North American Western Interior Research Articles

Unconformity generation and the shift from storm‐dominated to tide‐dominated processes in a Jurassic retroarc foreland basin: Insights from ichnology

AbstractAn enigmatic transition from the storm‐dominated, offshore to lower shoreface deposits of the Redwater Shale Member (Sundance Formation) to the overlying mixed tidal and aeolian Windy Hill Sandstone (Morrison Formation) in the Oxfordian of the North American Western Interior has long been a source of intrigue. Previously proposed drivers include the progradation of a large, tide‐dominated delta onto a storm‐dominated shelf, a complete reorganisation of the basin's hydrodynamics and climate, or the development of a regional unconformity (termed the J‐5). In south‐eastern Wyoming, the Redwater Shale is characterised as an offshore to distal shoreface deposit with glauconitic siltstones and sandstones punctuated by coquinoid and sandy tempestites and hosting a Cruziana Ichnofacies. The Windy Hill Sandstone, a time‐transgressive, sand‐rich, intertidal succession with classic Pteraichnus and stressed Skolithos Ichnofacies, sharply overlies the Redwater Shale and records an abrupt basinward shift in facies that accompanied at least tens of metres of sea‐level fall. New, detailed sedimentological, ichnological and architectural data collected across this transition in the study area provide fresh insights into the depositional history of these units and demonstrates the existence locally of a composite J‐5 unconformity. The unconformity developed as tectonically driven, prograding shoreline trajectories of the Redwater Shale gave way to degrading trajectories of the Windy Hill Sandstone, leading to a forced regression and formation of a regressive surface of marine erosion. The sharp juxtaposition of intertidal flat facies (Pteraichnus Ichnofacies) directly upon offshore to lower shoreface deposits (Cruziana Ichnofacies) is the key to recognising the unconformity and proves the value of the previously underutilised ichnological data.

An unusual plane tree from the Early Cretaceous of Kansas, USA

We present a new genus and species of platanaceous inflorescences from the Hoisington III locality of Barton County, Kansas, USA. This locality lies within the Albian Dakota Formation, which was formed from deposits along the eastern margin of the mid-Cretaceous North American Western Interior Seaway. The elongate inflorescences documented here are unusual for Platanaceae, which is known for its globose, unisexual inflorescences. Distefananthus hoisingtonensis Huegele and Wang gen. et sp. nov. has elongate, cylindrical, ellipsoid, pedunculate inflorescences bearing apocarpous achenes with long styles, borne in florets of 3-4, with over 100 fruits per inflorescence, and stamens with short filaments and hairy apical connectives. The small and relatively fixed number of parts per floret ((3-)4 fruits and c. 3-4 stamens) is seen in other Cretaceous Platanaceae inflorescences and may have facilitated insect pollination. The state of perianth development is unclear, and the fruits lack basal dispersal hairs. The Hoisington material encompasses a developmental sequence; younger inflorescences bear a mix of stamens and immature carpels (thin with long styles) while mature inflorescences bear developed fruits (thicker and often with broken styles) and have lost all stamens. Like the typical globose capitula, the elongate inflorescences probably originated from the reduction of a compound inflorescence. The elongate inflorescences may represent a less compressed, ancestral or pleisiomorphic condition for Platanaceae. The only other species from this family known for elongate inflorescences is from the Paleocene and Eocene of western North America. The inflorescences from Hoisington co-occur with Sapindopsis leaves. At this locality, Sapindopsis powelliana occurs at a similar frequency to the elongate inflorescences and is a possible candidate for the leaves associated with them. The Hoisington material marks the earliest known occurrence of bisexual inflorescences in Platanaceae. The discovery of Distefananthus hoisingtonensis offers new insight into the diversity of platanaceous inflorescences in the mid-Cretaceous, a critical early period in the history of Platanaceae.

A tropically hot mid-Cretaceous North American Western Interior Seaway

AbstractThe mid-Cretaceous thermal maximum (KTM) during Cenomanian to Santonian times from ca. 100 to 83 Ma is considered among Earth's warmest sustained intervals of the Phanerozoic. The time interval is also characterized by major paleoceanographic changes in the form of an oceanic anoxic event and the flooding of epicontinental seaways, such as the Western Interior Seaway in North America. We report carbonate clumped isotope (Δ47) paleotemperatures (TΔ47) of the KTM measured from Cenomanian oyster fossils of the Western Interior Seaway. Following screening of specimens for carbonate diagenesis and exclusion of geographic zones with evidence consistent with solid-state Δ47 reordering, a mean TΔ47 of 28–34 °C (95% confidence interval for the standard error of mean) for primary oyster calcite quantifies extreme mid-latitude warmth in North America. When combined with existing Campanian and Maastrichtian marine TΔ47 records, the new data constrain Late Cretaceous temperature trends underlying the evolution of North American faunal and stratigraphic records. These TΔ47 data from the peak KTM highlight the potential of this proxy to quantitatively resolve the upper thermal limits of Phanerozoic greenhouse climates.

The easternmost occurrence of Saurornitholestes from the Judith River Formation, Montana, indicates broad biogeographic distribution of Saurornitholestes in the Western Interior of North America

ABSTRACT Small-bodied dromaeosaurid and troodontid theropod dinosaurs are present in most of the terrestrial Late Cretaceous formations of the North American Western Interior, but are represented predominantly by shed teeth and more rarely by isolated skeletal elements and associated skeletons. The eudromaeosaurian dromaeosaurid Saurornitholestes is among the most commonly recovered small theropods from North America, known from abundant shed teeth, isolated cranial and postcranial elements, and several partial or near-complete skeletons from the Campanian aged Two Medicine Formation of Montana and the Dinosaur Park Formation of Alberta. Conversely, few Saurornitholestes specimens beyond shed teeth have been recovered from temporally equivalent units of the Judith River Formation of Montana. This study reports the first cranial material, a near-complete frontal, referable to Saurornitholestes from the Judith River Formation of Montana. This specimen is the easternmost occurrence of this taxon and was recovered approximately 400 km east of the localities which have produced referred Saurornitholestes specimens in the Two Medicine Formation, proximal to the Cordilleran thrust front. As such, this new specimen indicates that Saurornitholestes was biogeographically widespread and occupied both inland and coastal environments across the northern portions of the Late Cretaceous Western Interior of North America.

Regional chronostratigraphic synthesis of the Cenomanian-Turonian Oceanic Anoxic Event 2 (OAE2) interval, Western Interior Basin (USA): New Re-Os chemostratigraphy and 40Ar/39Ar geochronology

AbstractFluctuations in depositional conditions during the onset of severe climate events in Earth history predispose stratigraphic archives to hiatuses, often hindering complete reconstructions of paleoclimate events and their triggers. Several studies have proposed that a hiatus of unknown duration exists at the base of Oceanic Anoxic Event 2 (OAE2) in the North American Western Interior Basin at the base Turonian global boundary stratotype section and point (GSSP) in Pueblo, Colorado, which potentially influences integrated radioisotopic, biostratigraphic, and astrochronologic age models of the Cenomanian-Turonian boundary interval. To quantify the duration of this hiatus, refine the chronology of OAE2, and assess marine geochemical perturbations associated with the onset of the event, we present new 40Ar/39Ar dates from regional bentonites along with a new proximal-distal chemostratigraphic transect of the epeiric Western Interior Basin (WIB), including initial osmium isotope (Osi) and stable carbon isotope (δ13C) data. The new 40Ar/39Ar age determinations confirm and further constrain previous estimates of Cenomanian-Turonian boundary timing. Further, the regional chemostratigraphic synthesis demonstrates the conformity of the OAE2 successions correlated to Pueblo, shows that the duration of the lag between the onset of the Osi and δ13C excursions is ∼60 k.y., and thus constrains the magnitude of the pre-OAE2 hiatus in Pueblo to less than this value. The new astronomically tuned, conformable Osi record across the onset of OAE2 captures a geologically rapid onset of large igneous province volcanism, consistent with other records, such that the addition of CO2 to the ocean-atmosphere system may have driven changes in marine carbonate chemistry. Additionally, the refined chronostratigraphy of OAE2 and the Cenomanian-Turonian boundary in the central WIB improves correlation with other records, such as those in the Eagle Ford Group, Texas. The correlations highlight that discrepancies among OAE2 age models from globally distributed sections commonly stem from differing definitions of the event and uncertainties associated with astronomical tuning, in addition to stratigraphic preservation.

The early Paleogene stratigraphic evolution of the Huerfano Basin, Colorado

ABSTRACTSedimentary basins throughout the North American Western Interior contain a record of Late Cretaceous through Eocene deposition related to the Laramide orogeny. The typical stratigraphic progression includes an uppermost Cretaceous fluvio-deltaic geologic formation that is unconformably overlain by an alluvial or paludal Paleocene geologic formation. The Paleocene unit is usually characterized by drab overbank facies, and overlain by an interval of amalgamated fluvial sand bodies. The overlying Eocene geologic units are characterized by red bed overbank facies. These major stratigraphic changes have been variably linked to long-wavelength dynamic subsidence, local uplift, and climatic shifts. Herein, we evaluate the depositional history of the Huerfano Basin of south-central Colorado in this overarching context. Our study presents a detailed lithofacies analysis of the Poison Canyon, Cuchara, and Huerfano Formations integrated with a new bulk (1) organic carbon isotope record, n = 299 measurements (Data Supplement 1A); and (2) magnetic record, n = 247 measurements (Data Supplement 1B). We interpret that the Paleocene Poison Canyon Formation was deposited by a braided or coarse-grained meandering river system with relatively poorly drained floodplains. The Eocene Huerfano Formation was likely deposited by a coarse-grained meandering river system with a comparatively well-drained floodplain. This pattern mirrors other Laramide basins, and is likely related to a regional drying pattern linked to long-term warming during the early Paleogene. Age of the intervening Cuchara Formation is poorly resolved, but is an anomalously thick and coarse-grained fluvial unit, with evidence for extensive reworking of floodplain deposits and a moderate coarsening-upward pattern. The Cuchara Formation is associated with magnetic trends that suggest greater oxidation and weathering, and greater variability in rainfall patterns, as well as a subtle negative shift in carbon isotope values. This pattern indicates a period of widespread progradation within the basin, potentially related to a major Laramide uplift event that affected Colorado’s Wet Mountains, Front Range, and Sangre de Cristo Mountains.

Evolutionary stasis, ecophenotypy and environmental controls on ammonite morphology in the Late Cretaceous (Maastrichtian) Western Interior Seaway, USA

AbstractWe test for the presence of evolutionary stasis in a species of Late Cretaceous ammonoid cephalopod,Hoploscaphites nicolletii, from the North American Western Interior Seaway. A comprehensive dataset of morphological traits was compiled across the entire spatial and temporal range of this species. These were analysed in conjunction with sedimentologically and geochemically derived palaeoenvironmental conditions hypothesized to apply selective pressures. All changes in shell shape were observed to be ephemeral and reversable, that is, no unidirectional trend could be observed in any of the morphological traits analysed. Correlations between palaeoenvironmental conditions and morphological traits suggests ecophenotypic processes were at play; however, either environmental changes were too minor and/or provided no isolating mechanism to drive speciation. These data support mechanisms of stasis such as homogenizing gene flow or stabilizing selection under a fluctuating optimum (probably reflecting spatiotemporally heterogeneous palaeoenvironmental conditions). Finally, changes in shell size were not significantly associated with changes in shell‐specific δ18O, despite a correlation between shell size and δ18O averaged across horizons. This suggests a mismatch in scales of geochemical sampling that supports caution when making broad interpretations based on averaged geochemical data.

A unique Lower Paleocene shelf-sea chalk in the eastern U.S. Gulf coastal plain (Clayton Formation, western Alabama): Implications for depositional environment, sea-level dynamics and paleogeography

The deposition of carbonate oozes in shelf-sea settings so prevalent during the Cretaceous generally ceased by the onset of the Paleogene in response to sea-level fall. However, the Lower Paleocene (Danian) Clayton Formation exposed at Moscow Landing in western Alabama, U.S.A., contains a thin chalk bed—the Clayton chalk—that records a later, final phase of shelf-sea carbonate ooze deposition in the U.S. eastern Gulf coastal plain. Sedimentologic character, ichnology, and foraminiferal assemblages indicate deposition in an outer shelf setting, comparable to that for Cretaceous chalks exposed in the Gulf region (i.e., Campanian Demopolis Chalk), at depths of 90 m or more, during a brief transgressive pulse peaking circa 63.8–63.2 Ma. This transgression occurred ~1 Myr after the maximum expansion of the Danian Cannonball Sea in the North American western interior, presumably reflecting different controls on sea-level dynamics (eustasy vs. Laramide tectonism and isostasy). Although regional paleogeography for the Early Paleocene is poorly constrained, deposition of the Clayton chalk suggests that the eastern Gulf seaway at times extended much further north than implied by the presently mapped updip limit of preserved Paleocene marine strata. The apparent geographic restriction of this unique Paleocene shelf-sea chalk indicates that relative sediment starvation and associated carbonate ooze deposition may have been influenced by local structural controls operating along the eastern margin of the Mississippi embayment.

Astronomical pacing of relative sea level during Oceanic Anoxic Event 2: Preliminary studies of the expanded SH#1 Core, Utah, USA

AbstractProximal marine strata of the North American Western Interior Basin (WIB) preserve a rich record of biotic turnover during Oceanic Anoxic Event 2 (OAE2; ca. 94 Ma), a pronounced Late Cretaceous carbon cycle perturbation interpreted to reflect global warming, widespread hypoxia, and possible ocean acidification. To develop a more robust synthesis of paleobiologic and geochemical data sets spanning this Earth-life transition, we drilled the 131 m Smoky Hollow #1 Core (SH#1), on the Kaiparowits Plateau of southern Utah, USA, recovering the Cenomanian–Turonian Boundary (CTB) interval in the Tropic Shale Formation. A 17.5 m positive excursion in high-resolution bulk carbon isotope chemostratigraphy (δ13Corg) of SH#1 characterizes the most expanded OAE2 record recovered from the mid-latitudes of the WIB.Depleted values in a paired carbonate carbon isotope (δ13Ccarb) chemostratigraphy cyclically punctuate the OAE2 excursion. These depletions correspond to intervals in the core with a higher degree of carbonate diagenesis and correlate well to an existing sequence stratigraphic framework of flooding surfaces in the shoreface facies of the Markagunt Plateau (∼100 km west). We detect statistically significant evidence for astronomical cycles in the δ13Ccarb data set, imparted by diagenesis at flooding surfaces, and develop a floating astronomical time scale (ATS) for the study interval. Stable eccentricity cycles (405 k.y.) align with stratigraphic sequences and associated trends in sedimentation rate, and short eccentricity cycles (∼100 k.y.) pace nested parasequences. These results confirm an astronomical signal and, therefore, climatic forcing of relative sea level during OAE2 in the WIB. Furthermore, cross-basin correlation of the ATS and expanded δ13C chemostratigraphy of SH#1 suggests that these transgressive-regressive parasequences modulated siliciclastic sediment delivery in the seaway and contributed to deposition of prominent rhythmically bedded CTB units across the WIB, including the Bridge Creek Limestone. The presented approach to analysis of these proximal offshore siliciclastic facies links early diagenetic influences on chemostratigraphy to astronomically modulated sequence stratigraphic horizons, and helps to resolve rates of paleobiologic and paleoenvironmental change during a significant Mesozoic carbon cycle perturbation.

<em>Puercolestes</em> and <em>Betonnia</em> (Cimolestidae, Mammalia) from the early Paleocene (Puercan 3 Interval Zone) of northeastern Montana, U.S.A.

In northeastern Montana, fossil localities in the Garbani Channel Complex and other early Paleocene (Puercan 3 Interval Zone) localities are preserved in the Tullock Member of the Fort Union Formation. They document an early phase in the recovery of the terrestrial fauna of the North American Western Interior after the mass extinction marking the Cretaceous-Paleogene (K-Pg) boundary. The cimolestids Puercolestes simpsoni and Betonnia tsosia were typified on fragmentary jaws and isolated teeth found in Puercan 2 and 3 Interval Zones (Pu2, Pu3) in the Nacimiento Formation, San Juan Basin, New Mexico. The available samples of these genera from both New Mexico and Montana are small and dominantly consist of isolated teeth. Characters of upper cheek teeth, P4 and M1–M3, justify provisional recognition of Puercolestes sp. cf. Pu. simpsoni and the somewhat smaller Betonnia sp. cf. Be. tsosia in Pu3 local faunas in the Tullock Member. In contrast, discovery of characters distinguishing the isolated lower cheek teeth, p4s and m1–m3s, of these species must await recovery of dentulous dentaries documenting the patterns of morphological variation of their lower dentitions. Fossils from the Tullock Member add support to the current interpretation that cimolestids were taxonomically diverse and geographically widespread but relatively rare members of the faunas of the North American Western Interior during approximately the first million years of the Paleocene.

Organic matter source and thermal maturity within the Late Cretaceous Niobrara Formation, U.S. Western Interior

The Late Cretaceous sedimentary record of the North American Western Interior Seaway is characterized by cyclic deposition of organic carbon-rich sediments. One notable interval during the late Coniacian-Santonian is recorded by the Niobrara Formation. The organic carbon-rich interval within the Niobrara Formation has been identified as Oceanic Anoxic Event (OAE) 3. Understanding the reason for this distribution of organic carbon within the Niobrara Formation requires a refined understanding of the source and maturity of the organic matter. In this study, we present lipid biomarker records from the USGS Portland #1 core (Cañon City, CO) to constrain the thermal maturity of the organic matter and the differing contributions of organic matter sources. Sterane and hopane thermal maturity indices indicate that the samples are somewhat immature with respect to oil formation and that there is strong agreement between different proxies for thermal maturity. Based on the distribution of n-alkanes, steranes, and hopanes, there is a significant increase in the contribution of algal organic matter during and after OAE 3, coeval with increased organic carbon accumulation. Although a consistent terrestrial contribution is observed, it is only a minor source of organic matter at the Portland core location and does not drive increased organic matter accumulation during OAE 3. Of particular note is the consistent influence of even-over-odd predominantly mid-chain length (C21 to C25) organic matter. This observation within the brackish to marine, not methanogenic WIS represents an expansion of the depositional settings in which even-over-odd predominance has been observed in mid-chain length n-alkanes. Pristane (Pr) and phytane (Ph) abundances are inconsistent with a redox control on Pr/Ph ratios and suggest an increase in the delivery and/or preservation of phototrophic organic matter as the source for pristane and phytane in the Portland core.

Chapter 2: Inoceramid Bivalves from the Coniacian and Basal Santonian (Upper Cretaceous) of the Western Canada Foreland Basin

ABSTRACT Inoceramid bivalves are the dominant invertebrate fauna of the Coniacian and basal Santonian of the Western Canada Foreland Basin in western Alberta. In the upper lower Coniacian through to basal Santonian, six successive faunas are recognized, which provide the basis for corresponding, formally defined inoceramid zones. From bottom upward these are the zones of: Cremnoceramus crassus crassus /C. deformis deformis, Inoceramus gibbosus, Volviceramus koeneni, Volviceramus involutus, Sphenoceramus subcardissoides, and Sphenoceramus ex gr. pachti. Particular faunas represent assemblages known widely from the Euramerican biogeographic region, although they characterize mostly its northern, boreal area. The inoceramid-based biostratigraphic scheme allows correlation with other parts of the North American Western Interior and with parts of the Euramerican biogeographic region. The studied succession provides a good record of the Inoceramus gibbosus Zone, which characterizes the topmost lower Coniacian. ...

<em>Procerberus</em> (Cimolestidae, Mammalia) from the Latest Cretaceous and Earliest Paleocene of the Northern Western Interior, USA

Three species of the cimolestid Procerberus are currently recognized in the northern North American Western Interior in latest Cretaceous and earliest Paleocene (Puercan North American Land Mammal Age) faunas: P. formicarum, P. andesiticus, and P. grandis. Analysis of a new topotypic sample of P. formicarum from the Bug Creek Anthills locality provides an estimate of the range of variation of its postcanine dentition. The three currently recognized species occur in Puercan 1 (Pu1) interval zone faunas, but two other occurrences indicate that the genus originated and initially diversified prior to the Cretaceous/Paleogene boundary. Rare occurrences at several localities and entries in faunal lists suggest even greater taxonomic diversity. Limited evidence suggests continued diversification in the later Puercan and possible survival of the genus into the Torrejonian. Procerberus grandis or a closely related species may be a sister group of primitive taeniodonts.

Latest Cretaceous eggshell assemblage from the Willow Creek Formation (upper Maastrichtian – lower Paleocene) of Alberta, Canada, reveals higher dinosaur diversity than represented by skeletal remains

The Willow Creek Formation (upper Maastrichtian – lower Paleocene) of southwestern Alberta is a poorly fossiliferous formation that preserves a low end-Cretaceous dinosaur diversity compared with most correlative terrestrial deposits in the North American Western Interior. Although only three dinosaur taxa are known from skeletal remains (Tyrannosaurus rex, Hadrosauridae indet., and Leptoceratopsidae indet.), study of hundreds of dinosaur eggshells recovered from several sites in the formation reveals the presence of a more diverse dinosaur assemblage. Morphological and histological analyses of the eggshells indicate the presence of at least seven dinosaur ootaxa (Continuoolithus, Montanoolithus, Porituberoolithus, Prismatoolithus spp., Spheroolithus spp.). These ootaxa are referable to at least two ornithopod and five small theropod species, likely including dromaeosaurids, oviraptorosaurs, and troodontids. When considering the taxonomic affinity of eggshells and skeletal remains, the present study triples the known dinosaur diversity of the Willow Creek Formation, increasing the number of dinosaurs from three to at least nine species. Probable ornithopod eggshells comprise most of the eggshells preserved, although small theropods were likely an important component of the Willow Creek ecosystem, as most ootaxa can be ascribed to these dinosaurs. Although fossil bones are rarely found in the Willow Creek Formation, fossil eggshells are common compared with most other dinosaur-bearing formations in Alberta. The caliche-bearing deposits, indicative of arid to semi-arid conditions, typical of the formation were likely conducive to the preservation of calcareous eggshells.

Stable isotope paleohydrology and chemostratigraphy of the Albian Wayan Formation from the wedge-top depozone, North American Western Interior Basin

Understanding of the role of atmospheric moisture and heat transport in the climate system of the Cretaceous greenhouse world represents a major challenge in Earth system science. Stable isotopic paleohydrologic data from mid-Cretaceous paleosols in North America, from paleoequatorial to paleoArctic latitudes, have been used to constrain the oxygen isotope mass balance of the Albian hydrologic cycle. Over the range from 40°–50°N paleolatitude, sideritic paleosols predominate, indicating paleoenvironments with positive precipitation-evaporation (P-E) balances. Local exceptions occur on leeward side of the Sevier Orogen, where calcic paleosols in the wedge-top depozone record paleoenvironments with negative P-E balances in the orographic rain shadow. Stratigraphic sections in the Wayan Formation of Idaho (WF) were sampled from the wedge-top depozone. The units consist of stacked m-scale mudstone paleosols separated by m-scale sandstone-siltstone beds. Sections were sampled for organic carbon isotope profiles, and B-horizons from 6 well-developed paleosols were sampled for detrital zircons to determine maximum depositional ages. The first of these from the WF has produced a U-Pb concordia age of 101.0±1.1 Ma. This same WF section has produced a stratigraphic trend of upwardly decreasing δ13C values ranging from–24‰ upwards to–27‰ VPDB, suggesting correlation to the late Albian C15 C-isotope segment. Pedogenic carbonates from the WF principally consist of micritic calcite, with carbon-oxygen isotope values that array along meteoric calcite lines (MCLs) with δ18O values that range from–9.47‰ up to–8.39‰ VPDB. At approximately 42°N paleolatitude, these MCL values produce calculated paleoprecipitation values of–8.12‰ to–7.04‰ VSMOW, a range that is consistent with the estimates produced from other proxies at the same paleolatitudes across North America. These results indicate that despite the orographic rain shadow effect, the processes of meridional atmospheric moisture transport in this locale were similar to those in more humid mid-latitude paleoenvironments elsewhere in the continent.

Biostratigraphy and Inoceramus survival across the Cenomanian–Turonian (Cretaceous) boundary in the Ram River section, Alberta, Canada

AbstractThe biostratigraphy of the uppermost Cenomanian – Lower Turonian succession in the lower part of the Blackstone Formation exposed in the Ram River (Alberta, Canada), is interpreted in terms of the standard inoceramid/ammonite zonation of the interval. Four successive inoceramid zones are recognized, those ofInoceramus pictus,Mytiloides puebloensis,M. kossmati, andM. mytiloides, as established in the stratotype section at Pueblo, Colorado. Their correlation to Pueblo is confirmed by ammonite data.The mid-Early Turonian zone ofM. kossmatiyielded an assemblage ofInoceramus, with species showing close affinity to the latest Cenomanian lineages. This multi-species sample proves the survival ofInoceramuslineages into the otherwiseMytiloides-dominated Early Turonian, and indicates that their disappearance from the record of the North American Western Interior was not because of their extinction. It is suggested that the apparent lack ofInoceramusin Lower Turonian strata is due to an extremely low population abundance in the Early Turonian sea.

Micropaleontological evidence for redox changes in the OAE3 interval of the US Western Interior: Global vs. local processes

The Coniacian-Santonian interval has been proposed as the youngest of the Cretaceous ocean anoxic events (OAE3), but this designation has long been debated. OAE3 is associated with a long-lasting (∼3 myr) succession of black shales from the central and South Atlantic, Caribbean region, and the North American Western Interior; in the Western Interior it is characterized by an abrupt increase in total organic carbon (TOC) and corresponding trace metal indicators for anoxia. However, the modern concept of OAEs is predicated on detection of global carbon cycle perturbations as recorded by substantial carbon isotope excursions (CIE), and the protracted Coniacian-Santonian black shale interval does not have a large CIE. A more conservative definition of OAE3 might limit the event to the modest positive carbon isotope excursion restricted to the upper Coniacian Scaphites depressus Ammonite Zone. Trace metal proxies suggest that oxygen levels abruptly declined prior to the onset of this CIE in the Western Interior Sea (WIS), but it is unknown whether regional anoxic conditions were confined to sediments/pore waters, or how anoxia may have affected the biota. In an effort to characterize the oxygenation history of the WIS and to better understand the nature of the hypothesized OAE3, we present micropaleontological evidence of declining oxygen in bottom waters prior to the event using benthic foraminifera, which are sensitive to dissolved oxygen. Changes in benthic foraminiferal abundances suggest a decline in oxygen at least 1-myr prior to the CIE (including a nadir immediately below the start of the excursion), improving bottom water oxygen during the CIE, and re-establishment of persistent anoxia following the isotope excursion. Anoxia endured for nearly 3 myr in the central seaway, showing some signs of recovery toward the top of the Niobrara Formation. Our findings suggest that declining oxygen concentrations in the seaway eventually reached a tipping point, after which dissolved oxygen quickly dropped to zero.The late Coniacian CIE is an exception to the trend of declining oxygen in the WIS, and part of a larger pattern in the oxygenation history of the Niobrara Formation which suggests that it does not adhere to standard black shale models. Transgressive periods, including the Fort Hays Limestone and the lower limestone unit of the Smoky Hill Shale (which corresponds to the CIE) are relatively oxic, while periods of highstand (i.e., most of the Smoky Hill) correspond to deteriorating oxygen conditions. This contrasts with the standard black shale model for sea level and oxygen, where transgressions typically correlate with maximum TOC enrichment, interpreted to result from both sediment condensation and oxygen deficiency. The association of global carbon burial/anoxia (as indicated by carbon isotopes) with a regional increase in oxygen and decrease in organic matter preservation is reminiscent of the Cenomanian-Turonian Greenhorn Limestone, which contains OAE2. In both cases, the facies are not typical black shales, but instead have appreciable carbonate content. Western Interior redox trends support the rejection of the original concept of a protracted Coniacian-Santonian OAE3 because it is not a distinct “event.” Increasing local oxygen during the late Coniacian CIE also argues against a narrower OAE designation for this event, because the excursion can't be tied to anoxia here or anywhere else it has been described. Nevertheless, the Late Coniacian Event (as we prefer to call this CIE) still represents an important perturbation of the global carbon cycle. This is emblematic of the shift away from widespread, discrete anoxic events during the ongoing paleoceanographic reorganization of the Late Cretaceous, even as large carbon cycle perturbations continued.

Iron limitation in the Western Interior Seaway during the Late Cretaceous OAE 3 and its role in phosphorus recycling and enhanced organic matter preservation

The sedimentary record of the Coniacian–Santonian Oceanic Anoxic Event 3 (OAE 3) in the North American Western Interior Seaway is characterized by a prolonged period of enhanced organic carbon (OC) burial. This study investigates the role of Fe in enhancing organic matter preservation and maintaining elevated primary productivity to sustain black shale deposition within the Coniacian–Santonian-aged Niobrara Formation in the USGS #1 Portland core. Iron speciation results indicate the development of a reactive Fe limitation coeval with reduced bioturbation and increased organic matter preservation, suggesting that decreased sulfide buffering by reactive Fe may have promoted enhanced organic matter preservation at the onset of OAE 3. An Fe limitation would also provide a feedback mechanism to sustain elevated primary productivity through enhanced phosphorus recycling. Additionally our results demonstrate inconsistencies between Fe-based and trace metal redox reconstructions. Iron indices from the Portland core indicate a single stepwise change, whereas the trace metal redox proxies indicate fluctuating redox conditions during and after OAE 3. Using Fe speciation to reconstruct past redox conditions may be complicated by a number of factors, including Fe sequestration in diagenetic carbonate phases and efficient sedimentary pyrite formation in a system with limited Fe supply and high levels of export production.

Memorial for Paul L. Heller (September 16, 1952–July 6, 2016)

Paul Heller in 2011. With no warning, and far too early, we have lost our friend, mentor, and colleague Paul Heller. Paul was part of the New York Jewish diaspora of the 1950s and ’60s, working his way west via Western Washington University (M.S., 1978). He completed his Ph.D. at the University of Arizona (1983) with Bill Dickinson, and ultimately landed at the University of Wyoming, where he was a member of the faculty from 1983–2016. Having caught the bug for sedimentation and tectonics, Paul emerged as one of the leaders of the next wave of tectonic sedimentologists, who focused on how new insights on basin mechanics could be used to understand stratigraphic patterns. Some of his earliest work involved deep-marine sedimentation, springing from fieldwork in the Pacific Northwest, and culminating in an important paper proposing the submarine-ramp model for sandy turbidites. Paul also was a pioneer of the idea of isotopic provenance, using isotopes as fingerprints of the origin of sediments. But perhaps Paul's most influential early work was that on the interplay of lithospheric flexure and sedimentation. He proposed the idea, widely accepted now but controversial at the time, that mountain building in foreland basins might be marked distally not by gravels but rather by fine-grained sediments, if the increasing load creates accommodation faster than it can be filled with deposits. Paul and colleagues synthesized much of this new quantitative approach in a basin analysis short course offered through the American Association of Petroleum Geologists. It found another expression through Paul's central and inspirational involvement in the development of experimental stratigraphy. The interplay of tectonic and surface processes remained central to Paul's work, up through his most recent publication on the dynamic topography of the North American Western Interior. But of course basins record more than tectonics, and …

A new clade of putative plankton-feeding sharks from the Upper Cretaceous of Russia and the United States

ABSTRACTEorhincodon casei from Russia and Megachasma comanchensis from the United States are two Cretaceous taxa initially described as putative planktivorous elasmobranchs, but the type specimens of these two taxa were subsequently reinterpreted to represent taphonomically abraded teeth of an odontaspidid, Johnlongia Siverson (Lamniformes: Odontaspididae). Here, we redescribe the type materials of ‘E. casei’ and ‘M. comanchensis’ and describe additional specimens of these species from other Late Cretaceous localities in Russia and the United States. These specimens demonstrate that (1) the two fossil taxa are valid species; (2) they warrant the establishment of a new genus of presumed planktivorous sharks, Pseudomegachasma, gen. nov., to accommodate the two species; and (3) the new genus is sister to Johnlongia and together constitute a new subfamily Johnlonginae, subfam. nov., tentatively placed in the family Odontaspididae sensu stricto. This taxonomic placement indicates that the putative planktivorous clade was derived from a presumed piscivorous form (Johnlongia), with an implication that Pseudomegachasma, gen. nov., evolved a plankton-eating habit independent of the four known planktivorous elasmobranch clades (Rhincodontidae, Megachasmidae, Cetorhinidae, and Mobulidae). It also indicates that planktivorous diets evolved independently at least three times in the order Lamniformes (i.e., Megachasmidae, Cetorhinidae, and Odontaspididae), and more significantly, Pseudomegachasma, gen. nov., would represent the oldest known plankton-feeding elasmobranch in the fossil record. The present fossil record suggests that Pseudomegachasma, gen. nov., evolved in a relatively shallow-water environment in Russia in the early Cenomanian or earlier and subsequently migrated to the North American Western Interior Seaway by the mid-Cenomanian.http://zoobank.org/urn:lsid:zoobank.org:pub:D5D0400FD438-4A95-8301-DD47991572F6SUPPLEMENTAL DATA—Supplemental materials are available for this article for free at www.tandfonline.com/UJVP