera samples and examined. They contained 31 representatives of the family Atelestidae, 649 Brachystomatidae, 778 Empididae and 1,829 Hybotidae. The hybotid genera Drapetis Meigen, 1822, Platypalpus Macquart, 1827, and Tachydromia Meigen, 1803, were studied in more detail. Three species are described here as new to science: Platypalpus pseudoarticulatoides Grootaert, n. sp., P. pseudoniveisetoides Grootaert, n. sp. and Tachydromia corsicana Grootaert, n. sp. In addition, some rare species with a limited distribution in Europe such as Drapetis hirsuticercis Stark, 2003, Platypalpus longimanus (Corti, 1907), Platypalpus parvicauda (Collin, 1926), and Platypalpus stigma (Collin, 1926) are discussed here. Platypalpus excavatus Yang & Yao in Yang et al., 2007, is considered as a new objective synonym of Tachydromia excisa Becker, 1907.

Thyreophagus calusorum sp. n. (Acari, Acaridae), a new parthenogenetic species from the USA, with a checklist of Thyreophagus species of the world

The genus Thyreophagus is distributed worldwide, containing species occurring in stored food, house dust, bark, subcortical habitats, scale insect associations, and nests of wasps and bees. As species of this genus prefer concealed habitats, the actual species diversity is currently underappreciated. In addition, some described taxa are poorly known and their species boundaries are not entirely clear. Only a few species are known from both adult and deutonymphal stages. Using high-resolution light microscope and scanning electron microscopy, we describe a new partenogenetic species, Thyreophagus calusorum sp. n. from Florida (USA) based on adult stage (female) and heteromorphic deutonymph. This species was reared in the lab and its morphological variation was also studied. We did not find any substantial variation in the shape of the female spermatheca, but in the heteromorphic deutonymph, we did find extensive variation in the shape of tibial setae hT I (a taxonomically important character). Furthermore, we summarized taxonomic data and compiled an annotated checklist of species of Thyreophagus, with 35 species-level taxa. We clarified the following nomenclatorial questions: type species designation in Monieziella Berlese, 1897 was done by A. Berlese in G. Leonardi 1900 (Tyroglyphus entomophagus Laboulbène) not by A. Jacot (1936) as it was though previously; Monieziella mali Berlese, 1897 is an unjustified emendation and junior objective synonym of the name Acarus malus Shimer, 1868, currently Hemisacoptes malus (Shimer, 1868). Three subspecies of Thyreophagus were elevated to the species status due to their distinct morphology: Thyreophagus italicus Vacante, 1989, stat. nov. (=Thyreophagus entomophagus italicus Vacante, 1989); Thyreophagus mauritianus (Fain, 1982), stat. nov. (=Michaelopus gallegoi mauritianus Fain, 1982), and Thyreophagus ponticus Kadzhaya, 1973, stat. nov. (=Thyreophagus entomophagus ponticus Kadzhaya, 1973).

The identity of Euchirograpsus polyodous (Stebbing, 1921) (Decapoda, Brachyura, Plagusiidae), with a key to the genus

Abstract The identity of the poorly known plagusiid crab Euchirograpsus polyodous (Stebbing, 1921) is clarified on the basis of a good series of specimens from Zanzibar. The species proves to be a senior synonym of Euchirograpsus madagascariensis Türkay, 1978. The species is characterized by the more elongate and rugose third maxillipeds, distinctly serrate flexor margins of the second and third ambulatory legs, presence of only five major pairs of corneous setae on the ambulatory dactylus, and structure of the male first gonopod. The holotype of Euchirograpsus madagascariensis Türkay, 1978, is here designated as the neotype of Pachygrapsus polyodous Stebbing, 1921, making both names objective synonyms. A key to the nine species of Euchirograpsus is provided.

Kannaphallus leptosomus n. sp. (Monogenea: Heteraxinidae: Cemocotylinae) parasitizing Scyris indica (Carangiformes: Carangidae) from Malaysia.

A new monogenean species, Kannaphallus leptosomus n. sp., from the gills of the diamond trevally, Scyris indica Rüppell, caught off Terengganu, Peninsular Malaysia is described with molecular data. The new species differs from other species of the genus by the morphology of the largest clamp, the presence of a penis gun-associated needle, the unarmed genital atrium size, and the presence of two independent vaginal tubes. Previous records of Kannaphallus species were compiled, and four nominal species including one species incertae sedis (K. virilis Unnithan, 1957; K. lateriporis Mamaev, 1988; K. leptosomus n. sp.; and K. mochimae Fuentes Zambrano, 1998 incertae sedis) and two undescribed species recorded as invalid names were listed. Cemocotylelloides carangis Ramalingam, 1969 was considered a junior objective synonym of K. univaginalis Ramalingam, 1960; this species was treated as Cemocotylelloides univaginalis n. comb. In addition, Unnithan's materials including type specimens of K. virilis could not be found in specimen repositories in India, and it appears that the specimens were probably not deposited in any institution or have been subsequently lost. Based on the phylogenetic analysis of 28S rDNA sequences, Heteraxinidae was broadly divided into two clades, and Heteraxininae and Cemocotylinae were shown as polyphyletic groups, respectively.

Revision of the "celia clade" of Pseudodebis Forster, 1964, with Two New Species and Notes on Papilio phorcys Fabricius, 1793 (Lepidoptera: Nymphalidae: Satyrinae).

The species-level classification of the "celia clade" of the nymphalid butterfly genus Pseudodebis Forster, 1964, is revised as part of ongoing revisionary work on this genus. The "celia clade" contains three species, of which two, Pseudodebis darrenthroopi Nakahara & Willmott, n. sp. and P. tigrillo Nakahara & Willmott, n. sp., are described and named herein based on morphology and molecular data. Consequently, we increase the described species diversity of Pseudodebis to 13, with a remarkable six species occurring in the trans-Andean region. We discuss five specific epithets associated with the clade and designate a neotype for Papilio celia Cramer, 1780, and lectotype for Taygetis keneza Butler, 1869, based on the same specimen, thus retaining the status of the former name as a senior objective synonym. We also provide a brief historical review for Papilio phorcys Fabricius, 1793, an enigmatic name currently synonymized under Papilio celia. Nevertheless, we were unable to locate the syntype(s) for this name and the identity of Papilio phorcys remains uncertain, so we retain the current synonymy as a parsimonious approach.

Checking the checkered taxonomy of Plötz's checkered skippers (Hesperiidae: Pyrgini).

We present an analysis of the names proposed by Carl Plötz in 1884 for the New World species in the genus Pyrgus Hübner, [1819] facilitated by the genomic sequencing of extant primary type specimens comparatively with a larger sample of more recently collected specimens of these species and their relatives. The changes to nomenclature suggested here are only caused by the identity of primary type specimens as revealed by their phenotypes or through genomic sequencing. All neotypes are designated to stabilize nomenclature in agreement with the current usage of these names, which in unison agrees best with the information available about them. Lectotypes are designated for the following 5 taxa: Pyrgus (Scelothrix [sic]) bellatrix Plötz, 1884 (type locality Argentina: Buenos Aires), Pyrgus (Pyrgus) willi Plötz, 1884 (type locality in Brazil: Minas Gerais), Pyrgus (Pyrgus) albescens Plötz, 1884 (type locality in Mexico), Pyrgus (Syrichthus [sic]) lycurgus Plötz, 1884 (type locality in "Central America", likely southern Mexico), and Pyrgus occidentalis Skinner, 1906 (type locality USA: Texas, San Antonio). Neotypes are designated for the following 4 taxa: Pyrgus (Pyrgus) adepta Plötz, 1884 (Herrich-Schäffer in litt.) (type locality Colombia: Bogota), Pyrgus (Scelothrix [sic]) dion Plötz, 1884 (type locality Colombia: Bogota), Pyrgus (Scelothrix [sic]) adjutrix Plötz, 1884 (Herrich-Schäffer in litt.) (type locality in Mexico: Nuevo Leon), Pyrgus (Pyrgus) insolatrix Plötz, 1884 (Herrich-Schäffer in litt.) (type locality in "Central America", likely southern Mexico). As a result, P. lycurgus and P. insolatrix are objective synonyms. The following are junior subjective synonyms: P. dion of Burnsius adepta (Plötz, 1884), Pyrgus (Syrichthus [sic]) varus Plötz, 1884 of Burnsius orcus (Stoll, 1780) and P. adjutrix of Burnsius oileus (Linnaeus, 1767). Heliopetes (Heliopyrgus) willi (Plötz, 1884) is a species-level taxon and not a subspecies of Heliopetes (Heliopyrgus) domicella (Erichson, [1849])). Genomic analysis of the lectotypes of P. albescens, P. lycurgus, and P. occidentalis establishes them as conspecific with Burnsius communis (Grote, 1872), thus depriving a distinct species currently identified as Burnsius albescens from its name, that becomes a name for Burnsius communis albescens (Plötz, 1884) in accord with its lectotype identity; P. lycurgus and P. insolatrix are its junior subjective synonyms, but P. occidentalis is a junior subjective synonym of B. communis communis. A new name Burnsius albezens Grishin sp. n. (type locality USA: Arizona, Cochise Co., Portal) is proposed for the species misidentified as B. albescens. Furthermore, genomic comparisons reveal two other new species and one new subspecies of Burnsius Grishin, 2019: B. burnsi Grishin sp. n. (type locality Mexico: Veracruz, Huatusco), B. adepta inepta Grishin ssp. n. (type locality Ecuador: Pichincha, Tandapi), and B. orcynus Grishin sp. n. (type locality Curaçao: Hato Field) that are cryptic and can be confidently identified only by their genotype.

Vaginulus Férussac, 1822, not Vaginula auctt. is the valid name for a genus of slugs (Mollusca, Gastropoda, Systellommatophora, Veronicellidae)

There has been, and to some extent remains, confusion over the valid genus name for a group of slugs in the family Veronicellidae, either Vaginulus Férussac, 1822 or Vaginula auctt., often cited as of Berthold, 1827 or sometimes as of Blainville, 1828. We show that Vaginula was first made available as an emendation of Vaginulus Férussac by Hoffman in 1925 (Article 33.2.1 of the Code) and that all previous usages of the name were therefore incorrect subsequent spellings. However, Hoffman's emendation was unjustified (Art. 33.2.3). Vaginulus Férussac is the valid name and Vaginula Hoffman is a junior objective synonym. Vaginula Hoffman is also a junior homonym of Vaginula Risso, 1826 (Protozoa, Foraminifera) and of Vaginula Sowerby, 1833 (Gastropoda, Pteropoda).