Number Of Identified Specimens Research Articles

Living in a coast–grassland ecotone: Zooarchaeology of a late Holocene shell midden in the Uruguayan Atlantic Coast

La Esmeralda is a group of Donax hanleyanus shell middens (A, B, and C) located on the Uruguayan Atlantic coast dating to the late Holocene. This paper analyzes the vertebrate remains recovered during the excavation of shell midden A. The remains were taxonomically and anatomically identified, and quantified using NISP (number of identified specimens), MNI (minimum number of individuals), MNE (minimum number of elements), and MAU (minimum animal units). Additionally, bone surfaces were examined to identify anthropogenic and natural modifications. Several vertebrate species, including marine (pinnipeds, birds, fish) and terrestrial (deer, armadillos), were captured, processed, and consumed at the site. La Esmeralda can be chronologically linked to other coastal and inland sites located near the coast, which together reflect diverse forms of landscape occupation and various economic, domestic, and ritual activities. Collectively, these settlements show a sustained use of the Atlantic coast–grassland ecotone during the late Holocene, possibly within a coastal–inland mobility circuit. The significance of coastal and continental resources in this region has been discussed for over two decades. Although isotopic studies of human remains suggests that marine resources did not play a significant role in the diet, the material record shows the simultaneous exploitation of both types of resources. This work provides new information to continue enriching this discussion.

Exploitation of Pigs during the Late Medieval and Early Modern Period in Croatia

This study investigated the historical consumption of pork in Croatia through a comprehensive analysis of pig bones from six medieval archaeological sites, comprising three castles and three monasteries dating from the 13th to the 16th century A.D. Employing a detailed morphological analysis of pig bones and teeth, the study quantified the number of identified specimens (NISP) per site. On bones and teeth, identification of sex and age was performed, and butchery patterns were documented. The results revealed a significant presence of pork in the diet of castle and monastery inhabitants, with pig bones being the most abundant animal remains. Age analysis suggested a prevalence of juvenile pigs, predominantly males. The findings implied that from the 13th to the 16th century A.D., pork was a staple in the diet of castles and monasteries in continental Croatia, sourced from pig breeding outside these sites. Typically, male pigs aged under two years were delivered for consumption to these establishments.

Tetrapod biodiversity in sambaquis from southern Brazil.

Fishermen-hunter-gatherers of sambaquis (Brazilian shell mounds) had an intimate affinity with marine-coastal environments, where they exploited a great variety of fish and mollusks that comprise the best documented fauna from sambaquis. However, other groups of animals as mammals, birds, reptiles, and amphibians are also present in these sites, but are relatively less studied. This paper is the first one focused exclusively on the Tetrapoda biodiversity of sambaquis and aimed to identify tetrapods of ten sites from southern Brazil. We present a faunal inventory and data regarding animal capture and environmental exploitation. We identified the specimens anatomically and taxonomically, analyzed them concerning fragmentation, and quantified the data for the number of identified specimens (NISP) and minimum number of individuals (MNI). Despite the high degree of fragmentation of remains, we identified 46 taxa. As expected, most were from marine animals: cetaceans (total NISP = 2,568 and MNI = 27), otariids (total NISP = 248 and MNI = 32), and seabirds (total NISP = 65 and MNI = 23), indicating great relevance of marine tetrapod fauna as a resource for sambaqui builders (79.39% of NISP). We thus document the close bond between fishermen-hunter-gatherers of sambaquis and the marine tetrapods in southern Brazil.

Animal Use Strategies in the Longshan Mountain Region of Northern China during the First Millennium BC: A Zooarchaeological Analysis of Yucun.

The first millennium BC saw the expansion of the Western Zhou dynasty in its northwestern frontier, alongside the rise and development of the Qin State in the Longshan Mountain region of northern China. Exploring the subsistence practices of these communities is crucial to gaining a better understanding of the social, cultural, and political landscape in this region at the time. While much of the research to date has focused on the Qin people, the subsistence practices of the Zhou people remain poorly understood. In this study, we analyzed animal remains from Yucun, a large settlement site associated with the Zhou people, located to the east of the Longshan Mountain. These animal remains were recovered in the excavation seasons of 2018-2020. Our results show that pigs, dogs, cattle, caprines, and horses, which were the major domestic animals at Yucun, accounted for over 90.8% of the animal remains examined in terms of the number of identified specimens (NISP) and 72.8% in terms of the minimum number of individuals (MNI), with cattle and caprines playing dominant roles. In terms of the taxonomic composition and the mortality profiles of pigs, caprines, and cattle, Yucun shared similarities with Maojiaping and Xishan, two contemporaneous Qin cultural sites located to the west of the Longshan Mountain, and differ from other farming societies in the middle and lower reaches of the Yellow River valley. Considering the cultural attributes and topographic conditions of these various sites, these findings imply that environmental conditions may have played a more significant role than cultural factors in shaping the animal-related subsistence practices in northern China during the first millennium BC.

Taphonomy of harpy eagle predation on primates and other mammals.

The goal of this study is to provide a taphonomic analysis of bone fragments found in harpy eagle nests in the Brazilian Amazonia, utilizing the largest sample of prey remains collected to date. Harpy eagle kill samples were collected from nine nests, between June 2016 and December 2020 in Mato Grosso, Brazil. We identified the specimens, calculated the number of identified specimens (NISP) and minimum number of individuals (MNI). These metrics were used to estimate bone survivability and fragmentation. A total of 1661 specimens (NISP) were collected, representing a minimum number of 234 individuals (MNI). We identified at least nine species of primates, which represent 63.8% of the individuals in the kill sample. Harpy eagles preyed mostly on the medium-sized capuchin and bearded saki monkeys (28.2% of the MNI), and two-toed sloths (17.7% of the MNI). The large woolly monkeys also represented a significant portion of the sample (11.5% of the MNI). Three distinct patterns of bone survivability were found, one characterizing two-toed sloths, another characterizing medium-sized monkeys, and a third typical of woolly monkeys. We conclude that harpy eagle predation leaves an identifiable signature on the prey with a bone survivability pattern specific to each taxon. The intertaxon variations observed in the taphonomic signatures of harpy eagle kills should be taken into account when evaluating the potential influence of these raptors as accumulators of bone material in both paleontological and neontological assemblages.

The Truth About Deer, Turtles, and Dogs: An Examination of the Ancient Maya Human-Faunal Interaction

This paper examines how Mayan individuals, families, and communities living in Mesoamerica from approximately 1200 B.C. to A.D. 1550 interacted with deer, turtles, and dogs. Data collated from archaeological, ethnohistoric, ethnographic, and written/artistic records is reviewed, and summarizes the symbolic, ritual, and ceremonial significance of these animals in the Preclassic to Postclassic Maya world.
 Deer, turtles, and dogs played important roles in the subsistence economy of Mayan individuals and family groups. Chi-square statistical analyses of the Number of Identified Specimens (NISP) and Minimum Number of Individuals (MNI) of deer, turtles, and dogs recovered from 10 Mayan archaeological sites (6 inland and 4 coastal) indicates that the availability of these animals differed at inland versus coastal locations and over time. These differences can be attributed to a combination of factors including, but not limited to, local ecosystems, changes in humidity, precipitation, and sunlight conditions, and variations in human exploitation patterns over time.

Microfauna relative abundance since the Late Pleistocene at Natural Trap Cave, Wyoming, U.S.A

Microvertebrate communities have shifted and adapted to a variety of climate extremes since the Late Pleistocene, which can inform us on how modern communities might respond to similar climatic changes. Here we evaluate the microvertebrate remains from Natural Trap Cave (NTC), Wyoming, in the Western US, as an initial evaluation of how these communities responded to changes in local climate throughout the Late Quaternary. We calculated the number of identified specimens (NISP) and the minimum number of individuals (MNI) to determine relative abundance, richness, and evenness across 3 layers at the site, from the Late Pleistocene, the Middle Holocene, and the Late Holocene. Community composition was largely affected by changes in the abundances of taxa rather than local extinctions or colonizations. Evenness was highest in the arid Late Pleistocene, 23,000 cal BP and Late Holocene, approximately 2,000 to 6,000 cal BP. Shifts in evenness were primarily driven by changes in abundance of Phrynosoma with abundances of Lagomorpha and Thomomys contributing to changes within the small mammal community. Raw richness was highest in the Middle Holocene when Cynomys and Perognathus appeared in the region while standardized richness was highest in the Late Pleistocene with these differences being caused by small Late Pleistocene sample sizes. We performed a principal coordinate analysis to compare the small mammal community of NTC to three other caves with well-represented small mammal fauna from the Late Quaternary. NTC grouped closest with Samwell Cave, in California, another cave where small mammal material is collected by packrats. Across all four caves, layers generally grouped based on cave affinity. However, Samwell and NTC layers were grouped by age rather than cave identity. Further comparison between these two caves found that evenness decreased similarly from the Late Pleistocene to the Holocene.

Looking for the earliest evidence of Ursus arctos LINNAEUS, 1758 in the Iberian Peninsula: the Middle Pleistocene site of Postes cave

Brown bears (Ursus arctos) diverged from the cave bear lineage c. 1.2 million years ago and likely originated in Asia, where the oldest fossils belong to a Middle Pleistocene chronology. Brown bear fossils from the Middle Pleistocene are scarce in the Iberian Peninsula, especially when compared to the cave bear record and they are mainly located in the southern half of the peninsula. The oldest evidence in the Iberian Peninsula is from c. 250 ka, which is 300 ka younger than the oldest European records (~550 ka; L’Arago). Here we present the brown bear fossil assemblage from Postes cave (Fuentes de León, Extremadura, southern Iberian Peninsula). Postes cave has yielded some of the oldest evidence for brown bears in Iberia with a minimum date of 244 191±2261 a BP (number of identified specimens (NISP) = 4). Additionally, the uppermost part of the Pleistocene deposit has yielded one of the largest Middle Pleistocene brown bear collections in the Iberian Peninsula (NISP = 73) with a chronology roughly between 193 and 244 ka BP, comprising both cranial and postcranial remains and including a complete hemimandible. The Postes mandible fits well within the Middle Pleistocene brown bear range of variation, as it is significantly different in size and shape from the Holocene brown bears, based on geometric morphometric analyses. We show that these differences are due to allometry. This site provides more insight into the Middle Pleistocene morphological variation of brown bears in western Europe and underscores the need for additional Middle Pleistocene fossils in the northern half of Iberia to further test palaeobiogeographical hypotheses of this species’ entrance into the largest of the southern European peninsulas.

A refitting experiment on long bone identification

AbstractRefitting is an important analytical tool in archaeology that can yield valuable information on site formation processes and on the range of activities practiced at a site, including tool production, tool curation, and discard behavior, among others. In the present paper, we use refit data from a control assemblage of red deer (Cervus elaphus) long bones to assess problems of specimen identification and representation in an experiment where bones were processed for marrow. Three goals motivated this experiment: (i) to assess how different methods of NISP (number of identified specimens) calculation affect comparisons of the relative abundances of long bone regions, (ii) to evaluate whether long bone shaft regions vary with respect to the probability of identification, and (iii) to ascertain the potential refit rate for a well‐preserved and fully‐collected sample of faunal specimens. Our results show no statistical differences in terms of patterns of skeletal representation between the two methods of NISP calculation (single vs. multiple NISP counts) that we assessed. Our data also indicate that the shape, particularly the cross‐section, of fragments clearly impacts the probability of identification and refitting. Moreover, the refitting experiment reveals that, in ideal conditions, a majority of specimens (>95%) from the NISP sample can be refitted, which leads to largely reconstructed skeletal elements. Thus, the comparatively very low refit rates recorded in archaeological sites, including samples that are well preserved, suggest that the often limited extent of excavations, along with offsite discard and/or extensive sharing of parts, substantially reduce the possibility of finding refits in a faunal sample.

Preliminary report on the faunal remains from Aragatsavan 1, Armenia

This paper presents the preliminary analysis of the zooarchaeological data from the settlement of Aragatsavan 1, Armenia. The excavation of the site is in progress and a complete stratigraphic sequence has not been obtained yet, while the excavated levels are dated to the second and third phases of the Trialeti-Vanadzor culture of the Middle Bronze Age period (19th -18/17th centuries BC). Taxonomic and metric analyses suggest that all the species are domestic. Sheep (Ovis aries) and goats (Capra hircus) constitute more than half of the number of identified specimens (NISP), followed by horses (Equus caballus) and cattle (Bos taurus). Pigs (Sus scrofa domesticus) are scarce. Butchery and cut marks have been detected on many of the samples. A certain proportion of bones is burnt. As zooarchaeological data are not numerous for the Middle Bronze Age period of Armenia and South Caucasus, the faunal remains from Aragatsavan 1 are important. They provide new information on the subsistence economy in the region during the 3rd - 2nd millennia BC.

Macromammal and bird assemblages across the late Middle to Upper Palaeolithic transition in Italy: an extended zooarchaeological review

Evidence of human activities during the Middle to Upper Palaeolithic transition is well represented from rock-shelters, caves and open-air sites across Italy. Over the past decade, both the revision of taphonomic processes affecting archaeological faunal assemblages and new zooarchaeological studies have allowed archaeologists to better understand subsistence strategies and cultural behaviors attributed to groups of Neandertal and modern humans living in the region.This work presents the preliminary results of a 5-years research programme (ERC n. 724046 – SUCCESS) and offers a state-of-the-art synthesis of archaeological faunal assemblages including mammals and birds uncovered in Italy between 50 and 35 ky ago. The present data were recovered in primary Late Mousterian, Uluzzian, and Protoaurignacian stratigraphic contexts from Northern Italy (Grotta di Fumane, Riparo del Broion, Grotta Maggiore di San Bernardino, Grotta del Rio Secco, Riparo Bombrini), and Southern Italy (Grotta di Castelcivita, Grotta della Cala, Grotta del Cavallo, and Riparo l'Oscurusciuto). The available Number of Identified Specimens (NISP) is analysed through intra- and inter-site comparisons at a regional scale, while aoristic analysis is applied to the sequence documented at Grotta di Fumane. Results of qualitative comparisons suggest an increase in the number of hunted taxa since the end of the Middle Palaeolithic, and a marked change in ecological settings beginning with the Protoaurignacian, with a shift to lower temperatures and humidity. The distribution of carnivore remains and taphonomic analyses hint at a possible change in faunal exploitation and butchering processing between the Middle and Upper Palaeolithic. A preliminary comparison between bone frequencies and the distribution of burned bones poses interesting questions concerning the management of fire. Eventually, the combined use of relative taxonomic abundance and aoristic analysis explicitly addresses time averaging and temporal uncertainty embedded in NISP counts and offers estimates of absolute change over time that can be used to support hypotheses emerging from taxon relative frequencies.

Behind the steps of ancient sheep mobility in Iberia: new insights from a geometric morphometric approach

In Western Europe, the transition from the middle Iron Age to the early Roman period implied changes in livestock practices, with the emergence of a specialized and selective animal husbandry. These changes have been related in Italy and south of France with changes in livestock management involving their mobility between ecologically complementary areas. The study of this question in the Iberian Peninsula has only been partially investigated through palaeoenvironmental analyses, and the information about the origin and significance of this phenomenon is very scarce. To shed new light on this topic we used an archaeozoological approach, with the application of geometric morphometrics. They were used to study size and shape variability in sheep astragali from 9 sites dating from the middle Iron Age to the early Roman period (5th c. BC–3rd c. AD) and located on the Pyrenees and on the north-eastern Iberian coast as a case study. The results we obtained, combined with Number of Identified Specimens (NISP) and kill-off patterns, showed local specificities in terms of breeding methods and sheep morphologies between the two areas during the middle Iron Age. On the contrary, sheep with similar size and the implementation and development of similar sheep husbandry practices in the Pyrenees and the north-eastern Iberian coast were documented during the early Roman period. These results suggest the existence of livestock links between these two areas during the Roman period, that could be involved a possible movement of sheep between the lowlands and the Pyrenees for the first time.

A Critical Review of Four Efforts to Resurrect MNI in Zooarchaeology

Evaluation of zooarchaeology’s quantitative units known as NISP (number of identified specimens) and MNI (minimum number of individuals) during the last three decades of the twentieth century suggested neither provided ratio scale measures of taxonomic abundances. Many researchers at that time began to use NISP as often as MNI to measure taxonomic abundances. In part because of a desire to determine the composition of human diet more precisely, and in light of the fact that different taxa have different numbers of identifiable bones, four zooarchaeologists have, since 1990, used assemblages with known ANI (actual number of individuals) to evaluate whether NISP or MNI provides the most accurate measure of ANI. ANI data from ethnoarchaeological and historical contexts suggest taxonomic abundance data quantified as NISP or MNI are ordinal scale at best, something previously shown to be highly probable. Experimental data used to evaluate the accuracy of NISP and MNI as measures of ANI are either inappropriate or not designed to assess which quantitative unit produces the most accurate measure. A new quantitative unit proposed as an alternative to MNE (minimum number of [skeletal] elements)—the NDE (number of distinct elements)—is said to provide proportional abundances of taxa but demonstrably undercounts skeletal parts and fails to provide ratio scale abundance data.

Observations on the history of zooarchaeological quantitative units: Why NISP, then MNI, then NISP again?

Suggestions regarding the history of use of zooarchaeological quantitative units known as the number of identified specimens (NISP) and minimum number of individuals (MNI) have previously had little empirical substantiation. Analysis of the North American zooarchaeological literature that appeared between 1900 and 1999 indicates there was (i) an increase in the use of MNI relative to NISP between 1900 and the 1970s, (ii) a subsequent decrease in the use of MNI and increase in the use of NISP, and (iii) an increase in the use of both NISP and MNI in the same publication after the 1970s. Inspection of a sample of articles indicates the last is the result of a shift to research questions requiring use of both units and also unresolved weaknesses in both NISP and MNI as measures of taxonomic abundances. The former reflects increased inquiry into taphonomic histories, the latter reflects the volume of programmatic literature evaluating strengths and weaknesses of NISP and MNI that peaked between about 1975 and 1995, and subsequently decreased with no clear consensus. Zooarchaeology textbooks of the last two decades present mixed signals regarding which quantitative unit to use.

Los Batanes: A trap for the Pyrenean wild goat during the Late Pleistocene (Spain)

Los Batanes cave (1025 m.a.s.l.) is a karstic system formed by pressure ducts, near the locality of Biescas in the Upper Gallego Valley (Huesca, Spain). Many fossil remains were retrieved from the cave sediments. The minimum sediment calendar age was determined to be 12.770 ± 60 BP. This date indicates that the deposit was formed before the Late Pleistocene-Early Holocene transition at the Pyrenees. Here we analyze the faunal assemblage of the site that is only conformed by remains of Pyrenean wild goat (Capra pyrenaica pyrenaica), which is a recently extinct subspecies of Iberian wild goat (C. pyrenaica). In the site, 1079 remains of Iberian wild goat have been recovered being one of the most numerous both in minimum number of individuals (MNI) and number of identified specimens (NISP) recovered until now in a natural trap in the Spanish Pyrenees. The population of Los Batanes shows a wide range of ages between juvenile and senile individuals and the MNI estimated from the number of right metatarsus is nine.The taphonomic features indicate that the cave acted as a trap for the goats which inhabited the vicinity of the cave. These goats probably stumbled and fell to the bottom of the pit and they could not get out, dying inside. Due to the origin of the accumulation numerous complete bones have been recovered. These fossil allow us to perform a biometric analysis that indicates that population of Los Batanes is in the range size of other goats from Late Pleistocene of Iberian Peninsula.

The Howieson's Poort fauna from Sibudu Cave: Documenting continuity and change within Middle Stone Age industries

The Howieson's Poort (HP; ∼65–59 ka) continues to be a source of interest to scholars studying human behavioral evolution during the Late Pleistocene. This is in large part because the HP preserves evidence for a suite of innovative technologies and behaviors (including geometric backed tools and engraved ostrich eggshell), but also because the disappearance of the innovative behaviors associated with this phase is not well understood. Here, I present taphonomic and taxonomic data on the full sample of macromammal remains excavated from the HP deposits at Sibudu Cave under the direction of Lyn Wadley. With a total number of identified specimens (NISP) of 5921, Sibudu provides the largest sample of HP fauna published to date. Taken as a whole, the data suggest a focus on a diverse range of prey. Ungulates dominate the assemblage, as do taxa that preferentially inhabit closed (particularly forested) environments. Small bovids are common throughout; blue duiker (Philantomba monticola) alone comprises ∼33% of the total NISP. A diverse smaller game assemblage is also present. Taphonomic data implicate humans as the primary contributor to the fauna; however, low levels of gastric etching (∼1% of the NISP) suggest that non-human agents may have played some role in the accumulation of the smaller game. Despite broad similarities in the fauna, a number of directional trends are evidenced. Most notably, the lowermost deposits of the HP contain the highest frequency of blue duiker and other small ungulates, taxa which prefer closed environments, and miscellaneous smaller game. All of these decline throughout the HP, and these differences are statistically significant. After considering possible explanations for these trends, I discuss the potential implications of the variation evidenced in the assemblage to our understanding of the onset—and disappearance—of this important substage of the MSA.

Problems of Identification and Quantification in Archaeozoological Analysis, Part I: Insights from a Blind Test

In archaeozoology, counts are generally considered as replicable data that accurately represent the initial abundances of elements, individuals, or taxa, although perhaps only at the ordinal scale. However, few studies have tested these assumptions with control data. To improve our knowledge of these issues, we conducted a blind test that involved the analysis of two large experimental samples composed of modern ungulate specimens of known element and taxon. Because the samples differed in level of fragmentation, the blind test provides substantial information on the impact of bone processing on faunal identification and quantification. Our results suggest that Number of Identified SPecimens (NISP) and Minimum Number of Elements (MNE) provide measures of abundance for whole assemblages and for samples limited to non-long bones that are both replicable and accurate at the ratio scale. However, the same metrics generally failed, even at the ordinal level, to predict abundances in analyses restricted to long bones and long bone portions. Given these mixed results, it seems judicious, in agreement with the current majority view among archaeozoologists, to treat faunal tallies as ordinal-level information. Despite issues of reproducibility and the difficulty of aggregating counts with MNE, the blind test also indicates that this measure is more robust at predicting skeletal abundances than NISP. Substantial variations in rates of long bone fragmentation and identification probably explain the poorer performance of NISP in the blind test.

Problems of Identification and Quantification in Archaeozoological Analysis, Part II: Presentation of an Alternative Counting Method

Archaeozoologists commonly use Number of Identified SPecimens (NISP) and Minimum Number of Elements (MNE) as measures of anatomical abundances. According to a blind test examining the reproducibility and accuracy of identifications of ungulate remains (Morin et al., Part I, Journal of Archaeological Method and Theory, doi: 10.1007/s10816-016-9300-4), NISP provides estimates of skeletal abundances that are less robust than those based on MNE. However, although results were improved with the latter method, MNE is not free of problems. Here, we show through an analysis of paired NISP-MNE data for 24 classes of elements that MNE is prone to inflate the representation of rare parts (as measured by NISP), a phenomenon more strongly expressed in certain elements than in others. Moreover, some elements show a wide scatter of points, which raises issues of data reproducibility. MNE is also known for being seriously affected by aggregation methods. These fundamental problems severely undermine the value of MNE as a measure of abundance. This article introduces an alternative counting method that avoids many of the weaknesses of MNE. This counting method, called the Number of Distinct Elements (NDE), focuses on the occurrence of pre-determined, invariant landmarks counted on mutually exclusive specimens. Preliminary experimental results suggest that NDE counts are robust predictors of skeletal, and perhaps taxonomic, abundances. Moreover, the NDE approach eliminates the complex and time-consuming task of spreading or drawing specimens to identify fragment overlap. Furthermore, NDE values are additive and easy to calculate. Given these features, the NDE approach represents a compelling alternative to MNE in archaeozoological analysis.

Evaluating three methods to estimate the number of individuals from a commingled context

An estimate of how many individuals are represented in a commingled assemblage is important to interpret the wider context (archaeologically or forensically), for further analyses, and for palaeodemographic studies. The aim of this study was to establish whether the Minimum Number of Individuals (MNI) and Minimum Number of Elements (MNE) estimates produced by three different methods (traditional MNI (White, 1953); zonation method (Knüsel and Outram, 2004); the landmark method (Mack et al., 2015)) are the same or, if different, to evaluate these differences. The methods were applied to an assemblage recovered from a Spanish medieval cemetery from Navarra and used to estimate the Number of Identified Specimens (NISP), the MNI and the MNE according to each method. Fragmentation analysis was also performed. The results indicate different values of MNE and MNI when applying different methods. White's MNI equaled 84; the MNI by zones 68; and the MNI by landmarks 61. All methods showed differences but the disparity between the traditional MNI and the MNI by landmarks was highest. Furthermore, the results indicate that different methods had a minimal impact on estimates of smaller bones. Individuals may be double counted by White's MNI count and the zonation method, when refitting exercises cannot be applied to all fragments from the same context or site, or if the 50% presence rule is not applied to the method. Finally, these findings have important implications for future analysis of commingled remains, because MNE and MNI estimates, as well as levels of fragmentation can impact on decisions made to further analyse the collection. Further research on a known collection is needed to identify the most reliable method to use.

Four thousand years of western Torres Strait fishing in the Pacific-wide context

Situated just north of Cape York Peninsula, Australia, the western Torres Strait islands reveals the earliest archaeological evidence for human occupation of Torres Strait at 9000years ago on Badu. The first evidence for marine resource use of fish and turtle dates to ~7200cal BP. We describe the salient information on marine exploitation from the ethnographic record, summarise the evidence for shark/ray and finfish exploitation from faunal assemblages excavated at six sites (dating to the past 4000years) using similar standardised techniques, then compare the archaeological data for fishing to other sites across the tropical Pacific Islands. Using the number of identified specimens (NISP) for all six western Torres Strait assemblages (n=1927), sharks and rays (taxa including Elasmobranchii, Myliobatidiformes or stingrays, four shark familes, and the porcupine ray or Urogymnus asperrimus) accounted for 59.9% of all identified elements. The most abundant finfish were wrasses (Labridae, mostly Bodianus sp.) at 21.3%, parrotfish (Scaridae) 6.3% and groupers (Serranidae) at 2.5%. Seven families provided the remaining ~10%. Fish size was estimated by measuring the diameter of finfish and shark/ray vertebrae, length of emperor otoliths and widths of pharyngeal grinding clusters of parrotfish and wrasses. Live length was commonly <10cm for finfish and ~1m for sharks. This small size of the fish represented in sites throughout western Torres Strait reflects a forager capture strategy probably undertaken primarily by women and children walking along the reef flat at low tide gleaning fish stranded in tide pools and shallow water channels. Ethnographically, small fish are targeted due to their superior ‘sweeter’ taste compared to larger fish. Complementary ethnographic and archaeological information indicates that the bulk of marine protein came from socially prestigious dugongs and especially turtles hunted by men. Small sharks may have been targeted when attracted to shoreline dugong/turtle butchering events. No archaeological evidence for fishing technology such as shell fishhooks has been recovered from archaeological excavations. The western Torres Strait islands are unique within the greater tropical Pacific islands region because of: 1) low fish species richness of the assemblages; 2) dominance of sharks and rays in the combined assemblages accounting for ~60% NISP; and 3) small reconstructed length of captured finfish and shark/rays; and, as a consequence, the lack of evidence for an intensive fishery. Further studies of fish bone assemblages from across the 150km of Torres Strait may demonstrate that this is a common pattern for the region but, if not, documenting the variability of the fishery will be an important task.